RECORD: Linsley, E. G. & Usinger, R. L. 1966. Insects of the Galápagos Islands. Proceedings of the California Academy of Sciences 33 (7): 113-196.

REVISION HISTORY: Transcribed (single key) by AEL Data, corrections by John van Wyhe 8.2009. RN1


[page] 113

PROCEEDINGS

OF THE

CALIFORNIA ACADEMY OF SCIENCES

FOURTH SERIES

Vol. XXXIII, No. 7, pp. 113-196; 1 table January 31, 1966

INSECTS OF THE GALÁPAGOS ISLANDS1

By
E. G. Linsley and R. L. Usinger
University of California, Berkeley

INTRODUCTION

When Charles Darwin visited the Galápagos Islands in September and October of 1835, he was impressed with the small size and dull colors of the insects he encountered, and of the insect fauna in general he commented that "excepting Tierra del Fuego, I never saw … so poor a country." G. R. Waterhouse, upon examining the insects which Darwin collected, reported that there was nothing in their appearance which would have led him to imagine that they had come from under the equator. Darwin described them as having a desert character, along with the plants and the birds.

Few entomologists who have visited the Archipelago would disagree with the generalization that the insect fauna is more desert than tropical in aspect, a fact in keeping with the arid nature of the islands, but tropical species are present, especially, but by no means exclusively, in the mangrove swamps along the shores and in the moist forests of middle elevations of the larger islands. The fauna is depauperate, as would be expected of oceanic islands populated by chance. For example, the continental British Isles have currently known more than three times as many families, ten times as many genera and thirty-two times as many species of insects as the Galápagos (table I). Although this differential will no doubt be reduced as the Galápagos fauna becomes better known, there is no reason to believe that it will ever be eliminated. However, although the fauna is poor in taxonomic entities, in favorable seasons it is rich in individuals, and the entomologists of the Galápagos International Expedition of 1964 brought back thousands of specimens.

In connection with our own studies we attempted to assemble as much as possible of the literature relevant to Galápagos insects prior to the Expedition, and to organize it in a systematic manner upon our return. The result gave us,

1 Contribution No. 27 from the Charles Darwin Foundation for the Galápagos Islands.

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TABLE I Comparison of recorded insect jauna of the Galápagos Archipelago with that of the British Isles.

Insect Orders Galápagos Archipelago British Isles*
Families Genera Species Families Genera Species
Collembola 2 2 2 9 62 261
Protura - - - 2 4 17
Thysanura 1 2 2 2 6 11
Diplura 1 1 1 1 1 12
Orthoptera 3 11 18 5 27 38
Blattaria 1 7 9 4 5 8
Mantodea 1 1 1 - - -
Dermaptera 1 2 2 3 7 9
Phasmida - - - - - -
Isoptera 2 2 6 - - -
Embioptera - - - - - -
Plecoptera - - - 6 15 32
Odonata 3 6 7 9 21 42
Ephemerida - - - 8 19 46
Corrodentia - - - 11 33 68
Mallophaga 2 25 61 7 58 252
Anoplura 1 2 2 4 15 34
Thysanoptera - - - 3 42 183
Hemiptera 14 34 41 30 221 499
Homoptera 3 27 38 19 252 912
Megaloptera - - - 2 3 6
Neuroptera 3 4 6 5 18 54
Coleoptera 39 107 192 80 947 3,690
Strepsiptera - - - 4 5 17
Mecoptera - - - 2 2 4
Trichoptera - - - 13 70 188
Lepidoptera 16 71 97 64 657 2,187
Diptera 27 68 102 75 1,132 5,199
Siphonaptera - - - 5 24 47
Hymenoptera 9 23 31 52 1,071 6,191
TOTALS 129 395 618 425 4,717 20,007

* From Kloet, G. S., and W. D. Hincks, 1945, A Check List a British Insects. 483 pp. Stockport, England.

hopefully, a fairly complete list of the insects which have been reported from the Islands and a partially annotated bibliography of Galápagos entomology. We have been encouraged by entomologists interested in the Galápagos to publish this material, even though we realize that it is incomplete and has some obvious deficiencies. Among the latter are the fact that we have passed no independent taxonomic judgement on the validity of published records and unless we were aware of subsequent revisions, or unless the specialists who have so kindly examined our list have independently evaluated the references, with few exceptions they appear in the form in which they were treated in the most recent Galápagos publication. Nevertheless, we hope that our provisional faunal list

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and provisional bibliography will be useful, and it is in this spirit that we offer it, whatever its shortcomings.

ACKNOWLEDGMENTS

This bibliography was prompted by the authors' participation in the Galápagos International Scientific Project, supported in part by Grant GE-2370 from the National Science Foundation and in part by the following: Belvedere Scientific Fund; California Academy of Sciences; California Maritime Academy: Charles Darwin Foundation; Government of Ecuador; Pacific Science Board, National Academy of Sciences; Shell Oil Company; the United States Army, Navy, and Air Force; the University of California Extension Division; and the Associates in Tropical Biogeography, University of California. Many specialists have reviewed sections and made valuable corrections and additions. Acknowledgment is made of this kind assistance in the introductory remarks under each order.

ECUADORIAN NAMES OF GALÁPAGOS ISLANDS WITH THEIR
ENGLISH SYNONYMS

Baltra = South Seymour
Chávez = Santa Cruz = Indefatigable
Española = Hood
Fernandian = Narborough
Floreana = Santa Maria = Charles
Genovesa = Tower
Isabela = Albemarle
Marchena = Bindloe
Pinzón = Duncan
Pinta = Abingdon
Rábida = Jervis
San Cristóbal = Chatham
Santa Cruz = Chávez = Indefatigable
Santa Fé = Barrington
Santa Maria = Floreana = Charles
Santiago = James

A BRIEF HISTORY OF ENTOMOLOGICAL EXPLORATION
IN THE GALáPAGOS ISLANDS

Charles Darwin collected the first specimens of insects in the Galápagos Islands for scientific purposes. A surprising number of others have followed him in the 130 years since the Beagle's visit in 1835. We realize that the following list is incomplete, but it is offered mainly as an aid in locating collections of Galápagos insects and in finding detailed accounts of entomological work, islands visited, collecting, dates, field notes, etc. In some cases the information

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presented was difficult to find, and we hope that the list may save others unnecessary searching of the literature.

The general pattern of presentation is: first a list of individual collectors, in chronological order, followed by a list of expeditions, also in chronological order. In each case, where known, the name of the collector (or expedition and vessel) is given, then the dates and islands visited. Usually the place of publication of entomological results is indicated by reference to the bibliography or by mention of the most important journal articles or book. The best general account for each expedition is also cited, as is the current location of Galápagos collections. The information presented is as complete as we could make it up to the time of the Galápagos International Scientific Project (January, 1964).

INDIVIDUAL COLLECTORS

HABEL, SIMEON—July 22, 1868 to January 1, 1869. New York. Hood, Floreana, Santa Cruz, Isabela, Bindloe, and Abingdon islands. General Account: Habel, in Salvin, O. 1876. On the Avifauna of the Galápagos Archipelago. Trans. Zool. Soc. London, vol. 9, pp. 447–510.

BAUR, GEORGE—June 9–September 6, 1891. Clark University, Worchester, Connecticut. Albemarle, Bindloe, Barrington, Charles, Chatham, Duncan, Hood, Indefatigable, James, Jervis, and Tower islands. Collected a dragonfly nymph reported by Asahina 1961, p. 1. General Account: 1892. Ein Besuch der Galápagos Inseln. Biologisches Centralblatt, vol. 12, pp. 221–250.

GERSCH, KARL—February-April, 1932. San Cristóbal and Santa Cruz islands. Publication: Campos, F. (1932).

WITTMER, H.—1934. Floreana Island. Publication: Richards (1941). Collection: Hancock Foundation, University of Southern California.

CONWAY, E. A.—1939. Floreana Island. Publication: Richards (1941). Collection: Hancock Foundation, University of Southern California.

VINTON, K. W.—July and August, 1948. Balboa High School. Canal Zone. Specimens: U. S. National Museum. Publication: Wheeler, M. R. (1961). Some Drosophilidae (Diptera) from the Galápagos Islands. Ann. Ent. Soc. Amer., vol. 54, pp. 750–751.

FÖRSTER, JUAN—March, April, May, 1959. San Cristóbal, Floreana, and Santa Cruz islands. Collection: Zool. Sammlgn. Bayerischen Staates, München. publication: Kernbach (1962).

SEKIGUCHI, KOICHI—December—January, 1959–1960. Floreana and San Cristóbal islands. Publication: Asahina (1961).

GALÁPAGOS EXPEDITIONS

Voyage of the Beagle. Charles Darwin. H.M.S. Beagle. September 16 to October 19, 1835. Chatham, Charles, Albemarle, and James islands. Re-

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sults published: Butler, et al. 1877; Walker, 1848–1873, etc. Insect collections: British Museum (Natural History). General account: Darwin, C. 1839. Narrative of the surveying voyages of His Majesty's ships Adventure and Beagle, between the years 1826 and 1836, describing their examination of the southern shores of South America, and the Beagle's circumnavigation of the globe. London, Henry Colburn. Vol. III. Journal and Remarks. 1832–1836. pp. xiv + 629, maps. Darwin, C. 1845. Journal of researches into the natural history and geology of the countries visited during the voyage of H.M.S. Beagle round the world, under the command of Capt. FitzRoy, R.N. 2d ed. London, John Murray, pp. viii + 519, 12 figs.

Kongliga Svenska Fregatten Eugenies Resa Omkring Jorden under Befäl af C. A. Virgin, Åren 1851–3. (Royal Swedish frigate Eugenie voyage around the world under the command of C. A. Virgin, years 1851–3). Eugenie. Dr. J. G. M. Kinberg. May 9, 1852 to May 20, 1852. Chatham, Indefatigable, Charles, James, Albemarle islands. Results published: Svenska Vetenskaps-academien, Stockholm. Insect collection: Naturhistoriska Riksmuseum, Stockholm. General account: Skogman, C. 1854–1855. Fregatten Eugenies Resa Omkring Jorden Åren 1851–1853, under befäl of C. A. Virgin. Stockholm, Adolf Bonnier, Förra Delen. pp. vi + 250; sednare delen, pp. v+224, col. pls. maps, and text figs.

"Agassiz Expedition." Alexander Agassiz. Steamer Hassler. June 10, 1873 to June 20, 1873. Charles Albemarle, James, Jervis, and Indefatigable islands. Insect collection: Museum of Comparative Zoology, Harvard. General account: Elizabeth Agassiz, "A cruise through the Galapagos." Atlantic Monthly, 1873 (May), pp. 579–584.

Voyage of the Peterel. Commander W. E. Cookson. H.M.S. Peterel. June 1875. Abingdon, Charles, and Albemarle islands. Results published: Proc. Zool. Soc. Lond. 1877. General account: Gunther, A. 1877. "Account of the zoological collection made during the visit of H.M.S. "Peterel" to the Galapagos Islands." Proc. Zool. Soc. Lond., pp. 64-91.

"Albatross" Expedition—1888. (U. S. Fish Commission). Leslie A. Lee, Charles H. Townsend. Steamer Albatross. April 4 to 16, 1888. Chatham, Hood, Gardner, Albemarle, James, Duncan, Indefatigable, and Abingdon islands. Results published: Proc. U. S. National Museum, vol. XII, 1889. Insect collections: U. S. National Museum. General account: Howard, L. O. 1890. Scientific results of explorations by the U. S. Fish Commission Steamer Albatross. No. V. Annotated catalogue of the insects collected in 1887-88. Proc. U. S. National Museum, vol. 12, pp. 185-207.

"Albatross" Expedition—1891. (U. S. Fish Commission). Alexander Agassiz. Steamer Albatross. March 28-April 4, 1891. Chatham, Charles, Duncan,

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James, Hood, and Albemarle islands. Results published: Bulletin Museum of Comparative Zoology, vol. 23, p. 25. General account: Agassiz, Alexander. 1892. Reports on the dredging operations off the west coast of Central America to the Galápagos to the west coast of Mexico, and in the Gulf of California in charge of Alexander Agassiz, carried on by the U. S. Fish Commission Steamer "Albatross," Lieut. Commander Z. L. Tanner, U.S.N., Commanding. II. General sketch of the expedition of the "Albatross," from February to May, 1891. Bull. Mus. Comp. Zool., 23:1-89, 22 pls. (insects, pp. 68-69). Scudder, S. H. 1893. No. 1. Reports on the dredging operations off the west coast of Central America to the Galápagos…VII. The Orthoptera of the Galápagos Islands. Bull. Mus. Comp. Zool., vol. 25. pp. 1-25, 3 pls.

Hopkins Stanford Galápagos Expedition, 1898-1899. R. E. Snodgrass and Edmund Heller. Schooner Julia E.Whalen. December 8, 1898 to June 23, 1899. Visited all large and many small islands. Results published: Proc. Washington Academy of Sciences, vols. 4 and 5. Insect collections: Stanford University; mostly destroyed. General account: Proc. Wash. Acad. Sci., vol. 3, pp. 363-364, 1901.

California Academy of Science Expedition. F. X. Williams. Schooner Academy. Sept. 24, 1905 to Sept. 25, 1906. Visited all the islands. Results: Proc. Calif. Acad. Sci. Insect collection: Calif. Acad. Sci., San Fransisco. General account: Slevin, J. R. 1931. Log of the schooner Academy, on a voyage of scientific research to the Galápagos Islands, 1905-1906. Occ. papers, Calif. Acad. Sci., no. 17, 162 pp.

Harrison Williams Galápagos Expedition. Wm. Beebe and Wm. Morton Wheeler. Yacht Noma. March 28, 1923, for 21 days (100 hours on land). San Cristóbal, Santa Cruz, Santiago, Isabela, and Tower islands. Results: Zoologica—New York Zoological Society and "Galapagos, World's End." Insect collections: American Museum of Natural History. General account: Beebe, Wm. 1923. Williams Galapagos Expedition, Zoologica, vol. 5, no. 1, pp. 1-20, figs. and maps.

St. George Expedition. C. L. Collenette, Cynthia Longfield, E. Cheesman. Steam Yacht St. George. July 24, 1924—arrived. James, Indefatigable, Eden, Charles, Albemarle, and Narborough islands. General account: Collenette, C. L. 1925. Sea Girt Jungles. London, pp. 72-116.

Arcturus Oceanographic Expedition, 1925. Wm. Beebe. Steam Yacht Arcturus. April 4 to April 28, 1925. South Seymour, Albemarle, Tower, and Hood islands. Results published: Zoologica, vols. 7 and 8. Beebe, Wm.

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1926. Zoologica, vol. 8, pp. 1–45. General account: Beebe, Wm. 1926. "Arcturus Adventure" with "Log of the Arcturus." Also Beebe, Wm. 1925. "Logging the Arcturus, a visit to the Galápagos." Bull. N. Y. Zool. Soc., no. 4, July, pp. 103–116; N. Y. Zool. Soc., vol. 28, pp. 119–129.

The Norwegian Zoological Expedition. 1925. Alf Wollebaek. Cutter Monsunen. August to December, 1925. Floreana, Chatham, and Santa Cruz islands. Results published: Med. Zool. Mus., Oslo. Insect collections: Zollogiske Museum, Oslo. General account: Mielche, Hakon, 1938. Let's see if the world is round. London, 330 pp.

Cornelius Crane Pacific Expedition(Field Museum of Natural History, Chicago). Karl P. Schmidt. Brigantine Yacht Illyria. January 6, 1929-January 16, 1929. Albemarle, Narborough, Indefatigable, S. Seymour, Eden, and Charles islands. Collections: Chicago Museum of Natural History. Results: Publ. Field Museum, Zool. Series under heading: Scientific results of the Crane Pacific Expedition. General account: Shurcliff, S. N. 1930. Jungle Islands. The "Illyria" in the South Seas, New York, G. P. Putnam's Sons pp. xv + 298, illus.

Gifford Pinchot Expedition. Dr. A. K. Fisher. Yacht Mary Pinchot. June and July, 1929 (5 weeks). Chatham, Charles, Albemarle, Tower, Indefatigable, Seymour, Duncan, Hood, Daphne Major, Barrington, and Narborough islands. Collections. U. S. National Museum. General account: Pinchot, G. 1929. "To the South Seas, the cruise of the Mary Pinchot to the Galápagos, the Marquesas, and the Tuamotu islands, and Tahiti. Philadelphia, The John C. Winston Co., xiii + 500 pp., 1 plate.

Vincent Astor Expedition. C. H. Townsend, Kermit Roosevelt, and Henry K. Svenson. Nourmahal. March 31 to April 15, 1930. Indefatigable, Albemarle, Tower, and Charles islands. General account: Townsend, C. H. 1930. The Astor Expedition to the Galápagos Islands. Bull. N. Y. Zoological Society, vol. 33, pp. 135–172.

Hancock Galápagos Expeditions. J. S. Garth. R/V Velero III. 1. December 31, 1931 to January 24, 1932. Chatham, Charles, Albemarle, James, Indefatigable, S. Seymour, and Tower islands. 2. January 24, 1933 to February 26, 1933. Hood, Charles, chatham, Barrington, Indefatigable, Albemarle, James, Duncan, S. Seymour, Abingdon, and Tower islands. 3. January 11 to January 31, 1934. Wenman, Albemarle, Narborough, Charles, Indefatigable, Chatham, S. Seymour, James, and Hood islands. 4. December 2, 1934 to December 19, 1934. Bindloe, Charles, Indefatigable, Albemarle, James, S. Seymour, Chatham, and Hood islands. 5. January 16 to January

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30, 1938. Tower, Chatham, S. Seymour, James, Albemarle, Charles, Indefatigable, and Hood islands. Results published: Richards, A. G., Jr. The noctuid moths of the Galapagos from the collections of the Allan Hancock Foundation. Report of the Allan Hancock Pacific Exped., vol. 5, pp. 233–251, 4 pls. General account: Fraser, C. McLean. 1943. Allan Hancock Pacific Exped., vol. 1, 445 pp., 115 charts, 128 pls.

Templeton Crocker Expedition of the California Academy of Sciences. Mr. Maurice Willows. Schooner Zaca. April 15 to June 16, 1932. Thirteen islands including all main islands. Results: Proc. Calif. Acad. Sci., series 4, vols. 21 and 22. Insect collections: California Academy of Sciences. General account: Crocker, Templeton. 1933. The Templeton Crocker Expedition of the California Academy of Sciences, 1932. No. 2. Introductory statement. Proc. Calif. Acad. Sci., ser. 4, vol. 21, no. 2, pp. 3–9, 1 pl.

Templeton Crocker Expedition to Polynesia in 1934–35. Templeton Crocker. Schooner Zaca. March 9–25, 1935. Albemarle, Tower, and Santa Cruz islands. Results: Proc. Calif. Acad. Sci., ser. 4, vols. 21 and 22. Insect collections: American Museum of Natural History and (Hemiptera) Calif. Acad. Sciences. General account: Van Duzee, E. P. 1937. No. 33. The Hemiptera of the Templeton Crocker Expedition to Polynesia in 1934–1935. Proc. Calif. Acad. Sci., ser. 4, vol. 22, pp. 111-126. See also: Chapin, J. P. 1936. Jour. Amer. Mus. Nat. Hist., vol. 37, pp. 31–38.

Darwin Memorial Expedition. Wolfgang Von Hagen. September 1935 to February 1936. Chatham, Santa Cruz, Barrington, Hood, Seymour, Duncan, Jervis, James, Abingdon, and Albemarle islands. Insect collections: Termites. American Museum of Natural History? General accounts: 1. Von Hagen, Victor Wolfgang, 1940. Ecuador the Unknown. Two and a half years' travels in the Republic of Ecuador and the Galápagos Islands. New York, Oxford Univ. Press, 296 pp., many plates. 2. Von Hagen, Victor Wolfgang, 1949. Ecuador and the Galápagos islands. Norman, Oklahoma, Univ. of Oklahoma Press, 290 pp., plates and maps.

Hans Hass Expedition. Irenäus Eibl-Eibesfeldt. Yacht Xarifa. November 1953 to January, 1954. Tower, Santa Cruz, Hood, Narborough, Albemarle, Barrington, San Cristóbal, and Floreana islands. Specimens: Senckenberg Museum. General account: I. Eibl-Eibesfeldt, 1961. Galápagos, The Noah's Ark of the Pacific (Transl. by A. H. Brodrick). New York, Doubleday and Company, Inc. 192 pp. 21 plates, 4 colored.

Deutschen Galápagos-Expedition. 1962/63. Eberhard Curio. December, 1962–1963. Santa Cruz. Isabela, Fernandina, Wenman, Abingdon, Tower and Hood. publication: Curio, E., 1964. Über das Nächtigen von Nocticanace galapagensis Curran. Zeit. für Tierpsychologie, Band 21, Heft 7, pp. 794–797.

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INSECTS RECORDED FROM THE GALÁPAGOS ISLANDS

Order COLLEMBOLA

Family ENTOMOBRYIDAE
Genus Lepidocyrtus
intermixtus Folsom, 1924, zoologica, vol. 5, p. 75, pl. 5, figs. 27-29. Baltra
Family NEANURIDAE
Genus Anurida
maritima Guerin, 1836, Icon. règne animal de Cuvier, vol. 7, no. 3, p. 11 (Achorutes). Floreana
Order THYSANURA
Principal references to Galápagos fauna: Banks (1901), Folsom (1924), Stach (1932), Paclt (1959).
Family LEPISMATIDAE
Genus Heterolepisma
insulare Banks, 1901, Proc. Washington Acad. Sci., vol. 3 p. 543, figs. 51-52 (Lepisma). intermedia Folsom, 1942, Zoologica, vol. 5, p. 67, pl. 3, figs. 1-8. Genovesa, Fernandina, Isabela, Santa Cruz, Baltra, Floreana
Genus Stylifera
galapagoensis Banks, 1901, Proc. Washington Acad. Sci., vol. 3, p. 541, figs. 47-50 (Lepisma). Santiago, Pinzón, Daphne Major, Santa Cruz, Baltra, Floreana, Española
Order DIPLURA
Family CAMPODEIDAE
Genus Lepidocampa
juradoi seclusa Condé, 1960, Bull. Mus. Nat. Hist. Paris, ser. 2, vol. 32, p. 172. Santa Cruz

Order ORTHOPTERA

The Principal references for the Galápagos Islands are Butler (1877), Scudder (1893), McNeill (1901), Snodgrass (1902), Hebard (1920), and Caudell (1932). The treatments of Sphingonotus and Schistocerca by Snodgrass(1902) are followed rather than those of Hebard (who synonymized most subspecies) because snodgrass was familiar with the insects in the field. Also, McNeill's (1901) treatment of Liparoselis is adopted in preference to Hebard (1920). The list was reviewed for overall classification but not for completeness of Galápagos references by Grant (personal communication).

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Family ACRIDIAE
Genus Closteridea
bauri Scudder, 1893, Bull. Mus. Comp. Zool., vol. 25, p. 9, figs. Isabela, San Cristóbal
Genus Sphingonotus
tetranesiotis Snodgrass, 1902, proc. Wash. Acad. Sci., vol. 4, p. 444 (synonym of fuscoirroratus Stål, fide Hebard, 1920).
subsp. tetranesiotis Snodgrass, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 444 (as var. charlesensis).
fuscoirrorata Stål. 1860, Freg. Eugenies Resa, Zool. 1, Ins. p. 345 (type locality first restricted to Puna I.,Ecuador, by Snodgrass).
Floreana
subsp. barringtonensis Snodgrass, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 445 (as variety) Santa Fé
subsp. hoodensis Snodgrass, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 446 (as variety) Espanñola
subsp. indefatigabilensis Snodgrass, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 447 (as variety) Santa Cruz, Seymour
trinesiotis Snodgrass, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 439 (synonym of juscoirroratus Stål, fide Hebard, 1920). subsp. trinesiotis snodgrass, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 439 (as var. chathamensis). San Cristóbal
subsp. indefatigabilensis Snodgrass, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 441 (as variety) Seymour, Santa Cruz
subsp. albermarlensis Snodgrass, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 443 (as variety) Isabela
Genus Schistocerca
intermedia Snodgrass, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 431 Pinó
literosa Walker, 1870, Cat. Derm. Salt. Brit. Mus.,vol. 4, p. 620.
subsp. literosa Walker, 1870, Cat. Derm. Salt. San Cristóbal

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Brit. Mus., vol. 4, p. 620. discoidalis Scudder, 1893, Bull. Mus. Comp. Zoo., vol. 25, p. 16 (as subspecies).
subsp. hyalina Scudder, 1893, Bull. Mus. Comp. Zoo., vol. 25, p. 16. Genovesa
subsp. punctata Scudder, 1893, Bull. Mus. Comp. Zool., vol. 25, p. 16. Española
melanocera Stål, 1861, Freg. Eugenies Resa, Zool. 1, Ins. p. 326 (Acridium) (all subspecies synonymized by Hebard, 1920). Genovesa
subsp. immaculata Snodgrass, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 223 (as variety) Santa Cruz, Baltra
subsp. lineata Snodgrass, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 423 (as variety) Isabela
subsp. melanocera Stål, 1861, Freg, Eugenies Resa, Zool. 1, Ins., p. 326. Floreana
subsp. minor Snodgrass, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 421 (as variety) Isabela
subsp. pallida Snodgrass, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 422 (as variety) Santa Fé
Genus Halmenus
eschatus Hebard, 1920, Proc. Calif. Acad. Sci., ser. 4, vol. 2, no. 2, p. 329. Wenman I.
robustus Scudder, 1893, Bull. Mus. Comp. Zool., vol. 25, p. 18.
subsp. choristopterus Snodgrass, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 450 (as species).
Floreana
subsp. cuspidatus Snodgrass, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 450 (as species). Isabela
subsp. robustus Scudder, 1893, Bull. Mus. Comp. Zool., vol. 25, p. 18. Santa Cruz, Santiago
Family TETTIGONIIDAE
Genus Anaulocomera
darwinii Scudder, 1893, Bull. Mus. Comp. Zool., vol. 25, p. 19, figs. San Cristóbal, Santa Cruz
Genus Liparoscelis
cooksonii Butler, 1877, Proc. Zool. Soc. London, P. Floreana, Isabela,

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87 (Agraecia). Genovesa
cooksoni ensifer McNeill, 1901, Proc. Wash. Acad. Sci., vol. 3, p. 498, fig. 39 (synon. fide Hebard, 1920). Hood, Santa Cruz
pallidus McNeill, 1901, Proc. Wash. Acad. Sci., vol. 3, p. 500, fig. 41 (synon. fide Hebard, 1920). Barrington
paludicola McNeill, 1901, Proc. Wash. Acad. Sci., vol. 3, p. 499, fig. 40 (synon. fide Hebard, 1920). Isabela
Genus Conocephalus
exitiosus McNeill, 1901, Proc. Wash. Acad. Sci., vol. 3, p. 501, fig. 42 (Xiphidium). Santa Cruz, Santiago
Genus Neoconocephalus
triops Linnaeus, 1758, Systema Naturae, Ed. 10, p. 430 (Gryllus-Tettigonia). San Cristóbal
insulanus Scudder (not Redtenbacher), 1893, Bull. Mus. Comp. Zool., vol. 25, p. 21, pl. III, figs. 2, 3 (Conocephalus).
insularum Karny, 1907, Revisio Conocephalidarum, p. 38 (Conocephalus).
Family GRYLLIDAE
Genus Cryptoptilum
erraticum Scudder, 1893, Bull. Mus. Comp. Zool., vol. 25, p. 23, pl. III, figs. 6, 7 (Cycloptilum). Floreana, Isabela, Hood, Abingdon
lepismoide McNeill, 1901, Proc. Wash. Acad. Sci., vol. 3, p. 505 (Cycloptilum). Isabela, Genovesa
Genus Hygronemobius
speculi McNeill, 1901, Proc. Wash. Acad. Sci., vol. 3, p. 503, figs, 43, 44 (Nemobius). Isabela
Genus Gryllus
assimilis Fabricius, 1775, Syst. Ent., p. 280 (Acheta). galapageius Scudder, 1893, Bull. Mus. Comp. Zool., vol. 25, p. 22, pl. III, fig. 8. Isabela, Floreana, San Cristóbal, Santa Cruz, Hood
Order BLATTARIA
Family BLATTIDAE
Genus Anisopygia
snodgrassii McNeill, 1901, Proc. Wash. Acad. Sci., vol. 3, p. 493, figs. 35–37 (Temnopteryx). Isabela
Genus Blattella
germanica Linnaeus, 1767, Systema Naturae, Ed. 12, p, 668 (Blatta). Hood

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Genus Symploce
lita Hebard, 1916, Trans. Am. Ent. Soc., vol. 42, p. 354, figs. San Cristóbal
Genus Leurolestes
pallidus Brunner, 1865, Nouv. Syst. Blatt., p. 286 (Nauphoeta). San Cristóbal
Genus Periplaneta
americana Linnaeus, 1758, Systema Naturae, ed. 10, pp. 1:424 (Blatta). Floreana, San Cristóbal
australasiae Fabricius, 1775, Syst. Ent., p. 271 (Blatta). San Cristóbal, Floreana
brunnea Burmeister, 1838, Handb. Ent. 2; Abt. 2, pt. 1, p. 503. San Cristóbal
Genus Pycnoscelus
surinamensis Linnaeus, 1767, Systema Naturae, Ed. 12, p. 687 (Blatta). Isabela, San Cristóbal, Floreana
Genus Nauphoeta
cinerea Olivier, 1789, Encycl. Method., Ins., vol. 4, p. 314 (Blatta).
circumvagans Scudder (not Burmeister), 1893, Bull. Mus. Comp. Zool., vol. 25, p. 7.
San Cristóbal, Floreana
Order MANTODEA
Family MANTIDAE
Genus Galapagia
solitaria Scudder, 1893, Bull. Mus. Comp. Zool., vol. 25, p. 8, figs. Isabela, Floreana, Seymour
Order DERMAPTERA
Family FORFICULIDAE
Genus Anisolabis
bormansi Scudder, 1893, Bull, Mus. Comp. Zool., vol. 25, p.5. San Cristóbal
Genus Euborellia
annulipes Lucas, 1847, Bull. Soc. Ent. France, ser. 2, vol. 5, p. lxxiv (Forficula). Isabela, San Cristóbal
Order ISOPTERA
Primary references for the Galápagos are Banks (1901) and Light (1935).
Family KALOTERMITIDAE
Genus Kalotermes
darwini Light, 1935, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 242, pl. 9, figs. 1, 2, 7, 8
(subg. Cryptotermes).
Santiago, Santa Cruz, Gardner, Floreana

[page] 126

galapagoensis Banks, 1901, Proc. Wash. Acad. Sci., vol.3. p. 544, figs. 55, 56 (Calotermes). Wenman, Genovesa
immigrans Snyder, 1922, proc. U. S. Nat. Mus., vol. 61, no. 20, pp. 2–4. (Kalotermes s. str.). Fernandina, Santiago, Rábida, Santa Cruz, Baltra, San Cristóbal, Isabela, Genovesa
fatulus Light, 1935, Proc. Calif, Acad. Sci., ser. 4, vol. 21, p. 245, pl. 9, figs. 4, 6
(subg. Cryptotermes).
Isabela, Santiago
pacificus Banks, 1901, Proc. Wash. Acad. Sci., vol. 3, p. 555, fig. 57 (Calotermes). Fernandina, Baltra, Isabela, Santiago
Family RHINOTERMITIDAE
Genus Heterotermes
orthognathus Light, 1933, Univ. Calif. Publ. Entom., vol. 6, p. 113. Santa Cruz
Order ODONATA
The principal references are Currie (1901), Calvert (1901–08), and Asahina (1961). The list was checked by Lieftinck (personal Communication).
Family COENAGRIONIDAE
Genus Anomalagrion
hastatum Say, 1839, Jour. Acad. Nat. Sci. Phila. delphia, vol. 8, p. 38 (Agrion). San Cristóbal, Isabela
Family AESHNIDAE
Genus Aeshna
galapagoensis Currie, 1901, Proc. Wash. Acad. Sci., vol. 3, p. 382, figs. 29, 30. San Cristóbal, Isabela
Genus Anax
amazili Burmeister, 1839, Handb. d. Ent., vol. 2, p. 841 (Aeschna). San Cristóbal, Floreana
Family LIBELLULIDAE
Genus Pantala
flavescens Fabricius, 1798, Ent, Syst. Suppl., p. 285 (Libellula). Santa Cruz, Baltra, Floreana
hymenaea Say, 1839, Jour. Acad. Nat. Sci. Philadelphia, vol. 8, p. 18 (Libellula) Floreana
Genus Tramea
cophysa darwini Kirby, 1889, Trans. Zool. Soc. London, vol. 12, p. 315, pl. 51, fig. 1. Isabela, Floreana, San Cristóbal, Española
Genus Cannacria
fumipennis Currie, 1901, Proc. Wash. Acad. Sci., vol. 3, p. 387, figs. 31–34 Isabela

[page] 127

Order MALLOPHAGA

The principal references to Galápagos species are Kellogg and Kuwana (1902), Kellogg (1906), and Ewing (1924). Dr. Theresa Clay has kindly reviewed the manuscript and checked or corrected many of the names after studying types and other specimens in the Kellogg collection. Mr. B. C. Nelson assisted Dr. Clay by locating Galápagos slides in the Kellogg collection at Berkeley and sending them to London. New synonymy and confirmed or corrected identifications by Dr. Clay are marked with an asterisk [*]. The synonymy of Actornithophilus milleri is from Timmermann (1954), and generic placement of Philopterus domesticus and P. insulicola is by Carriker (1957). Menopon rusticum, Kellogg, 1896 (not Giebel, 1874), is a Colpocephalum (Clay, in litt.). For interpretation of the species of Brueelia see Hopkins (1951). Synonymy of Nirmus curvilineatus Kellogg & Kuwana is given by Clay (1958), that of Lipeurus limitatus (Halipeurus) by Edwards (1961), and that of Docophorus peristictus by Ward (1953).

Suborder AMBLYCERA
Family MENOPONIDAE
Genus Actornithophilus
grandiceps Piaget, 1880, Pédiculines, p. 558, pl. 46, fig. 7 (Colpocephalum)* Santa Cruz
incisus Piaget, 1880, Pédiculines, p. 569, pl. 47, fig. 9 (Colpocephalum)
funebris Kellogg, 1906 (not Kellogg, 1896), Trans. Amer. Ent. Soc., vol. 32, p. 320 (Colpocephalum)*
milleri Kellogg & Kuwana, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 483, pl. 30, fig. 6 (Colpocephalum)*
Wolf, Marchean, Fernandina, Isabela, Daphne, Baltra
Genus Austromenopon
albemarlei Kelloggg & Kuwana, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 488, pl. 31, fig. 5 (Menopon) Isabela
becki Kellogg, 1906, Trans. Amer. Ent. Soc., vol. 32, p. 322 (Menopon)* Daphne
narboroughi Kellogg & Kuwana, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 485, pl. 31, fig. 2 (Menopon).
numerosum Kellogg, 1906 (not Kellogg, 1896), Trans. Amer. Ent. Soc., vol. 32, p. 321 (Menopon)*
Darwin, Marchena, Fernandina, Baltra
navigans Kellogg, 1896, Proc. Calif. Acad. Sci., ser. 2, vol. 6, p. 156, pl. 14, figs. 4, 5 (Menopon) Galápagos Islands

[page] 128

Genus Colpocephalum
flavescens Haan, 1829, Mem. Mus. Hist. Nat. Paris, vol. 18, p. 309 (Liotheum). Wolf, Santa Cruz
heterosoma Piaget, 1880, Pédiculines, p. 572, pl. 48, figs. 3, 4.* Santa Cruz
spineum Kellogg, 1899, Occ. Pap. Calif. Acad. Sci., no. 6, p. 38, pl. 4, fig. 1.* Wolf, Isabela
unciferum Kellogg, 1896, Proc. Calif. Acad. Sci., ser. 2, vol. 6, p. 140, pl. 12, figs. 1–3. Galápagos Islands
Genus Cuculiphilus
snodgrassi Kellogg & Kuwana, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 486, pl. 31. fig. 3 (Menopon).
galapagensis Kellogg & Kuwana, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 487, pl. 31, fig. 4 (Menopon).
San Cristóbal, Española
Genus Eidmanniella
aurifasciata Kellogg, 1899, Occ. Pap. Calif. Acad. Sci., vol. 6, p. 43, pl. 4, fig. 5 (Menopon).
fuscofasciatum Kellogg, 1906 (not Piaget, 1880), Trans. Amer. Ent. Soc., vol. 32, p. 321.*
Wolf, Marchena, Daphne, Baltra, Santa Cruz
singularis Kellogg & Kuwana, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 485, pl. 31, fig. 1 (Menopon).* Wolf
Genus Myrsidea
incerta Kellogg, 1896, Proc. Calif. Acad. Sci., ser. 2, vol. 6, p. 553, pl. 73, fig. 2 (Menopon). Wolf, Marchena, Fernandina, Isabela, Santiago, Santa Cruz, Santa Fé, San Cristóbal Española
Genus Trinoton
lituratum Burmeister, 1838, Handbuch Ent., vol. 2, p. 441 (Trinotum). Santa Cruz
querquedulae Linnaeus, 1758, Systema Naturae, Ed. 10, p. 612 (Pediculus).
luridum Kellogg, 1906 (not Burmeister, 1838), Trans. Amer. Ent. Soc., vol. 32, p. 320.*
Santa Cruz
Family RICINIDAE
Genus Ricinus
angulatus Kellogg, 1896, Proc. Calif. Acad. Sci., ser. 2, vol. 6, p. 515, pl. 70, fig. 5 (Physostomum). Fernandina

[page] 129

Suborder ISCHNOCERA
Family PHILOPTERIDAE
Genus Anatoecus
icteroides Nitzsch, 1818, Germar's Mag. Ent., vol. 3, p. 290 (Philopterus). Santa Cruz
Genus Brueelia
chelydensis Hopkins, 1951, Ann. Mag. Nat. Hist., ser. 12, vol. 4, p. 337.
galapagensis Kellogg & Kuwana, 1902, Proc. Wash. Acad. Sci., 4:474 (Nirmus vulgatus var.).
Fernandina, Isabela, San Cristóbal, Española
galapagensis Kellogg & Kuwana, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 464, pl. 28, fig. 4 (Docophorus).
galapagensis Kellogg & Kuwana, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 471, pl. 29, fig. 5 (Nirmus).
Fernandina, Isabela, San Cristóbal
interposita Kellogg, 1896, Occ. Pap. Calif. Acad. Sci., no. 6, p. 23, pl. 2, fig. 7 (Nirmus). Fernandina, Isabela
vulgata Kellogg, 1896, Occ. Pap. Calif. Acad. Sci., no. 6, p. 495, pl. 67, fig. 5 (Nirmus). Isabela
Genus Columbicola
?macrourae Wilson, 1941, Jour. Parasitol., vol. 27, no. 262, figs. 5–8 (Esthiopterum).
baculus Kellogg & Kuwana, 1902 (not Nitzsch, 1818), Proc. Wash. Acad. Sci., vol. 4, p. 478 (Lipeurus).*
Wolf, Marchena, Isabela
Genus Craspedorrhynchus
taurocephalus Kellogg, 1896, Proc. Calif. Acad. Sci., ser. 2, vol. 6, p. 65, fig. 1 (Docophorus). Baltra
Genus Degeeriella
regalis Giebel, 1866, Zeitschr. Ges. Naturwiss., vol. 28, p. 364 (Nirmus).
curvilineata Kellogg & Kuwana, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 470, pl. 29, fig. 4 (Nirmus).
fusca Kellogg, 1906 (not Denny, 1842), Trans. Amer. Ent. Soc., vol. 32, p. 318 (Nirmus).*
Fernandina, Isabela, Marchena

[page] 130

Genus Docophoroides
? brevis Dufour, 1835, Ann. Soc. Ent. France, vol. 4, p. 674, pl. 21, fig.3 (Philopterus). Fernandina
irroratae Timmerman, 1962, Zeitschr. f. Parisitenk., vol. 21, p. 435. Galápagos Islands
levequei Timmerman, 1963, Bonn. zool. Beitr., vol. 14, p. 158. Española
Genus Halipeurus
attenuatus Edwards, 1961, Jour. Parasitol., vol. 47, p. 148, figs. 3–7 M. nr. Galápagos Islands
driversus Kellogg, 1896, Proc. Calif, Acad. Sci., ser. 2, vol. 6, p. 123, pl. 8, figs. 3, 4 (Lipeurus).
limitatus Kellogg, 1906, Trans, Arner, Ent. Soc., vol. 32, p. 319 (Lipeurus).
Darwin, Wolf, Isabela
intermedius Edwards, 1961, Jour. Parasitol., vol. 47 p. 151, figs. 3–7 S.
major Kellogg & Kuwana, 1902 (not Piaget, 1880), Proc. Wash. Acad. Sci., vol. 4, p. 447 (Lipeurus diversus var.).
Galápagos Islands
pelagicus Denny, 1842, Mon. Anopl. Brit., vol. 58, p. 173, pl. 14, fig. 2 (Lipeurus)
exiguus Kellogg & Kuwana, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 479, pl. 30, fig. 2 (Lipeurus).
languidus Kellogg & Kuwana, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 475, pl. 28, fig. 8 (Lipeurus).
Darwin, Wolf, Isabela
Genus Pectinopygus
annulatus Piaget, 1880, Pédiculines: p. 340, pl. 27, fig. 10 (Lipeurus).
fuscomarginatus, Kellogg, 1906 (not Denny 1842), Trans. Amer. Ent. Soc., vol. 32, p. 318 (Nirmus).*
potens Kellogg & Kuwana, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 477, pl. 30, fig. 1 (Lipeurus).
Darwin, Wolf, Isabela, Daphne,Baltra, santa Cruz
fregatiphagus Eichler, 1843, Zool. Anz. (Epifrega).*
major Kellogg, 1899, Occ. Pap. Calif. Acad. Sci., no. 6, p. 30, pl. 3, fig. 3 (Lipeurus gracilicorne Var.).
Wolf

[page] 131

nannopteri Ewing, 1924, Zoologica, vol. 5, p. 82, fig. 12 (Esthiopterum).* Galápagos Islands
sulae Rudow, 1869, Beitr. Kenntn. Malloph., p. 43 (Lipeurus).
helleri Kellogg & Kuwana, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 479, pl. 30, fig. 3 (Lipeurus).*
Darwin, Wolf, Marchena, Isabela
Genus Perineus
miriceps Kellogg & Kuwana, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 480, pl. 30, fig. 4 (Lipeurus). Isabela
oblongus Kéler, 1957, Beitr. Ent., vol. 7, p. 523, figs. 27–28. Galápagos Islands
Genus Phioceanus
becki Kellogg, 1903, Biol. Bull. Woods Hole, vol. 5, Wolf p. 88, figs. 1,2.* Wolf
Genus Philopterus
breviformis Kellogg & Kuwana, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 463, pl. 28, fig. 3 (Docophorus).* Isabela, santa Cruz
communis Nitzsch, 1818, Germar's Mag. Ent., vol. 3, p. 290 Fernandina, Española
domesticus Kellogg, 1896, Proc. Calif. acad. Sci., ser. 2, vol. 6, p. 475, pl. 65, fig. 4 (Docophorus). Santa Cruz
insulicoal Kellogg & Kuwana, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 28, fig. 6 (Docophorus) Isabela
Genus Physconelloides
galapagensis Kellogg & Kuwana, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 481, pl. 30, fig. 5 (Goniocotes).* Wolf, Marchena, Fernandia, Isabela
Genus Quadraceps
birostris Giebel, 1874, Insecta Epizoa, p. 174 (Nirmus).
gloriosa Kellogg & Kuwana, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 467 (Nirmus).*
Isabela, Baltra, Santa Fé
paludicola Kellogg & Kuwana, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 469, pl. 29, fig. 3 (Nirmus).* Isabela
ridgewayi Kellogg, 1906, Trans. Amer. Ent. Soc., vol. 32, p. 317 (Nirmus).* Santa Cruz
separatus Kellogg & Kuwana, 1902, Trans. Wash. Acad. Sci., vol. 4, p. 472, pl. 29, fig. 6 (Nirmus).* Wolf, Marchena, Isabela, Española

[page] 132

strepsilaris Denny, 1842, Mon. Anopl. Brit., pp. 52, 135, pl. 11, fig. 4 (Nirmus).
lepidus Kellogg & Kuwana, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 473, pl. 29, fig. 7 (Nirmus).*
Fernandina, Santa Fé
Genus Saemundssonia
albemarlensis Kellogg & Kuwana, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 465, pl. 28, fig. 5 (Docophorus).
melanocephalus Kellogg & Kuwana, 1902 (not Burmeister, 1838), Proc. Wash. Acad. Sci., vol. 4, p. 462 (Docophorus).*
Darwin, Wolf, Fernandina, Isabela, Baltra, Gardner
lari O. Fabricius, 1780, Fauna Groenlandia, p. 219 (Pediculus).* Darwin
minor Kellogg & Kuwana, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 460 (Docophorus validus var.). Darwin, Fernandina, Isabela, Santa Fé
platygaster Denny, 1842, Mon. Anopl. Brit., pp. 43, 83, pl. 2, fig. 5 (Docophorus).
peristicta Kellogg & Kuwana, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 462, pl. 28, fig. 2 (Docophorus).
Darwin, Isabela, Santa Fé
phaetona Osborn 1890, Proc. U. S. Nat. Mus., vol. 12, p. 189 (Docophorus).* Wolf, Daphne, Santa Cruz
platycephalus Kellogg & Kuwana, 1902, Proc. Wash. Acad. Sci., vol. 4, p. 461, pl. 8, fig. 1 (Docophorus).*
Genus Trabeculus
mirabilis Kellogg, 1896, Proc. Calif. Acad. Sci., ser. 2, vol. 6, p. 138, pl. 11, fig. 7, 8 (Giebelia).* Wolf, Isabela
Order ANOPLURA
Family HAEMATOPINIDAE
Genus Hoplopleura
nesoryzomydis Ferris, 1921, Stanford Univ. Publ., Biol. Ser., vol. 2, no. 2, p. 90.
quadridentata Ferris, 1916 (not Neumann), Psyche, vol. 23, p. 116.
Galápagos Islands
Genus Polyplax
spinulosa Burmeister, 1839, "Rhynchota" Gen. Ins., p. 8 (Pediculus). Floreana

[page] 133

Order HEMIPTERA

The principal references to Galápagos Hemiptera-Heteroptera are by Stål (1859), Butler (1877), Heidemann (1901), Barber (1925, 1934), and Van Duzee (1933, 1937). Changes since Barber's comprehensive list (1934) are Dallasiellus (Froeschner, 1960), Dicranocedhalus (Scudder, 1957), Liorhyssus and Rhopalus (Harris, 1943, Iowa State College Jour. Sci., vol. 17, pp. 197–204), pentacora sphacelata (Drake, 1954), Trigonotylus (Carvalho and Wagner, 1957), Polymerus nigritulus and Cyrtopeltis modesta (Carvalho, 1957–1960), and Halobates (Herrring, 1961). Trichocorixa (Hutchinson, 1931) was not identified to species. It is recorded as Trichocorixa reticulata Guerin (Usinger).

Family CYDNIDAE
Genus Dallasiellus
murinus Van Duzee, 1933, Proc, Calif. Acad, Sci., ser. 4,vol. 21,p. 26 (Geotomus). Isabela, San Cristóbal
Family PENTATOMIDAE
Genus Acrosternum
viridans Stål, 1859, Fregatten Eugenies Resa, Zool., vol. 1, p. 228 (Rhaphigaster). San Cristóbal, Isabela, Santa Cruz, Santiago
Genus Thyanta
setigera Ruckes, 1957, Pan-Pac. Ent., vol. 33, p. 179. perditor Heidemann, 1901 (not Fabricius, 1974), Proc, Wash, Acad. Sci., vol. 3,p. 365. Santa Cruz, Isabela, San Cristóbal, Floreana, Jervis, Gardner, James, N. Seymour
similis Van Duzee, 1933, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 26. Rábida, Floreana
Genus Podisus
sordidus Stål, 1859, Fregatten Eugenies Resa, Zool., vol. 1.p. 221 (Arma). Santa Cruz, Isabela, San Cristóbal, Floreana
Genus Trincavellius
galapagoensis Butler, 1877, Proc. Zool. Soc. London, 1877, p. 88 (Sciocoris). Floreana
GenusAlcaeorrhyncus
grandis Dallas, 1851, List Hemipt. Brit. Mus., vol. 1, p. 91 (Canthecona). Floreana
Family COREIDAE
Genus Anasa
obscura Dallas, 1852, List Hemipt. Brit. Mus., vol.2, p. 505. San Cristóbal
GenusDicranocephalus
insularis Dallas, 1852, List Hemipt. Brit. Mus., vol. Santa Cruz, Santiago,

[page] 134

2, p. 482 (Stenocephalus). Isabela, Santiago, Santa Cruz
Genus Harmostes
disjunctus Barber, 1925, Zoologica, vol, 5, p. 241. serratus Heidemann (not Fabricius), 1901, Proc. Wash. Acad, Sci. vol. 3,p. 365.
Santa Cruz, Isabela
Genus Liorhyssus
hyalinus Fabricius, 1794, Ent. Syst., vol. 4, p. 168 (Lygaeus). Genovesa, Daphne Major
Genus Rhopalus
lugens Signoret, 1859. Ann. Soc. Ent. France, ser. 3, vol. 7, p. 92 (Corizus). Galápagos
Genus Jadera
sanguinolenta Fabricius, 1775, Syst. Ent., p. 721 (Cimex) Wenman
Family NEIDIDAE
Genus Aknisus
galapagensis Barber, 1934, Meded. Zool. Mus., Oslo, vol. 42, p. 284.
tenellus Heidemann, 1901, Proc. Wash. Acad. Sci vol. 3, p. 366 (not Stål) (Jalysus).
San Cristóbal
Family LYGAEIDAE
Genus Nysius
naso Van Duzee, 1933, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 27 (Nysuis-Ortholomus). Floreana
marginalisDallas, 1852, List Hemipt. Brit. Mus., vol. 2, p. 556 (? Nysius).
galapagensis Stål, 1860, Freg. Eugenies Resa, Ins., p.252 (Cymus).
Floreana, Santiago, Daphne Major
Genus Heraeus
pacificusBarber, 1925, Zoologica, vol. 5, p. 245. Santiago
GenusOrthaea
insularisBarber, 1925, Zoologica, vol. 5, p.246. Baltra, Santiago,Isabela, Santa Cruz
Family PYRRHOCORIDAE
Genus Dysdercus
concinnus Stål, 1861, Öfv. Vet. Akad. Förh., p. 198. Santa Cruz
Family TINGIDAE
Genus Corythaica
cytharina Butler, 1877, Proc. Zool. Soc. Lond., 1877, Santiago, Daphne

[page] 135

p. 90 (Monanthia).
renormata Barber, 1925, Zoologica, vol. 5, p. 251.
Major
Family REDUVIIDAE
Genus Ghilianella
galapagensis Heidemann, 1901, Proc. Wash. Acad. Sci., vol, 3, p. 367. Isabela, Española
Genus Repipta
annulipes Barber, 1925, Zoologica, vol.5, p. 250. species? Heidemann, 1901. Proc. Wash. Acad. Sci., vol. 3, p. 366 (nymph) (Cosmoclopius-Harpactor). Santa Cruz, Isabela
Family NABIDAE
Genus Nabis
punctipennis Blanchard, 1852, Gay, Hist. de Chili, Zool., vol. 7, p.161. Isabela, Floreana, Santiago
Family CIMICIDAE
Genus Cimex
lectularius Linnaeus, 1758, Syst, Nat., 10th ed., p. 441. Floreana
Family MIRIDAE
Genus Trigonotylus
lineatus Butler, 1877, Proc. Zool. Soc. Lond., 1877, p. 89 (Miris). Isabela, Floreana, Seymour
Genus Psallus
insularis Barber, 1925, Zoologica, vol. 5, p. 250. Santiago
Genus Fulvius
geniculatus Van Duzee, 1933, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 29. Rábida, San Cristóbal, Santiago, Fernandina, Floreana
Genus Diaphnidia
crockeri Van Duzee, 1933, Proc. Calif. Acad. Sci., ser. 4, vol. 21,p. 29. Santiago
Genus Dagbertus
darwini Butler, 1877, Proc. Zool. Soc. Lond., 1877, p. 89 (Capsus). Floreana
quadrinotatus Walker, 1873, Cat. Hem. Heter. Brit. Mus., vol. 6, p. 113 (Capsus). Floreana, Santiago
spoliatus Walker, 1873, Cat. Hem. Heter. Brit. Mus., vol. 6, p. 112 (Capsus). Floreana, Santiago

[page] 136

Genus Polymerus
nigritulus Walker, 1873, Cat. Hem. Heter. Brit. Mus., vol. 6, p. 112 (Capsus). Floreana
vegatusVan Duzee, 1933, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 28 (Poeciloscytus). Isabela
Genus Creontiades
castaneumVan Duzee, 1933, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 27. San Cristóbal
fucosus Barber, 1925, Zoologica, vol. 5, p. 248. Santiago, Santa Cruz
willowsiVan Duzee, 1933, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 28. Santiago, Santa Cruz, Rábida
Genus Cyrtopeltis
modesta Distant, 1893, Biol. Centr.-Amer. Rhynch., vol. 1, p. 447 (Neosilia).
geniculatus Reuter, 1876, Öfv. K. Vet. Akad. Förh., vol. 36, no. 2, p. 83 (Engytatus).
Santa Cruz
Genus Europiella
mella Van Duzee, 1937, Proc. Calif. Acad. Sci., ser. 4, vol. 22, p. 117. Santa Cruz
Family SALDIDAE
Genus Pentacora
sphacelata Uhler, 1877, Bull. U. S. Geol. Geogr. Survey, vol. 3, p. 434 (Salda).
rubromaculata Heidemann, 1901, Proc. Wash. Acad. Sci., vol. 3, p. 368 (Salda).
Isabela
Family GERRIDAE
Genus Halobates
micans Eschscholtz, 1822, Entomographien, p. 107 wüllerstorffi Frauenfeld, 1867, Verh. K. K. Zool. Bot. Ges. Wien, vol. 17, p. 458. near Santiago, Galápagos
robustus Barber, 1925, Zoologica, vol. 5, p. 253. Santa Cruz, Floreana, Isabela
sobrinus White, 1883, Voy. Challenger, Rpt. Zool., vol. 7, no. 19, p. 46. near Galápagos
Family CORIXIDAE
Genus Trichocorixa
reticulata Guerin, 1857, in Sagra, Hist. Cuba, Ins., p. 423 (Corisa). Galápagos

Order HOMOPTERA

The leading publications on Galápagos Homoptera are Walker (1851), Stål

[page] 137

(1859), Butler (1877), Morrison (1924), Osborn (1924), and Van Duzee (1933, 1937). Reclassification of the Walker and Butler species way by Muir (1919) and W. E. China (in Champion, 1924). The most recent placement of Fulgoridae is given in Metcalf (1943, 1954, 1958). Fennah (personal communication) placed Issus varius Walker in Philatis. Linnavuori (1959) is the authority for generic placement of Deltocephalus, Scaphytopius, Amplicephalus, and Exitianus. W. E. China (personal communication) corrected the name Jassus of Van Duzee and Osborn. D. A. Young (personal communication) examined the Van Duzee types of Cicadellidae and his generic assignments have been followed. The modern classification of Coccidae is after Ferris (Atlas of Scale Insects of North America, Stanford University). H. L. McKenzie kindly reviewed the manuscript on Coccidae.

Family FULGORIDAE
Sub-family CIXIINAE
Genus Nymphocixia
unipunctata Van Duzee, 1923, Proc. Calif. Acad. Sci., ser. 4, vol. 12, p. 189. Isabela, Santa Cruz
Genus Oliarus
galapagensis Van Duzee, 1933, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 33. Isabela, Santa Cruz
Sub-family ISSINAE
Genusphilatis
brevicepsVan Duzee, 1933, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 33. Floreana
cinerea Osborn,1924, Zoologica, vol. 5, p. 78 Genovesa
major Osborn,1924, Zoologica, vol. 5, p. 79. Santa Cruz
producta stâl, 1859, Fregatten Eugenies Resa, Ins., p. 278 (Mycterodus Isabela, Seymour, Genovesa, Santa Cruz
serva Van Duzee, 1933, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 34. Isabela, ? Santiago
variaWalker, 1851, List Homopt. Brit. Mus., vol. 2, p. 372 (Issus). New combination. Santiago
vicina Van Duzee, 1933, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 34. Rábida
Genus Euthiscia
crockeri Van Duzee, 1937, Proc. Calif. Acad. Sci., ser. 4,vol. 22, p. 119. Santa Cruz
Genus Galapagosana
rostrifer Butler, 1877, Proc, Zool. Soc. Lond., 1877, p. 90 (Issus).

[page] 138

Sub-family DELPHACINAE
Genus Delphacodes
substitua Walker, 1851, List Homopt. Brit. Mus., vol. 2, p. 354 (Delphax). Floreana, Santiago
vicaria Walker, 1851, List Homopt. Brit. Mus., vol. 2, p. 355 (Delphax). Floreana, Santiago
Genus Ilburnia
simulans Walker, 1851, List Homopt. Brit. Mus., vol. 2, p. 355 (Delphax). Santigo, Floreana
Family CICADELLIDAE
Genus Deltocephalus
insularis Van Duzee, 1933, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 30 (possibly an Amplicephalus Linnavuori, 1959). San Cristóbal, Floreana
Genus Scaphytopius
acquinoctialis Van Duzee, 1933, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 30 (Platymetopius). Isabela, ? Santiago
retusus Van Duzee, 1937, Proc. Calif. Acad. Sci., ser. 4, vol. 22, p. 123 (Platymetopius). Santa Cruz
Genus Sanctonus
discalis Van Duzee, 1933, Proc. Calif Acad. Sci., ser. 4, vol. 21, p. 31 (Scaphoideus).
Genus Exitianus
digressus Van Duzee, 1933, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 32 (Athysanus). San Cristóbal, Santiago, Seymour, Rábida
Genus Agalliopsis
mera Van Duzee, 1937, Proc. Calif. Acad. Sci., ser. 4, vol. 22, p. 123. Santa Cruz
Genus Agallia
planus Butler, 1877, Proc. Zool. Soc. Lond., 1877, p. 91 (Jassus). Floreana
striolaris Butler, 1877, Proc. Zool. Soc. Lond., 1877, p. 91. (Jassus). Floreana
Genus Coelidia
infesta Van Duzee, 1937, Proc. Calif. Acad. Sci., ser. 4, vol. 22, p. 124 (Jassus). Santa Cruz
galapagoensis Osborn, 1924, Zoologica, vol. 5, no. 6, p. 77 (Jassus). Santiago, Santa Cruz
Genus Mesamia
obliquus Walker, 1851, List Homopt. Brit. Mus., vol. 3, p. 851 (Acrocephalus).

[page] 139

Genus Balclutha
lucidus Butler, 1877, Proc. Zool. Soc. Lond., 1877, p. 91 (Jassus). Santiago
Family COCCIDAE
Genus Margarodes
similis Morrison, 1924, Zoologica, vol. 5, p. 143, fig. 33. Santa Cruz, Eden, Baltra
Genus Orthezia
galapagoensis Kuwana, 1902, Jour, New York Ent. Soc., vol. 10, p. 28 Isabela, Santa Cruz, Duncan
Genus Eriococcus
papillosus Morrison, 1924, Zoologica, vol. 5, p. 145. Santa Cruz
Genus Asterolecanium
pustulans Cockerell, 1892. Jour. Inst. Jamaica, vol. 1, p. 143 (Asterodiaspis). Isabela
Genus Saissetia
hemisphaerica Targioni-Tozzetti, 1867. Studii sul. Cocc., vol. 11, p. 26 (Lecanium). Isabela, San Cristóbal
Genus Coccus
hesperidum var. pacificum Kuwana, 1902, Jour. New York Ent. Soc., vol. 10, p. 30(Lecanium). Abingdon, Isabela, Baltra
Genus Melanaspis
obtusa Ferris, 1941, Atlas of Scale Ins. N. America, S III, p. 362.
smilacis Kuwana, 1902. Jour. N. Y. Ent. Soc., vol. 10, p. 32 (not Comstock).
Isabela
Genus Hemiberlesia
lataniae Signoret, 1869, Ann. Soc. Ent. France, ser. 4, vol. 9, p. 124 (Aspidiotus). Abingdon
Genus Phenacoccus
parvus Morrison, 1924, Zoologica, vol. 5, p. 147. Tower
Genus Pseudococcus
galapagoensis Morrison, 1924, Zoologica, vol. 5, p. 148. Santa Cruz. (Eden)
insularis Morrison, 1924, Zoologica, vol. 5, p. 150. Baltra
Genus Ceroplastes
sp. Morrison, 1924, Zoologica, vol. 5, p. 152. Santa Cruz, (Eden), Baltra
Order NEUROPTERA
The principal references are Banks (1924) and Esben-Petersen (1934).

[page] 140

The list was checked by D. E. Kimmins (personal communication).

Family MYRMELEONIDAE
Genus Myrmeleon
perpilosus Banks, 1924, Zoologica, vol. 5, p. 177. sp., Esben-Peterson, 1934, Nyt Mag. f. Naturvidensk., vol. 74, p. 293, fig. 3 (larva). Santa Cruz Floreana
Genus Dimares
formosus Banks, 1908, Proc. Ent. Soc. Washington, vol. 9, p. 31.
nummatus Navas, 1912, Ann. Soc. Sci. Bruxelles, vol. 36, p. 229, fig. 16.
S. Elena
Family CHRYSOPIDAE
Genus Chrysopa
nigripilosa Banks, 1924, Zoologica, vol. 5, p. 177. Baltra
galapgoensis Banks, 1924, Zoologica, vol. 5, p. 179. Baltra
wollbaeki Esben-Petersen, 1934, Nyt Mag. f. Naturvidensk., vol. 74, p. 291, figs. 1–2. Floreana
Family HEMEROBIIDAE.
Genus Megalomus
darwini Banks, 1924, Zoologica, vol. 5, p. 179. Santa Cruz

Order COLEOPTERA

The major contributors to our knowledge of the Galápagos Coleoptera have been G. R. Waterhouse (1845), Boheman (1859), C. Waterhouse (1877), Mutchler (1925, 1938), Blair (1928, 1933), and Van Dyke (1953). The present summary is based largely on Van Dyke, supplemented by descriptions or taxonomic treatment of Galápagos beetles appearing since 1953. Three of the most doubtful of the records reported by Blair (1933), Doryphora guerina Stål, Diabrotica ventricosa Jacoby, and Physonota alutacea Boheman, were excluded by Van Dyke and we have not restored them to the list. However, other However, other dubious records remain and these are indicated with a question mark in front of the recorded Galápagos locality. R. F. Smith (personal communication) examined the type of Diabrotica limbata Waterhouse and found that it belongs to the genus Acalymma. This altered generic assignment is indicated below and the new combination should be credited to him.

Family CICINDELIDAE
Genus Cicindela
galapagoensis W. Horn, 1915, Genera Insectorum, Fasc. 82, p. 402. Isabela
vonhageni Mutchler, 1938, Amer. Mus. Nov., no. 981, p. 2, p1. 1, fig. 1. Santa Cruz

[page break]

Family CARABIDAE
Genus Calosoma
darwiniana Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 10, p1. 1, f. 3. Isabela
galapageium Hope, 1838, Trans. Ent. Soc. London, vol. 2, p. 130. Santiago
howardi Linell, 1898, Proc. U. S. Nat. Mus., vol. 21, p. 251.
galapagoum Howard (not Hope), 1889, Proc. U. S. Nat. Mus., vol. 12, p. 191.
galapageiumBreuning (not Hope), 1927, Koleopt. Rundschau, vol. 13, p. 140.
Darwin, Isabela, santiago, Pinzón, Santa Cruz, Baltra, San Cristóbal, Floreana
linelli Mutchler, 1925, Zoologica, vol. 5, p. 221.
galapageium Linell (not Hope), 1898, Proc. U. S. Nat. Mus., vol. 21, p. 250.
San Cristóbal
GenusScarites
galapagoensis Linell, 1898, Proc. U. S. Nat. Mus., vol. 21, p. 253. San Cristóbal
williamsi Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., vol. 22, p. 13, pl. 2, fig. 3. Isabela
Genus Bembidion
equatoriale Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 15. San Cristóbal
galapagoensis G. R. Waterhouse, 1845, Ann. Nat. Hist., vol. 16, p. 23 (Notaphus). Pinta, Santiago
Genus Tachys
beebei Mutchler, 1925, Zoologica, vol. 5, p. 223, fig. 42. Baltra
Genus Feronia
becki Van Dyke, 1953, Occ. Pap. Calif. Acad. sci., no. 22, p. 24. santiago
blairi Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 19. Santiago
calathoides G. R. Waterhouse, 1845, Ann. Nat. Hist., vol. 16, p. 21. San Cristóbal
duncani Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 21, pl. 2, fig. 4. Pinzón
galapagoensis G. R. Waterhouse, 1845, Ann. Nat. Hist., vol. 16, p. 21. Santiago
insularis Boheman, 1858, Fregatten Eugenies Resa, vol. 1, p. 14. Isabela

[page] 142

mutchleri Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 20. Isabela
waterhousei Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 17. Floreana
williamsi Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 21. Santa Cruz
Genus Agonum
albemarli Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 27. Isabela
chathami Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 26, pl. 2, fig. 9. San Cristóbal
darwini Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 25, pl. 2, figs. 7, 8. San Cristóbal
Genus Selenophorus
galapagoensis G. R. Waterhouse, 1845, Ann. Nat. Hist., vol. 16, p. 22. San Cristóbal, Floreana, Española, Gardner
obscuricornis G. R. Waterhouse, 1845, Ann. Nat. Hist., vol. 16, p. 22 (Amblygnathus). Isabela
Wenmani Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., vol. 22, p. 29. Wolf
Family DYTISCIDAE Genus Copelatus galapagoensis G. R. Waterhouse, 1845, Ann. Nat. Hist., vol. 16, p. 23. Isabela, San Cristóbal
Genus Rhantus
sp. (signatus Fabricius?) Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 32. San Cristóbal
Genus Thermonectes
basillaris galapagoensis Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 33.
incisus var. C. Waterhouse, 1877, Proc. Zool. Soc. London, vol. 5, p. 79, 82 (Acilius).
basillaris Mutchler (not Harris), 1925, Zoologica, vol. 5, p. 225.
Isabela, Baltra, San Cristóbal, Floreana Genus Eretes sticticus Linnaeus, 1767, Systema Naturae, ed. 12, p. 666 (Dytiscus).
occidentalis C. Waterhouse (not Erickson), 1877, Proc. Zool. Soc. London, vol. 5, p. 77(Eunectes).
Santa Cruz, San Cristóbal, Floreana

[page] 143

Family GYRINIDAE
Genus Gyrinus
galapagoensis Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 34. San Cristóbal
Family HYDROPHILIDAE
Genus Ochthebius
batesoni Blair, 1933, Ann. Mag. Nat. Hist., ser. 10, vol. 11, p. 473. San Cristóbal?
Genus Tropisternus
lateralis Fabricius, 1775, Systema Entomologia, p. 28 (Hydrophilus). Santiago, San Cristóbal, Floreana
Genus Enochrus
obscurus Sharp, 1882, Biologia Centr-Americana, Coleoptera, vol. 1, no. 2, p. 69 (Philhydrus). San Cristóbal, Floreana
waterhousei Blair, 1933, Ann. Mag. Nat. Hist., ser. 10, vol. 11, p. 473. ? San Cristóbal
Genus Galapagodacnum
darwini Blair, 1933, Ann. Mag. Nat. Hist., ser. 10, vol. 11, p. 474 (Coelostoma). Pinta, Brattle, Genovesa
Family STAPHYLINIDAE
Genus Bledius
aequatorialis Mutchler, 1925, Zoologica, vol. 5, p. 225, fig. 43. Baltra
Genus Criophilus
villosus Gravenhorst, 1802, Coleopt. Microptera Brunsvicensis, p. 160. Isabela, San Cristóbal
Family HISTERIDAE
Genus Carcinops
galapagoensis Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 38. Pinta, Genovesa, Santa Cruz
tenellus Erickson, 1834, in King, Jahrb. Ins., vol. 1, p. 170 (Paramalus). Isabela, Santiago
Genus Saprinus
batesoni Blair, 1933, Ann. Mag. Nat. Hist., ser. 10, vol. 11, p. 475. Eden, ? San Cristóbal
modestior Marseul, 1845, Mon., p. 493, pl. 19, fig. 110. Pinta

[page] 144

Family LYCIDAE
Genus Calocladon
testaccum Gorham, 1881, Biologia Centr.-Americana, Coleoptera, vol. 3, no. 2, p. 28, pl. 2, fig. 20. ? San Cristóbal
Family CANTHARIDAE
Genus Chauliognathus
sulphureus C. Waterhouse, 1878, Trans. Ent. Soc. London, 1878, p. 331. San Cristóbal
Family MELYRIDAE
Genus Ablechrus
flavipes C. Waterhouse, 1877, Proc. Zool. Soc. London, p. 79.
darwinii C. Waterhouse, 1877, Proc. Zool. Soc. London, p. 81.
San cristóbal
Family CLERIDAE
Genus Pelonium
longfieldae Blair, 1928, Ann. Mag. Nat. Hist., ser. 10, vol. 1, p. 677. Isabela, Santiago, San Cristóbal, Floreana
Genus Necrobia
rufipes De Geer, 1775, Mem., vol. 5, p. 165, pl. 15, fig. 4. Cosmopolitan
Family OEDEMERIDAE
Genus Oxacis
galapagoensis Linell, 1898, Proc. U. S. Nat. Mus., vol. 21, p. 266. Isabela, Santiago, San Cristóbal, Floreana, Española
pilosa Champion, 1890, Biologia Centr-Amer., Coleoptera, vol. 4, no. 2, p. 156, pl. 7, fig. 15. Santiago
Genus Alloxacis
collenettei Blair, 1928, Ann. Mag. Nat. Hist., ser. 10, vol. 1, p. 673. Isabela, Santiago, Santa Cruz, Baltra
hoodi Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 43. Española
seymourensis Mutchler, 1925, Zoologica, vol. 5, p. 226. Baltra
Family MORDELLIDAE
Genus Mordellistena
galapagoensis Van Dyke, 1953, Occ. Pap. Calif Acad. Sci., no. 22, p. 44. Genovesa

[page] 145

Family MELOIDAE
Genus Cissites
maculata Swederus, 1787, Svenska Vetensk. Avhandl. Natursk., p. 199, pl. 8, fig. 8 (Cucujus). Santa Cruz, Baltra, San Cristóbal
Family ELATERIDAE
Genus Conoderus
galapagoensis Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 45, pl. 6, fig. 4. Isabela, San Cristóbal, Floreana
Genus Physorinus
batesoni Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 51. Isabela
blairi Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 50. Santiago
chathami Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 51. San Cristóbal
dichroa Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 47. Santiago
hoodi Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 49. Española
quirsfeldi Mutchler, 1938, Amer. Mus. Novitates, no. 981, p. 4. Santa Cruz
ruficeps Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 48. Isabela
Genus Anchastus
galapagoensis G. R. Waterhouse, 1845, Ann. Nat. Hist., vol. 16, p. 25 (Physorhinus). Galápagos Islands
williamsi Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 53. Santa Cruz
Genus Heterocrepedius
puberulus Boheman, 1858, Fregatten Eugenies Resa, vol. 1, p. 66. Galápagos Islands
Genus Aeolus
fuscatus Steinhal, 1875, Coleopt. Hefte, vol. 14, p. 125. Santa Cruz, Floreana
Genus Grammophorus
galapagoensis Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., ser. 4, vol. 22, p. 55. Pinzón, Santa Cruz
Genus Coptostethus
williamsi Mutchler, 1925, Zoologica, vol. 5, p. 227, fig. 44. Pinta, Marchena, Baltra

[page] 146

Family BUPRESTIDAE
Genus Chrysobothris
williamsi Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 56, pl. 6, fig. 6. Isabela, Pinzón, San Cristóbal
Genus Mastogenius
cuneaticollis Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 58. Pinzón, Santa Cruz
galapagoensis Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 57. Gardner
Family DERMESTIDAE
Genus Dermestes
carnivorus Fabricius, 1775, Syst. Ent., p. 55. Isabela, San Cristóbal
maculatus De Geer, 1774, Mem. Ins., vol. 4, p. 223. Santiago
Family OSTOMIDAE
Genus Temnochila
galapagoensis Mutchler, 1938, Amer. Mus. Novitates, no. 981, p. 5, fig. 4. Isabela, Santiago, Santa Cruz
Genus Tenebroides
sp., Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 61. Isabela
Family NITIDULIDAE
Genus Stelidota
insularis Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 61. Santa Cruz, San Cristóbal
Genus Haptonchus
luteolus Erickson, 1843, Zeitschr. f. Ent., vol. 4, p. 272 (Epuraea). Isabela
Family CUCUJIDAE
Genus Silvanus
tropicalis Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 63. Santa Cruz
Family MONOTOMIDAE
Genus Bactridium
insularis Van Dyke, 1953, Occ, Pap. Calif. Acad. Sci., no 22, p. 63. Santa Cruz
Family DISCOLOMIDAE
Genus Fallia
colorata John, 1959, Senckenbergiana Biol., vol. 40, p. 295, pl. 35, figs. 2a–2e. Santa Cruz

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galapagana John, 1959, Senckenbergiana Biol., vol. 40, p. 295, fig. 2. Santa Cruz
Family COLYDIIDAE
Genus Bitoma
exarata Pascoe, 1866, Jour. Ent., vol. 2, p. 91, pl. 5, fig. 8 (Coniophaea).
Isabela
Family LANGURIIDAE
Genus Comptocarpus
longicollis Motschulsky, 1860, in Schrenck, Reisen u. Forschungen im Amurlande, vol. 2, no. 2, p. 244. San Cristóbal
Family COCCINELLIDAE
Genus Scymnus
galapagoensis G. R. Waterhouse, 1845, Ann. Nat. Hist., vol. 16, p. 41. Fernandina, Santa Cruz
Genus Cycloneda
sanguinea Linnaeus, 1763, Centurea Ins., p. 10. Genovesa, Santa Cruz, Floreana
Genus Olla
abdominalis Say, 1824, Jour. Acad. Nat. Sci., Philadelphia, vol. 4, p. 95. Santa Cruz
Genus Adalia
galapagoensis Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 65. Isabela, Floreana
Genus Psyllobora
bisigma Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no 22, p. 66. Isabela
Family ALLECULIDAE
Genus Lobopoda
galapagoensis Linell, 1898, Proc. U. S. Nat. Mus., vol. 21, p. 266.
Allecula sp., Howard, 1889, Proc. U. S. Nat. Mus., vol. 12, p. 191.
Floreana
Genus Allecula
galapagoensis Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., vol. 22, p. 67. Santiago, Santa Cruz
insularis Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 68. Isabela, Santa Cruz

[page] 148

Genus Ctesia
pedinoidesMäklin, 1875, Acta Soc. Sci. Fennica, vol. 10, p. 681. ? San Cristóbal
Family TENEBRIONIDAE
Genus Stomion
cribricollis Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 75, pl. 3, fig. 3 Darwin, Wolf
Galapagoensis G. R. Waterhouse, 1845, Ann. Nat. Hist., vol. 16, p. 29.
carinipenne Linell, 1898, Proc. U. S. Nat. Mus., vol. 21, p. 262.
carinatipenne Blair, 1933, Ann. Mag. Nat. Hist., ser. 10, vol. 11, p. 477.
piceum Linell, 1898, Proc. U. S. Nat. Mus., vol. 21, p. 262.
punctipennis Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 72 (galapagoensis subsp.).
San Cristóbal, Floreana, Española
helopoides G. R. Waterhouse, 1845, Ann. Nat. Hist., vol. 16, p. 30. Genovesa, San Cristóbal, Floreana, Española, Gardner
laevigatum G. R. Waterhouse, 1845, Ann. Nat. Hist., vol. 16, p. 30.
bauri Linell, 1898, Proc. U. S. Nat. Mus., vol. 21, p. 263.
Santiago, Floreana
linelli Blair, 1933, Ann. Mag. Nat. Hist., ser. 10, vol. 11, p. 478.
laevigatum Linell (not Waterhouse), 1898, Proc. U. S. Nat. Mus., vol. 21, p. 263.
mutchleri Blair, 1933, Ann. Mag. Nat. Hist., ser. 10, vol. 11, p. 477.
Genovesa, Eden, Santa Cruz
longicornis Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 79, pl. 3, fig. 7. Española
longulum Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 76, pl. 3, fig. 4. Isabela, Santa Cruz
obesum Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 74. Pinzón, Brattle, Santa Fé
rugosum Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 80. Pinta
Genus Perepitragus
fuscipes Latreille, 1833, Humb. et Bonpl. Voy., vol. 2, p. 64, pl. 34, fig. 5. San Cristóbal

[page] 149

Genus Ammophorus
abingdoni Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no 22, p. 96, pl. 4, fig. 7. Pinta
antennatus Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 90. Santa Fé
bifoveatus bifoveatus G. R. Waterhouse, 1845, Ann. Nat. Hist., vol. 16, p. 31. Santiago
bifoveatusbarringtoni Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 96. Santa Fé
cooksoni C. Waterhouse, 1877, Proc. Zool. Soc. London, vol. 5, p. 80.
caroli Linell, 1892, Proc. U. S. Nat. Mus., vol. 21, p. 264.
Brattle, San Cristóbal, Floreana
denticollis Boheman, 1858, Fregatten Eugenies Resa, vol. 1, p. 89. ? Galápagos Islands
galapagoensis G. R. Waterhouse, 1845, Ann. Nat. Hist., vol. 16, p. 30.
laevis Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 86, pl. 4, fig. 2 (galapagoensis subsp.).
subpunctatus Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 86, pl. 4, fig. 2 (galapagoensis subsp.).
San Cristóbal
insularis Boheman, 1858, Fregatten Eugenies Resa, vol. 1, p. 89. Isabela, San Cristóbal
obscurus G. R. Waterhouse, 1845, Ann. Nat. Hist., vol. 16, p. 32. Isabela, Pinzó, Eden, Santa Cruz
simplex Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 87. San Cristóbal
Genus Pedonoeces
apicalis Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 105. Genovesa
barringtoni Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 109, pl. 5, fig. 9. Santa Fé
batesoni Blair, 1933, Ann. Mag. Nat. Hist., ser. 10, vol. 11, p. 479. Eden
blairi Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 103. Floreana
bauri Linell, 1898, Proc. U. S. Nat. Mus., vol. 21, p. 265. San Cristóbal
caudatus Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 104, pl. 5, fig. 2. Española

[page] 150

costatus G. R. Waterhouse, 1845, Ann. Nat. Hist., vol. 16, p. 35. Santiago
duncani Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 110. Pinzón
galapagoensis Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p.107. Santiago
lugubris Boheman, 1858, Fregatten Eugenies Resa, vol. 1, p. 91, pl. 1 fig 5(Tessaroma). Pinta, Isabela, Eden, Santa Cruz, Española
morio Boheman, 1858, Fregatten Eugenies Resa, vol. 1, p. 92 (Tessaroma). Santiago, Santa Cruz
opacus Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 114. Rábida
pubescens G. R. Waterhouse, 1845, Ann. Nat. Hist., vol. 16, p. 36. San Cristóbal
spatulus Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 108, pl. 5, fig. 5. Española, Gardner
uniformis Van Dyke, 1953, Occ Pap. Calif Acad. Sci., no 22, p. 105. San Cristbóal
wenmani Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 100, pl.5, fig. 1. Wolf
williamsi Van Dyke, 1953, Occ Pap. Calif. Acad. Sci., no. 22, p. 112. Santa Cruz
Genus Phaleria
manicata Boheman, 1858, Fregatten Eugenies Resa, vol. 1, p. 92. Isabela, Floreana, Española
Genus Gnathocerus
cornutus Fabricius, 1798, Entom. Syst., Suppl., p. 51 (Trogosita.) Isabela, Santiago
Genus Alphitobius
laevigatus Fabricius, 1781, Species Ins., vol. 1, p. 304. Folreana
Genus Rhacius
costipennis Blair, 1933, Ann. Mag. Hist., ser. 10, vol. 11, p. 480. Santiago
Genus Prateus
dentatus Van Dyke, 1953, Occ. Pap. Calif, Acad. Sci., no. 22, p. 119. Santa Cruz
Family ANOBIIDAE
Genus Trichodesma
denticollis Blair, 1928, Ann. Mag. Nat. Hist., ser. 10, vol. 1, p. 675. Floreana

[page] 151

Genus Thaptor
galapagoensisBlair, 1928, Ann. Mag. Nat. Hit., ser. 10, vol. 1, p. 676. Isabela, Santiago, Santa Cruz, Floreana
Genus Eupactus
alutaceus Blair, 1928, Ann. Mag. Nat. Hist., ser. 10, vol. 1, p. 677. Santiago
georgicus Blair, 1928, Ann. Mag. Nat. Hist., ser. 10. vol. 1, p. 676. Floreana
Family BOSTRICHIDAE
Genus Tetrapriocera
longicornis Olivier, 1795, Entomologie, vol. 4, 77, p. 15, pl. 3, fig. 18 (Apate). Pinzón, Santa Cruz
Genus Micrapate
scabratus Erickson, 1847, Wiegm. Arch. f. Naturg., vol. 13, no. 1, p. 87 (Rhizopertha). San Cristóbal
Genus Amphicerus
cornutus galapaganus Lesne, 1910, Bull. Mus. Hist. Nat. d'Hist. Naturelle, vol. 16, p. 184 (Schistoceros). Wolf, Pinta, Isabela, Pinzón, Santa Cruz, Española
sp., G. R. Waterhouse, 1845, Ann. Nat. Hist., vol. 16, p. 36(Apate).
unicantus C. Waterhouse, 1877, Proc. Zool. Soc. London, p. 82 (Bostrichus)
punctipennis Linell, 1898, Proc. U.S Nat. Mus., vol. 21, p. 256.
Family SCARABAEIDAE
Genus Copris
lugubris Boheman, 1858, Fregatten Eugenies Resa, vol. 1, p. 42. ? Galápagos Islands
Genus Ataenius
aequatorialis Petrovitz, 1961, Entomologische Arb., vol. 12, p. 355. Santa Cruz
arrowi Hinton, 1936, Ann. Mag. Nat. Hist., ser. 10, vol. 17, p. 414. Pinta, Isabela, Santa Cruz
cribrithorax, Blair 1933 (not Bates, 1887), Ann. Mag. Nat. Hist., ser. 10, vol. 11, p. 476.
dampieri Petrovitz, 1961, Entomologische Arb., vol. 12, p. 356. Floreana
scutellaris Harold, 1867, Coleopterologische Hefte, vol. 18, p. 82. Pinta, Isabela, San Cristóbal, Floreana

[page] 152

Genus Trox
galapagoensis Van Dyke, 1953, Occ. Pap. Calif. Aced. Sci., no. 22, p. 124, pl. 6, fig. 7. Darwin
suberosus Fabricius, 1775, Systema Ent., p. 31. Pinta, Isabela, Santiago, Santa Cruz, San Cristóbal, Floreana, Española
seymourensis Mutchler, 1925, Zoologica, vol. 5, p. 229. Pinta, Isabela, Baltra
Genus Neoryctes
galapagoensis G. R. Waterhouse, 1845, Ann. Nat. Hist., vol. 16, p. 26 (Oryctes). Isabela, San Cristóbal, Floreana
linelli Mutchler, 1938, Amer. Mus. Novitates, no. 981, p. 10, figs. 10–12. San Cristóbal
sp., Van Dyke,1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 126. Santa Cruz
Family PASSALIDAE
Genus Passalus
interruptus Linnaeus, 1767, Systema Naturae, ed. 12, p. 560 (Lucanus).
tlascala Linell, 1898 Proc. U. S. Nat. Mus., vol. 21, p. 257.
Floreana
Family CERAMBYCIDAE
Genus Parandra
galapagoensis Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p, 127. Isabela, Santiago, Santa Cruz
Genus Stenodontes
galapagoensisMutchler, 1925, Amer. Mus. Novitates, no. 981, p. 11, fig. 6. Santa Cruz
molarius Bates, 1879, Biol. Centr.-Americana, vol. 5, p. 9 (Mallodon). Santa Cruz, San Cristóbal, Floreana
Genus Strongylaspis
kraepelini Lameere, 1903, Mem. Soc. Ent. Belgique, vol. 11, p. 28. Santiago
Genus Achryson
galapagoensis Linell, 1898, Proc. U. S. Nat. Mus., vol. 21, p. 259. Baltra, San Cristóbal
Genus Eburia
amabilis Boheman, 1859, Fregatten Eugenies Resa, vol. 1, p. 150. ? Galápagos Islands

[page] 153

lanigera Linell, 1898, Proc. U. S. Nat. Mus., vol. 21, p. 259.
amabilis Howard, 1890, Proc. U. S. Nat. Mus., vol. 12, p. 192.
Isabela, Santiago, Rábida, Pinzón, Eden, San Cristobal, Floreana, Española, Gardner
proletaria Erickson, 1847, Arch. f. Naturgesch., vol. 13, p. 140.
bauri Linell, 1898, Proc. U. S. Nat. Mus., vol. 21, p. 260
Santiago, San Cristóbal
Genus Compsa
apicalis Blair, 1933, Ann. Mag. Nat. Hist., ser. 10, vol. 11, p. 481. Santiago
Genus Desmiphora
hirticollis Olivier, 1795, Entomologie, vol. 4, no. 68, p. 11, pl. 4, fig. 37 (Saperda.) Santiago
Genus Estoloides
galapagoensis Blair, 1933, Ann Mag. Nat. Hist., ser. 10, no. 11, p. 482 (Estola.) Isabela, Santa Cruz
Genus Estola
cribrata Blair, 1933, Ann. Mag. Nat. Hist., ser. 10, vol. 11, p. 483. Isabela
duncani Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 135. Pinzón
insularis Blair, 1933, Ann. Mag. Nat. Hist., ser. 10, no. 11, p. 483. Isabela, Santiago, Santa Cruz
Genus Acanthoderes
galapagoensis Linell, 1898, Proc. U. S. Nat. Mus., vol. 21, p. 261.
vonhageni Muthler, 1938, Amer. Mus. Novitates, no. 981, p. 14 (galapagoensis var.).
Isabela, Santiago, Santa Cruz, San Cristóbal
Genus Leptostylus
galapagoensis Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 136, pl. 7, fig. 1. Isabela, Santiago
Williamsi Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 137. Santiago, Rábida, Santa Cruz
Genus Taeniotes
hayi Mutchler, 1938, Amer. Mus. Novitates, no. 981, p. 13, fig. 3. ? Galápagos Islands
Family CHRYSOMELIDAE
Genus Metachroma
labrale Blair, 1933, Ann. Mag. Nat. Hist., ser. 10, vol. 11, p. 484. Isabela, Santiago, Santa Cruz

[page] 154

Genus Gynandrobrotica
ventricosa Jacoby, 1887, Biol. Centr.-Amer., Coleoptera, vol. 6, p. 543, pl. 32, fig. 1 (Diabrotica). ? San Cristóbal
Genus Acalymma
limbata C. Waterhouse, 1877, Proc. Zool. Soc. London, 1877, p. 81 (Diabrotica). ?Galápagos Islands
Genus Docema
darwiniMutchler, 1924, Zoologica, vol. 5, p. 230. Genovesa, Eden
galapagoensis G. R. Waterhouse, 1845, Ann. Mag. Nat. Hist., vol. 16, p. 39 (Haltica). Isabela, Rábida, Floreana
Genus Longitarsus
galapagoensis Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 140. Santa Cruz, San Cristóbal, Floreana
lunatus C. waterhouse, 1877, Proc. Zool. Soc. London, 1877, p. 81.
Family BRUCHIDAE
Genus Spermophagus
galapagoensis Blair, 1928, Ann. Mag. Nat. Hist., ser. 10, vol. 1, p. 678. Santiago, Española
Genus Bruchus
fuscomaculatus Blair, 1928, Ann. Mag. Nat. Hist., ser. 10, vol. 1, p. 679. Santiago
Family ANTHRIBIDAE
Genus Ormiscus
variegatus G. R. Waterhouse, 1845, Ann. Nat Hist., vol. 16, p. 37. Pinta, Isabela Gardner, Floreana
Family CURCULIONIDAE
Genus Amphideritus
cuneiformis G. R. Waterhouse, 1845, Ann. Nat. Hist, vol. 16, p. 38(Otiorhynchus). San Cristóbal, Floreana
Genus Pantomorus
blairi Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 146. Santiago
caroli Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 147. Floreana
conwayensis Mutchler, 1938, Amer. Mus. Novitates, no. 981, p. 15. Santa Cruz
crockeri Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 147. Genovesa

[page] 155

galapagoensis Linell, 1898, Proc. U. S. Nat. Mus., vol. 21, p. 268. San Cristóbal
williamsi Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 150. Isabela
Genus Gerstaeckeria
galapagoensis barringtoni Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 152. Santa Fé
galapagoensis galapagoensis Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 151, pl. 7, fig. 8. Pinta
galapagoensis hoodensis Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 153. Española
galapagoensis seymourensis Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 153. Baltra
Genus Gerais
batesoni Blair, 1933, Ann. Mag. Nat. Hist., ser. 10, vol. 9, p. 485. San Cristóbal
Genus Lembodes
subcostaus Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 154, pl. 7, fig. 2. Isabela, Pinzón
Genus Anchonus
galapagoensis G. R. Waterhouse, 1845, Ann. Nat. Hist., vol. 16, p. 39. Santiago
Genus Dryotribus
mimeticus Horn, 1873, Proc. Amer. Philos. Soc., 13: 433. Fernandina
Genus Micrancylus
gracilis Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 156. Pinta
Genus Neopentarthrum
cunicollis Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 157. Isabela, Pinzón
glabrum Van Dyke, 1953, Occ. Pap. Calif. Acad. Sci., no. 22, p. 158. Pinta, Eslpañola
mutchleri Van Dyke, Occ. Pap. Calif. Acad, Sci., no. 22, p. 158. Pinta
towerensis Mutchler, 1925, Zoologica, vol. 5, p. 231, fig. 45. Genovesa
Family PLATYPODIDAE
Genus Platypus
santacruzensis Mutchler, 1925, Zoologica, vol. 5, p. 232, fig. 24. Isabela, Santa Cruz

[page] 156

Family SCOLYTIDAE
Genus Pycnarthurm
insulare Blair, 1933, Ann. Mag. Nat. Hist., ser. 10, vol. 11, p. 478. Genovesa, Isabela, Española

Order LEPIDOPTERA

Comprehensive works on Galápagos Lepidoptera are by Williams (1911) for butterflies and hawk moths and Schaus (1923) for other moths. Meyrick (1926) described additional Microlepidoptera, some of which have been revised by Clark (1958). Richards (1941) dealt with the Noctuidae collected on Hancock expeditions. Beebe (1923) gives field notes for his collections of Lepidoptera. Burns (1964) clarifies the Van Duzee record of Erynnis (Hesperiidae). Kernbach (1962) gives the most recent review of Sphingidae. E. G. Munroe (personal communication) has indicated generic placements in the Pyralidae, J. A. Powell in the Tortricidae, and R. J. Collins (personal communication) has checked bibliographic references throughout. In Washington, sections of the manuscript were reviewed as follows:

D. R. Davis (USNM)—Tortricidae (Olethreutinae), Tineidae; W. D. Duckworth (USNM)—Pyralidae (Phycitinae), Yponomeutidae; W. D. Field (USNM)—Pieridae, Nymphalidae, Danaidae, Lycaenidae, Hesperidae, Springidae, Arctiidae; R. W. Hodges (USDA)—Pyralidae (Pyraustinae, Crambinae), Pterophoridae, Tortricidea (Tortricinae), Gelechiidae, Balstobasidae; E. L. Todd (USDA)—Noctuidae, Geometridae.

Suborder RHOPALOCERA
Family PIERIDAE
Genus Phoebis
Sennae Linnaeus, 1766, Syst. Nat., Ed. 12, p. 764 (Papilio Danaus) (from Sennae Edwards fide Williams, 1911). Isabela, San Cristóbal, Floreana, Santa Cruz, Santiago, Fernandina, Eden, Daphne, Blatra, Pinta
Family NYMPHALIDAE
Genus Agraulis
vanillae galapagensis Holland, 1889, Proc. U. S. National Mus., vol. 12, p. 194 (as var.). Isabela, Foreana, San Cristóbal, Santa Cruz, Santiago, Fernandina, Pinta, Baltra, Eden
Genus Vanessa
virginiensis Drury, 1773, Ill. Exot. Ent., 2, pl. 13, fig. 2 (Papilio). Isabela
carye Hübner, 1945, Samml. exot. Schmet., vol. 1, pl. 41 (Vanessa). San Cristóbal

[page] 157

Family DANAIDAE
Genus Danaus
plexippus Linnaeus, 1758, Syst. Nat. Ed. 10, p. 471 (Papilio Danaus). San Cristóbal
Family LYCAENIDAE
Genus Leptotes
parrhasioides Wallengren, 1860, Wien. Ent. Monat., vol. 4, p. 37 (Lycaena). Floreana, San Cristóbal, Isabela, Fernandina, Santiago, Española, Pinzón, Santa Cruz, Eden, Baltra
Family HESPERIIDAE
Genus Urbanus
dorantes galapagensis Williams, 1911, Proc. Calif. Acad. Sci., ser. 4, vol. 1, p. 303. San Cristóbal, Isabela, Española, Pinzón, Floreana, Santiago, Santa Cruz, Genovesa, Eden, Baltra
Genus Erynnis
funeralis Scudder and Burgess, 1870, Proc. Boston Soc. Nat. Hist., vol. 13, p. 293 (Nisoniades).
persius Van Duzee, 1933 (not Scudder, 1864), Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 145 (Thanaos). (Van Duzee suggests an error in labeling).
San Cristóbal
Genus Heteropia
sp. near imalena Butler, 1874, Lep. Exot., p. 109, pl. 40, fig. 1 (Telegonus). (Van Duzee, 1933, suggests error in labeling). Isabela
Suborder HETEROCERA
Family SPHINGIDAE
Genus Agrius
cingulatus Fabricius, 1775, Syst. Ent., p. 545 (Sphinx). San Cristóbal, Isabela, Santa Cruz, Baltra, Floreana

[page] 158

.
Genus Protoparce
sexta leucoptera Rothschild and Jordan, 1903, Nov. Zool. Suppl., vol. 9, p. 79. San Cristóbal, Isbela, Santa Cruz
rustica calapagensis Holland, 1889, Proc. U. S. Nat. Mus., vol. 12, p. 195.
nigrita Rothschild and Jordan, 1903, Nov. Zool. Suppl., vol. 9, p. 86 (rustica ab.).
Postscripta Clark, 1926, Proc. New England zool. Club, vol. 9,p. 70 (rustica subsp.).
Floreana, San Cristóbal, Española, Isabela, Baltra, Santa Cruz, Genovesa
Genus Erinnyis
alope dispersa Kernbach, 1962, Opusc. Zool., vol. 63, p. 9. Santa Cruz
ello encantada Kernbach, 1962, Opusc. Zool., vol. 63, p. 10. Santa Cruz, Floreana, San Cristóbal, Isabela
Obscura conformis Rothschild and Jordan, 1903, Nov. Zool. Suppl., vol. 9, p. 369. Isabela, Baltra, Santa Cruz, Floreana
Genus Epistor
lugubris delanoi Kernbach, 1962, Opusc. Zool., vol. 63, p. 11. Santa Cruz, Isabela, Floreana
Genus Pholus
fasciatus tupaci Kernbach, 1962, Opusc. Zool., vol. 63, p. 12. Santa Cruz
labruscae yupanquii Kernbach, 1962, Opusc. Zool., vol. 63, p. 13. Santa Cruz, Floreana
Genus Xylophanes
norfolki Kernbach, 1962, Opusc. Zool., vol. 63, p. 14. Santa Cruz
tersa Linnaeus, 1771, Mant. plant., vol. 2, p. 538 (Sphinx). San Cristóbal
Genus Celerio
lineata florilega Kernbach, 1962, Opusc. Zool., vol. 63, p. 16. Santa Cruz, Floreana, San Cristóbal, Isabela, Baltra
Family ARCTIIDAE
Genus Utetheisa
galapagensis Wallengren, 1860, Wien. Ent. Monat., vol. 4, p. 161 (Euchalia). Isabela, Santa Cruz, Santiago, Floreana, Baltra
ornatrix Linnaeus, 1758, Syst. Nat., Ed. 10, vol. 1, p. 511 (Phalaena noctua). Isabela, Baltra

[page] 159

Family NOCTUIDAE
Subfamily AGROTINAE
Genus Agrotis
ipsilon Hufnagel, 1766, Berl. Mag., vol 3, p. 416 (Phalaena). Isabela, Floreana
galapagosensis Kohler, 1961, An Soc. Cient. Argentina, vol. 172, p. 71–72 (subgen. Scotia). Floreana
Genus Feltia
subterranea Fabricius, 1794, Ent. Syst., vol. 3, no. 2, p. 70 (Noctua). Santiago, Isabela
Genus Peridroma
conwayi Richards, 1941, Hancock Pac. Exp., vol. 5, p. 235. Floreana
saucia Hübner, 1803–1808, Samml. Eur. Schmett. Noct. f. 378 (Noctua). Floreana
Genus Heliothis
cystiphora Wallengren, 1860, Wien. Ent. Monat., vol. 4, p. 172 (Anthoecia).
inflata Wallengren, 1860, Wien. Ent. Monat., vol. 4, p. 172 (Anthoecia) onca Wallengren, 1860, Wien. Ent. Monat., vol. 4, p. 172 (Anthoecia).
Santa Cruz, Santiago, Isabela, San Cristóbal, Baltra
virescens Fabricius, 1781, Spec. Ins., vol. 2, p. 216 (Noctua). San Cristóbal, Isabela, Genovesa
Genus Euxoa
williamsi Schaus, 1923, Zoologica, vol. 5, p. 32. Santa Cruz, Satiago, Isabela, Baltra
Genus Lycophotia
oceanica Schaus, 1923, Zoologica, vol. 5, p. 32. Baltra
Subfamily HADENINAE
Genus Pseudaletia
cooperi Schaus, 1923, Zoologica, vol. 5, p. 33 (Cirphis) Santa Cruz
Subfamily AMPHIPYRINAE
Genus Magusa
orbifera Walker, 1857, Cat. Lep. Brit. Mus., vol. 11, p. 761 (Xylina). Isabela

[page] 160

Genus Platysenta
apameoides Guenée, 1852, Sp. Gen., Noct., vol. 1, p. 229 (Perigea) Santiago
ebba Schaus, 1923, Zoologica, vol. 5, p. 36 (Perigea) n. comb. Santiago, Santa Cruz, Baltra
ruthae Schaus, 1923, Zoologica, vol. 5, p. 35 (Perigea). Santa Cruz, Isabela
Genus Harrisonia
williamsi Schaus, 1923, Zoologica, vol. 5, p. 36. Baltra, Santa Cruz
Genus Prodenia
dolichos Fabricius, 1794, Ent. Syst., vol. 3, p. 95 (Noctua). Floreana
eridania form linea Fabricius, 1794, Ent. Syst., vol. 3, p. 106 (Noctua). Floreana
Genus Spodoptera
frugiperda J.E. Smith, 1797, Lep. Ins. Georgia, vol. 2, p. 191, pl. 96 (Phalaena). Santa Cruz
roseae Schaus, 1923, Zoologica, vol. 5, p. 33 (Trachea), n. comb. Santiago, Isabela, Baltra, Santa Cruz
GenusCatabena
sp.? Schaus, 1923, Zooolgica, vol. 5, p. 25. Genovesa
GenusAmyna
insularum Schaus, 1923, Zoologica, vol. 5, p. 37. Isabela, Santa Cruz, Santiago, Floreana, San Cristóbal, Baltra
Subfamily ACONTIINAE
Genus Heliocontia
margana Fabricius, 1794, Ent. Syst., vol. 3, no. 2, p. 257 (Pyralis). Santa Cruz, Baltra
Genus Spragueia
creton Schaus, 1923, Zoologica, vol. 5, p. 38. Genovesa, Baltra
plumbeata Schaus, 1923, Zoologica, vol. 5, p. 38. Santa Cruz
Subfamily PLUSIINAE (=PHYTOMETRINAE)
Genus Pseudoplusia
includens Walker, 1857, Cat. Lep. Brit. Mus., vol. 12, p. 914 (Plusia).Santa Cruz, Floreana, Sanitago, San Cristóbal, Baltra
Genus Autoplusia
egena galapagensis Schaus, 1923, Zoologica, vol. 5, p. 41 (as form) (Syngrapha). Santiago

[page] 161

Subfamily EUTELIINAE
Genus Paectes
indefatigabilis Schaus, 1923, Zoologica, vol. 5, p. 38. Santa Cruz, Baltra
isabel Schaus, 1923, Zoologica, vol. 5, p. 39. Santa Cruz
Subfamily SARROTHRIPINAE
Genus Characoma
nilotica Rogenhofer, 1882, Verh. Zool. Bot. Ges. Wien., vol. 31, p. 26 (Sarrothripa). San Cristóbal
Subfamily CATOCALINAE (sensu Richards=NOCTUINAE Schaus)
Genus Melipotis
harrisoni Schaus, 1923, Zoologica, vol. 5, p. 42. Santa Cruz, Isabela, Baltra, Santiago
indomita Walker, 1857, Cat. Lep. Brit. Mus., vol. 13, p. 1161 (Bolina).
nigrescens Williams, 1911, Proc. Calif. Acad. Sci., ser. 4, vol. 1, p.?
Santa Cruz, Baltra, Isabela, Floreana
Genus Anticarsia
gemmatalis Hübner, 1818, Zutr. Exot. Schmentt., vol. 1, p. 26, figs. 153, 154. Santa Cruz, San Cristóbal, Baltra
Genus Gonodonta
biarmata galapagensis Todd, 1959, Tech. Bull. 1201, U. S. Department of Agriculture, p. 20. Floreana, San Cristóbal, Santa Cruz
Genus Epidromia
zephyritis Schaus, 1923, Zoologica, vol. 5, p. 43. Floreana, Santa Cruz
Genus Psorya (? Smyra—see Richards, 1941)
hadesia Schaus, 1923, Zoologica, vol. 5, p. 44. Baltra, Floreana
Genus Zale
sp. (viridans group) Richards, 1941, Hancock Pac. Exp., vol. 5, p. 243. Floreana
Genus Ascalapha
odorata Linnaeus, 1758, Syst. Nat., ed. 10, p. 505 (Phalaena Bombyx). Floreana
Genus Anomis
professorum Schaus, 1923, Zoologica, vol. 5, p. 42. Isabela, Baltra, Santa Cruz, San Cristóbal
Genus Mocis
incurvalis Schaus, 1923, Zoologica, vol. 5, p. 41. Floreana, Santa Cruz, Baltra
latipes Guenée, 1852, Sp. Gen., Noct., vol. 3, p. 314 (Remigia) Santa Cruz, Baltra

[page] 162

Genus Rivula
dubiosa Schaus, 1923, Zoologica, vol. 5, p. 44. Santa Cruz
Family GEOMETRIDAE
Genus Camptogramma
stellata Guenée, 1857, Uranides et Phalenites, vol. 2, p. 443 (Scotosia). Santa Cruz, Baltra
Genus Cosymbia
impudens Warren, 1904, Nov. Zool., vol. 11, p. 487 (perixera) Santa Cruz, Santiago, Isabela, Gardner
Genus Sericosema
lignata Warren, 1905, Nov. Zool., vol. 12, p. 362. San Cristóbal
Family PYRALIDAE
Subfamily PYRAUSTINAE
Genus Hymenia
recurvalis Fabricius, 1775, Syst. Ent., 644 (Phalaena). Baltra
Genus Pilocrocis
ramentalis Lederer, 1863, Wien. Ent. Monat., vol. 7, p. 430. Santiago, Baltra
Genus Herpetogramma
chathamalis Schaus, 1923, Zoologica, vol. 5, p. 45 (pilocrocis) (n. comb.). San Cristóbal, Santa Cruz, Baltra
Genus Asciodes
gordialis Guenée, 1854, Delt. et Pyral., p. 374. Santa Cruz, Isabela, San Cristóbal, Baltra
Genus Pleuroptya
silicalis Guenée, 1854, Delt. et Pyral., p. 349 (Botys) (n. comb.). Santa Cruz, Santiago, Isabela, San Cristóbal, Baltra
Genus Polygrammodes
elevata Fabricius, 1793, Ent. Syst. III, 2, no. 325, p. 216 (phalaena). Santiago
Genus Pyrausta
eneanalis Schaus, 1923, Zoologica, vol. 5, p. 45. Santa Cruz, Baltra, Genovesa
Genus Piletocera
bufalis Guenée, 1854, Delt. et Pyral., p. 245 (Stenia). Santa Cruz
Genus Beebea
guglielmi Schaus, 1923, Zoologica, vol. 5, p. 46. San Cristóbal

[page] 163

Subfamily CRAMBINAE
Genus Euchrombius
ocelleus Haworth 1811, Lep. Brit., p. 486 (Palparia). Santa Cruz, Baltra
Subfamily PHYCITINAE
Genus Etiella
zinckenella (Treit.) 1832, Schmett. Eur., vol. 9. no. l, p. 201. Santa Cruz, Baltra
Genus Fundella
agapella Schaus, 1923, Zoologica, vol. 5, p. 47 Isabela
GenusFulrada
carpasella Schaus, 1923, Zoologica, vol. 5, p. 47 (Piesmopoda). Santa Cruz
Genus Tota
galdinella Schus, 1923, Zoologica, vol. 5, p. 46 (Elasmopalpus). Santa Cruz, Baltra, Santiago
Genus Nicetiodes
apianella Schaus, 1923, Zoologica, vol, 5, p. 48 Santa Cruz, Baltra, Genovesa
Family PTEROPHORIDAE
Genus Trichoptilus
derelictus Meyrick, 1926, Trans. Ent. Soc. Lond., vol. 74, p. 276. Floreana
Genus Pterophorus
nephogenes Meyrick, 1926, Trans. Ent. Soc. London, vol. 74, p. 276. Floreana, Isabela
sp. Schaus, 1923, Zoologica, vol. 5, p. 30. Daphne Major, Santa cruz
Family TORTRICIDAE
Subfamily TORTRICINAE
Genus Platynota
colobota Meyrick, 1926, Trans. Ent. Soc. Lond., vol. 74, p. 276. Isabela
Subfamily OLETHREUTINAE
Genues Strepsicrates
smithiana Walsingham, 1892, Proc. Zool. Soc. London, p. 506. Baltra
spp. Schaus, 1923, Zoologica, vol. 5, p. 31 (3 species). Baltra, Santiago
Genus Crocidosema
plebiana Zeller, 1847, Isis, p. 721. Baltra, Isabela,

[page] 164

synneurota Meyrick, 1926, Trans. Ent. Soc. Lond., vol. 74, p. 276. Santa Cruz
Family GELECHIIDAE
Genus Aristotelia
howardi Walsingham, 1909, Biol. Centr. Am. Ins. Lep.-Heter., IV, p. 23 Baltra
naxia Meyrick, 1926, Trans. Ent. Soc. London, vol. 74, p. 277. Floreana
Genus Stegasta
bosqueela Chambers, 1875, Canad. Ent., vol. 7, p. 92 (Oecophora). Santa Cruz
Genus Gelechia
gnathodoxa Meyrick, 1926, Trans. Ent. Soc. London, vol. 74, p. 277. Santiago
protozona Meyrick, 1926, Trans. Ent. Soc. London, vol. 74, p. 277. Isabela
Family BLASTOBASIDAE
Genus Blastobasis
crotospila Meyrick, 1926, Trans. Ent. Soc. Lonton, vol. 74, p. 278. Santiago
Family YPONOMEUTIDAE
Genus Atteva
hysginiella Wallengren, 1861, Freg. Eugenies Resa, vol. 2, p. 386
sylpharis Butler, 1877, Proc. Zool. Soc. Lond., 1877, p. 87.
Isabela, santa Cruz, Santiago, San Cristóbal, Baltra, Daphne Major
monerythra Meyrick, 1926, Trans. Ent. Soc. London, vol. 74, p. 278 Floreana
Family TINEIDAE
Genua Xylesthia
pruniramiella Clemens, 1859, Proc. Acad. Nat. Sci. Philadclphia, vol. 11, p. 259. Santiago

Order DIPTERA

Comprehensive reports on Galápagos Diptera may be found in Coquillett (1901), Johnson (1924), and Curran (1932, 1934). Authority for the synonymy of pseudolfersia diomedae Coquillett with Olfersia aenescens Thomson, is Maa (1963), and for that of Ceratopogon galapagensis Coquillett with Forcipomyia fuliginosa Meigen, Wirth (1956). This section has been read by R. H. Foote, C. W. Sabrosky, G. C. Steyskal, Alan Stone, and W. W. Wirth.

[page] 165

Family CHIRONOMIDAE
Genus Chironomus
sp., Johnson, 1924, Zoologica, vol. 5, p. 86. Baltra
sp., Curran, 1934, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 149 San Criostóbal
Genus Tanypus
sp., Johnson, 1924, Zoologica, vol. 5, p. 86. Baltra
Genus Thalassomya
longipes Johnson, 1924, Zoologica, vol. 5, p. 86 (Galapagomyia). Santa Cruz, Floreana
Genus Clunio
schmitti Stone & Wirth, 1947, Proc. Ent. Soc. Washington, vol. 49, p. 217. Fernandina
Family TIPULIDAE
Genus Erioptera
(Subgenus Trimiera)
pilipes Fabricius, 1787, Mantissa Insectorum, vol. 2, p. 324 (Tipula). San Cristóbal
Genus Limonia
(Subgenus Dicranomyia)
galapagoensis Alexander, 1962, Opuscula zoologica, vol. 61, p. 1. Santa Cruz, San Cristóbal
(Subgenus Geranomyia)
stigmaticaphilippi, 1865, Verhandl. Kais. Kon. Zool.-bot. Ges. Wien, vol. 15, p. 598 (Plettusa). Isabela
tibialis Loew, 1851, Linn. Ent. vol. 5, p. 397 (Aporosa). Santa Cruz, San Cristóbal, Floreana
Family CULICIDAE
Genus Aedes
taeniorhynchus Wiedemann, 1821, Diptera Exot., p. 43 (Cules). Genovesa, Isabela, Santiago, R´bida, Santa Cruz, Baltra, San Cristóbal, Floreana
Family CERATOPOGONIDAE
Genus Forcipomyia
fuliginosa Meigen, 1818, Syst. Beschr. Eur. Zweifl. Ins., vol. 1, p. 86 (Ceratopogon).
galapagensis Coquillett, 1901, Proc. Washington Acad. Sci., vol. 3, p. 372(Ceratopogon).
Isabela, Baltra, Santa Cruz, Gardner

[page] 166

Family SCIARIDAE
Genus Sciara
sp., Curran, 1934, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 149. Baltra
Family STRATIOMYIDAE
Genus Pelagomyia
dubia Curran, 1932, Nyt Mag. Naturvidensk., vol. 71, p. 348.
albitalus Coquillett (not Williston), 1901, Proc. Washington Acad. Sci., vol. 3, p. 373.
Isabela, Santiago, Santa Cruz, Baltra, San Cristóbal, Floreana
Genus Nemotelus
acutirostris Loew, 1863, Berliner Ent. Zeitschr., vol. 7, p. 8. Isabela
albiventris Thomson, 1869, Freg, Eugenies Resa, Zool., vol. 1, p. 462. Isabela
Family TABANIDAE
Genus Tabanus
vittiger Thomson, 1869, Freg. Eugenies Resa, Zool., vol. 1, p. 451. Isabela, Santa Cruz, San Cristóbal, Floreana
Family BOMBYLIIDAE
Genus Villa
brachialis Thomson, 1869, Freg. Eugenies Resa, Zool., vol. 1, p. 484 (Anthrax). Isabela
curvirostris Thomson, 1869, Freg. Eugenies Resa, Zool., vol. 1, p. 483 (Anthrax). Galápagos Islands
primitiva Walker, 1849, List Diptera Brit. Mus., vol. 2, p. 257 (Anthrax).
lateralis Thomson, 1869, Freg. Eugenies Resa, Zool., vol. 1, p. 482 (Anthrax).
nudiuscula Coquillett (not Thomson), 1901, Proc. Washington Acad. Sci., vol. 3, p. 373 (Anthrax).
Santa Cruz, Santa Fé, Floreana
Genus Lepidanthrax
tincta Thomson, 1869, Freg. Eugenies Resa, Zool., vol. 1, p. 483 (Anthrax). Santa Cruz, Bartholomew
Family ASILIDAE
Genus Ommatius
marginellus Fabricius, 1781, Spec. Ins., vol. 2, p. 464 (Asilus). Santiago, Santa Cruz

[page] 167

Family EMPIDIDAE
Genus Drapetis
zonalis Curran, 1932, Nyt Mag. Naturvidensk., vol. 71, p. 351. Santa Cruz, Floreana
Family DOLICHOPODIDAE
Genus Condylostylus
dentaticauda Van Duzee, 1933, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 66. Santiago
tibialis Thomson, 1869, Freg. Eugenies Resa, Zool., vol. 1, p. 507 (Auchineura). Fernandina, Isabela
Genus Chrysotus
brevicornis Van Duzee, 1933, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 68. Santa Cruz
brevispina Van Duzee, 1933, Proc. Calif. Acad. Sci., ser. 4 vol. 21, p. 68. Isabela, Santa Cruz, San Cristóbal
latifacies Van Duzee, 1933, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 69. Santa Cruz
Genus Asyndetus
interruptus Loew, 1861, Ent. Monatschr., vol. 5, p. 37. Pinta
versicolor Johnson, 1924, Zoologica, vol. 5, p. 88. Baltra
Genus Aphrosylus
setosus Curran, 1933, Nyt Mag. Naturvidensk., vol. 71, p. 352. Floreana
Genus Paracleius
? pusillus Macquart, 1844, Diptera Exot., suppl. 1, p. 121 (Dolichopus). Fernandina, Isabela
Genus Vanduzeeia
cheesmani Parent, 1934, Mem. Soc. Sci. Nat. & Math. Cherbourg, vol. 41, p. 258. Santiago
Family PHORIDAE
Genus Dohrniphora
cornuta Bigot, 1856, Sagra's Hist. Fis. Nat. Cuba. vol. 7, p. 348.
venusta Coquillett, 1895, Canadian Ent., vol. 27, p. 107.
crockeri Van Duzee, 1933, Proc. Calif. Acad. Sci., ser. 4, vol. 21. p. 70.
Baltra, Floreana
willowsi Van Duzee, 1933, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 70. Baltra

[page] 168

Genus Megaselia
scalaris Loew, 1866, Berliner Ent. Zeitschr., vol. 10, p. 53 (phora). Genovesa
Family SYRPHIDAE
Genus Allograpta
splendens Thomas, 1869, Freg. Eugenies Resa, Zool., vol. 1, p. 501 (syrphus). Santiago, San Cristóbal, Floreana
Genus Baccha
clavata Fabricius, 1794, Ent. Syst., vol. 4, p. 298 (Syrphus)
facialis Thomson, 1869, Freg. Eugenies Resa, Zool., vol. 1,p. 504.
Isabela, Santiago, San Cristóbal, Floreana
Genus Mesograpta
crockeri Curran, 1934,Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 154.
duplicata Cogquillett (not Wiedemann), 1901, Proc. Washington Acad. Sci., vol. 3, p. 374.
Isabela, Santiago, San Cristóbal, Floreana
Genus Xanthandrus
sp., Curran, 1934, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 155. Pinta
albomaculatus smith, 1877, Proc. Zool. Soc. London, 1877, p. 84 (Syrphus). Floreana
Family OTITIDAE
Genus Acrosticta
scrobiculata Loew, 1867, Berliner Ent. Zeitschr., vol. 11, p. 293. Floreana
Genus Euxesta
galapagensis Curran,1934, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 156. Española
nitidiventris Loew, 1873, Mon. No. Amer. Diptera, vol. 2, p. 157. Isabela, Santa Cruz, Floreana
notata Wiedemann, 1830, Ausser. Zweifl. Jnsekten, vol. 2, p. 462 (Ortalis). Floreana
Genus Pareuxesta
hyalinata Coquillett, 1901, Proc. Washington Acad. Sci., vol. 3, p. 377. Isabela
latifasciata Coquillett, 1901, Proc. Washington Acad. Sci., vol. 3, p. 376.
intermedia Coquillett, 1901, Proc. Washington Acad. Sci., vol. 3, p. 377.
Genovesa, Isabela

[page] 169

obscura Coquillett, 1901, Proc. Washington Acad. Sci., vol. 3, p. 377. Daphne Major, Isabela
Family TEPHRITIDAE
Genus Paroxyna
crockeri Curran, 1934, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 157, fig. 1. Isabela, Santiago, Rábida, Santa Cruz, Baltra, Floreana
Genus Euaresta
aesia Walker, 1849, List Diptera Brit. Mus., vol. 4, p. 1006 (Trypeta). Galápagos Islands
Family SPHAEROCERIDAE
GenusLeptocera
discalis Malloch, 1912, Smithsonian Misc. Coll., vol. 59, p. 433. Floreana
GenusSphaerocera
galapagensis Curran, 1934, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 158. San Cristóbal
Family DROSOPHILIDAE
Genus Drosophila
immigrans Sturtevant, 1921, Carnegie Inst. Washington, Publ. 301, p. 83. Santiago, Floreana
nebulosa Sturtevant, 1916, Ann. Ent. Soc. Amer., vol. 9, p. 327. San Cristóbal
simulans Sturtevant, 1919, Psyche, vol. 26, p. 153. San Cristóbal, Baltra
willistoni Sturtevant, 1916, Ann. Ent. Soc. Amer., vol. 9. p. 327. Baltra
sp., Wheeler, 1961, Ann. Ent. Soc. Amer., vol. 54, p. 750 Santa Cruz
Genus Gitona
braziliensis Costa Lima, 1950, Arthropoda, vol. 1, p. 251. Santa Cruz
Family EPHYDRIDAE
Genus Dimecoenia
gilvipes Coquillett, 1901, Proc. washington Acad. Sci., vol. 3, p. 377 (Ephydra). Isabela
Genus Scatella
galapagensis Curran, 1934, Proc. Calif. Acad. Sci., ser.4, vol. 21, p. 161. San Cristóbal

[page] 170

Family CANACEIDAE
Genus Canace
maritima Wirth, 1951, Occ. Pap. Bernice P. Bishop Mus., vol. 20, p. 263. Bartholomew
snodgrassii Coquillett, 1901, Proc. Washington Acad. Sci., vol. 3, p. 378. Fernandina, Isabela
Genus Nocticanace
galapagensis Curran, 1934, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 160 (Procanace). Isabela, Santa Cruz, Wenman, Tower, Abingdon, Hood
Family CHLOROPIDAE
Genus Hippelates
galapgensis Curran, 1932, Nyt Mag. Naturvidensk., vol. 71, p. 356. Floreana
pusio Loew, 1872, Berliner Ent. Zeitschr., vol. 16, p. 279. Isabela
Genus Cadrema
pallida Loew, 1865, Berliner Ent. Zeitschr., vol. 9, p. 184 (Hippelates). Baltra
Genus Oscinella
galapagensis Curran, 1932, Nyt Mag. Naturvidensk., vol. 71, p. 357. Floreana
Family MILICHIIDAE
Genus Desmometopa
m-nigrum Zetterstedt, 1848, Diptera Scand., vol. 7, p. 2743 (Agromyza).
Family TETHINIDAE
Genus Trixoscelis
costalis Coquillett, 1901, Proc. Washington Acad. Sci., vol. 3, p. 378 (Rhicnoessa). Isabela, Floreana, Española
Genus Tethina
insulans Curran, 1932, Nyt Mag. Naturvidensk., vol. 71, p. 359. Floreana
Family ODINIIDAE
Genus Odinia
williamsi Johnson, 1924, Zoologica, vol. 5, p. 90. Pinta, Rábida, Santa Cruz, Baltra, Santa Fé, Floreana

[page] 171

Family AGROMYZIDAE
Genus Cerodontha
dorsalis Loew, 1863, Berliner Ent. Zeitschr., vol. 7, P. 54 (Odontocera). Santa Cruz, Floreana
Family MUSCIDAE
Genus Xenocoenosia
devia Curran, 1934, Proc. Calif. Acad. Sci., ser. 4, vol. 21, P. 164. Santiago, San Cristóbal
Genus Fannia
Canicularis Linnaeus, 1761, Fauna Suecica, P. 1871 (Musca). Isabela
sp., Curran, 1934, Proc. Calif. Acad. Sci., ser. 4, vol. 21, P. 164 (? Pusio Wiedemann). San Cristóbal
Genus Ophyra
aenescens Wiedemann, 1830, Ausser. Zweifl. Insekten, vol. 2, P. 435 (Anthomyia). Fernandina, Isabela, Baltra, Floreana
setia Walker, 1849, List Diptera Brit. Mus., vol. 4, p. 956 (Anthomiya.)
Genus Synthesiomyia
nudiseta Wulp,1833, Tijf. v. Ent., vol. 26, p. 42 (Cytoneura.) Isabela, Floreana
brasiliana Brauer & Bergenstamm, 1893, Denkschr. Math. Nat. Kais. Akad. Wiss. Wien, vol. 60, P. 96.
Genus Lispe
sp., Curran, 1934, Proc. Calif. Acad. Sci., ser. 4, vol. 21, P. 165. Floreana
Genus Musca
domestica Linnaeus, 1758, Systema Naturae, ed. 10, P. 596. Baltra, Floreana
Genus Stomoxys
Calcitrans Linnaeus, 1758 Systema Naturae, ed. 10, P. 604. Baltra, San Cristóbal, Floreana
Genus Morellia
Ochricornis Wiedemann, 1830, Aussereur. Zweifl. Insekten, vol. 2, P. 408. Floreana
Family CALLIPHORIDAE
Genus Viridinsula
Pionia Walker, 1849, List Diptera Brit. Mus., vol. 4, P. 880 (Musca). Santa Cruz, Floreana

[page] 172

deceptor Curran, 1934, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 166.
pionia Curran (not Walker), 1932, Nyt Mag. Naturvidensk., vol. 71, p. 361.
Isabela, Seymour, Floreana, Española
Genus Cochliomyia
macellaria Fabricius, 1775, Systema Ent., p. 776.
phanda Walker, 1849, List Diptera Brit. Mus., vol. 4, p. 869 (Musca).
quadrisignata Thomson, 1869, Freg. Eugenies Resa, Zool., vol. 1, p. 554 (Lucilia).
Isabela, Santa Cruz, Floreana
Family SARCOPHAGIDAE
Genus Opsophytopsis
inoa Walker, 1849, List Diptera Brit. Mus., vol. 4, p. 832 (Sarcophaga).
insularis Townsend, 1917, Insecutor Inscitae Mens., vol. 5, p. 163.
Darwin, Fernandina, Isabella, Santiago, Santa Cruz, San Cristóbal, Floreana, Española
Genus Sacrophagula
occidua Fabricus, 1794, Ent. Syst., p. 315 (Musca). Santiago, San Cristóbal, Floreana
Genus Prosthetocirca
cana Townsend, 1917, Proc. Biol. Soc. Washington, vol. 30, p. 196. Santa Cruz, Española
Genus Sarcophaga
isla Curran, 1934. Proc. Calif. Acad. Sci., 4th ser., vol. 21, p. 169, fig. 3. Española, Gardner
plinthopyga Wiedemann, 1830, Aussereur. Zweiflug. Insekten, vol. 2, p. 360.
quadriforceps Curran, 1932, Nyt Mag. Naturvidensk., vol. 71, p. 363.
Floreana
reversa Aldrich, 1916, Sarcophaga and Allies, p. 127. Baltra
taitensis Schiner, 1868, Novara Reise, Diptera, p. 314.
obtusifrons Thomson, 1869, Freg. Eugenies Resa, Zool., vol. 1, p. 536.
Isabella, Pinzón, San Cristóbal, Floreana
violenta Walker, 1849, List Diptera Brit. Mus., vol. 4, p. 826.
galapagensis Townsend, 1917, Proc. Biol. Soc. Washington, vol. 30, p. 195.
Isabella, Santa Cruz, Baltra, Santa Fé
williamsi Curran, 1934, Proc. Calif. Acad. Sci., 4th ser., vol. 21, p. 170, fig. 4. Floreana

[page] 173

Family TACHINIDAE
Genus Galapagosia
minuta Curran, 1934, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 172. Santa Cruz
Family HIPPOBOSCIDAE
Genus Olfersia
aenescens Thomson, 1869, Freg. Eugenies Resa, Zool., vol. 1, p. 610.
diomedae Coquillett, 1901, Proc. Washington Acad. Sci., vol. 3, p. 379 (Pseudolfersia).
Isabela, Santa Cruz, Española
fossulata Macquart, 1843, Mem. Soc. Sci. Lille, 1842, p. 434. Wolf, Daphne Major
spinifera Leach, 1817, on genera and species Eprobosc. Ins., p. 11, pl. 26 (1–3); 1818, Mem. Wernerian Nat. Hist. Soc., vol. 2, p. 557, pl. 24, figs. 1–3. Genovesa, Floreana
Genus Lynchia
nigra Perty, 1833, Delectus Anim. Artic. Brasil, vol. 3, p. 190, pl. 37, fig. 15 (Hippobosca).
intertropica Walker, 1849, List Diptera Brit. Mus., vol. 4, p. 1144 (Ornithomyia).
americanus, Johnson (not Leach), 1924, Zoologica, vol. 5, p. 91Ornithoponus).
Santa Cruz
albipennis Say, 1823, Jour. Acad. Nat. Sci. Philadelphia, vol. 3, p. 101 (Olfersia).
intertropicus, Johnson (not Walker), 1924, Zoologica, vol. 5, p. 91 (Ornithoponus).
Genovesa, Fernandina, Santiago, Santa Cruz, Seymour
Genus Microlynchia
galapagoensis Bequaert, 1955, Entom. Americana, vol. 35, p. 384. Santa Cruz, Española
pusilla Speiser, 1902, Zeitschr. Syst. Hym. Dipt., vol. 2, p. 157 (Lynchia). Española

Order HYMENOPTERA

The leading references on Hymenoptera other than ants are by Rohwer (1924) and Williams (1926). Formicidae were treated by Wheeler (1919 and 1924) and by Stitz (1932). Dr. William L. Brown kindly reviewed the present manuscript and made valuable corrections and proposed new synonymy.

Family CHALCIDIDAE
Genus Chalcis
sp., Rohwer, 1924, Zoologica, vol. 5, p. 175. Baltra

[page] 174

Family BETHYLIDAE
Genus Scleroderma
galapagense Brues, 1919, Proc. Calif. Acad. Sci., ser. 4, vol. 2, p. 309. Santiago
Family THYNNIDAE
Genus Agriomyia
vagans Smith, 1877, Proc. Zool. Soc. London, 1877, p. 83. Floreana
Family MUTILLIDAE
Genus Photopsis
galapagensis Williams, 1926, Proc. Calif. Acad. Sci., ser. 4, vol. 18, p. 350. Baltra
Family FORMICIDAE
Genus Cylindromyrmex
striatus tibialis Stitz, 1932, Nyt Mag. f. Naturvidensk., vol. 71, p. 367. Floreana
williamsi Wheeler, 1924, Zoologica, vol. 10, p. 101, fig. 19b, c.
striatus Wheeler (not Mayr, 1870), 1919, Proc. Calif. Acad. Sci., ser. 4, vol. 2, p. 266.
Santa Cruz, Baltra
Genus Hypoponera
beebei Wheeler, 1924, Zoologica, vol. 10, p. 107, fig. 21. Genovesa
Genus Odontomachus
bauri Emery, 1891, Ann. Soc. Ent. France, vol. 60, p. 591.
haematoda Stitz (not Linnaeus), 1932, Nyt Mag. Naturvidensk., vol. 71, p. 369.
San Cristóbal, Floreana
Genus Pheidole
sp., Wheeler, 1919, Proc. Calif. Acad. Sci., ser. 4, vol. 2, p. 271. Isabela
williamsi seymourensis Wheeler, 1924, Zoologica, vol. 5, p. 108. Baltra, Daphne
williamsi williamsi Wheeler, 1919, Proc. Calif. Acad. Sci., ser. 4, vol. 2, p. 267. Santa Cruz
Genus Monomorium
floricola Jerdon, 1851, Madras J. Litt. Sci., vol. 17, p. 107 (Atta). Genovesa

[page] 175

*floreanum Stitz, 1932, Nyt Mag. Naturvidensk., vol. 71, p. 368, figs. 1, 2. Floreana
pharaonis Linnaeus, 1758, Systema Naturae, ed. 10, vol. 1, p. 580 (Formica). Pinta, Santa Cruz
Genus Solenopsis
geminata galapageia Wheeler, 1919, Proc. Calif. Acad. Sci., ser. 4, vol. 2, p. 272. Floreana
globularia pacifica Wheeler, 1919, Proc. Calif. Acad. Sci., ser. 4, vol. 2, p. 272. Genovesa, Isabela, Daphne, Santa Cruz
globularia rubida Wheeler, 1919, Proc. Calif. Acad. Sci., ser. 4, vol. 2, p. 273. Española
saevissima F. Smith, 1855, Trans. Ent. Soc. London, 1855, p. 166, pl. 13, fig. 18 (Myrmica). Santa Cruz
Genus Tetramorium
guineense Fabricius, 1793, Ent. Syst., vol. 2, p. 357 (Formica). Genovesa, Isabela, Santa Cruz, San Cristóbal
simillimum F. Smith, 1851, List Hymenoptera Brit. Mus., vol. 6, p. 118 (Myrmica). Santiago, Floreana
Genus Crematogaster
brevispinosa chathamensis Wheeler, 1933, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 58 (subgenus Orthocrema). San Cristóbal
Genus Conomyrma
pyramica albemarlensis Wheeler, 1919, Proc. Calif. Acad. Sci., ser. 4, vol. 2, p. 275. Isabela, Rábida, Eden, Santa Cruz, Baltra
Genus Tapinoma
melanocephalum Fabricius, 1793, Ent. Syst., vol. 2, p. 353 (Formica). Genovesa, Santa Cruz, San Cristóbal, Española
Genus Paratrechina
fulva nesiotis Wheeler, 1919, Proc. Calif. Acad. Sci., ser. 4, vol. 2, p. 277 (Prenolepis). Isabela, Santiago, Santa Cruz, Española
longicornis Latreille, 1802, Hist. Nat. Fourmis, p. 113 (Formica). Santa Cruz, San Cristóbal, Floreana, Española, Gardner
vividula edenensis Wheeler, 1924, Zoologica, vol. 5, p. 111 (Prenolepis v. guatemalensis variety). Eden

* New Synonymy, fide W. L. Brown, 1964.

[page] 176

vividula itinerans Forel, 1901, Mitt. naturh. Mus. Hamburg, vol. 18, p. 81 Santa Cruz, San Cristóbal
Genus Camponotus
Subgenus Pseudocolobopsis
macilentus albemarlensis Wheeler, 1919, Proc. Calif. Acad. Sci., ser. 4, vol. 2, p. 284. Isabela
macilentus altinota Stitz, 1932, Nyt Mag. Naturvidensk., vol. 71, p. 370, fig. 3. Floreana
macilentus barringtonensis Wheeler, 1919 Proc. Calif. Acad. Sci., ser. 4, vol. 2, p. 282. Santa Fé
macilentus bindloensis Wheeler, 1919, Proc. Calif. Acad. Sci., ser. 4, vol. 2, p. 286. Marchena
macilentus castellanus Wheeler, 1924, Zoologica, vol. 10, p. 116, fig. 25. Genovesa
macilentus duncanensis Wheeler, 1919, Proc. Calif. Acad. Sci., ser. 4, vol. 2, p. 283. Pinzón, Floreana
macilentus hoodensis Wheeler, 1919, Proc. Calif. Acad. Sci., ser. 4, vol. 2, p. 285. Española
macilentus jacobensis Wheeler, 1919, Proc. Calif. Acad. Sci., ser. 4, vol. 2, p. 280. Santiago
macilentus macilentus F. Smith, 1877, Proc. Zool. Soc. London, 1877, p. 83. Floreana
macilentus narboroensis Wheeler, 1919, Proc. Calif. Acad. Sci., ser. 4,` vol. 2, p. 286. Fernandina
macilentus pervicus Wheeler, 1924, Zoologica, vol. 10, p. 115. Santa Cruz
macilentus sapphirinus Wheeler, 1924, Zoologica, vol. 10, p. 114, fig. 24. Santa Cruz, Baltra
macilentus vulcanalis Wheeler, 1919, Proc. Calif. Acad. Sci., ser. 4, vol. 2, p. 284. Isabela
macilentus wollebaeki Stitz, 1932, Nyt Mag. Naturvidensk., vol. 71, p. 371, fig. 4. Floreana
Subgenus Myrmocladoecus
planus fernandinensis Wheeler, 1919, Proc. Calif. Acad. Sci., ser. 4, vol. 2, p. 296. Fernandina
planus fidelis Wheeler, 1919, Proc. Calif. Acad. Sci., ser. 4, vol. 2, p. 295. Santa Fé
planus hephaestus Wheeler, 1933, Proc. Calif. Acad. Sci., ser. 4, vol. 21, p. 59. Isabela
planus indefessus Wheeler, 1919, Proc. Calif. Acad. Sci., ser. 4, vol. 2, p. 294. Santa Cruz

[page] 177

planus isabelensis Wheeler, 1919, Proc. Calif. Acad. Sci., ser. 4, vol. 2, p. 293. Isabela
planus peregrinus Emery, 1893, Ann. Soc. Ent. France, vol. 63, p. 91. San Cristóbal, Floreana
planus pinzonensis Wheeler, 1919, Proc. Calif. Acad. Sci., ser. 4, vol. 2, p. 297. pinzón
planus planus F. Smith, 1877, Proc. Zool. Soc. London, 1877, p. 83. Floreana
planus sansalvadorensis Wheeler, 1924. Zoologica, vol. 10, p. 119, fig. 27. Santiago
planus santacruzensis Wheeler, 1919, Proc. Calif. Acad. Sci., ser. 4, vol. 2, p. 294. Santa Cruz, Baltra
Family VESPIDAE
Genus Odynerus
galapagensis Williams, 1926, Proc. Calif. Acad. Sci., ser. 4, vol. 2, p. 353 (subgenus Pachyodynerus). San Cristóbal
Family POMPILIDAE
Genus Aporinellus
galapagensis Rohwer, 1924, Zoologica, vol. 5, p. 174. Baltra
Family SPHECIDAE
Genus Nitela
darwini Turner, 1916, Ann. Mag. Nat. Hist., ser. 8, vol. 18, p. 345. Floreana
Genus Tachysphex
galapagensis Rohwer, 1924, Zoologica, vol. 5, p. 173. Isabela, Baltra
Family APIDAE
Genus Xylocopa
darwini Cockerell, 1926, Ann. Mag. Nat. Hist., ser. 9, vol. 17, p. 659.
mordax Smith (not Smith, 1874), 1877, Proc. Zool. Soc. London, 1877, p. 83.
brasilianorum Maidl (not Linnaeus, 1767), 1912, Ann. K. K. Naturhist. Hofmus., vol. 26, p. 312.
transitoria Brues (not Perez, 1901), 1924, Zoologica, vol. 5, p. 125.
colona Rohwer (not Lepeletier, 1841), 1924, Zoologica, vol. 5, p. 173.
Isabela, Santiago, Pinzón, Santa Cruz, Baltra, San Cristóbal, Floreana, Gardner

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ANNOTATED BIBLIOGRAPHY OF GALÁPAGOS ENTOMOLOGY

AGASSIZ, A.
1892. General sketch of the expedition of the Albatross from February to May, 1891. Bulletin of the Museum of Comparative Zoology, vol. 23, pp. 1–89.
ALEXANDER, C. P.
1962. The crane flies of the Galapagos Islands (Tipulidae, Diptera). Opuscula Zoologica, vol. 61, pp. 1–5, 6 figs. [Limonia (Dicranomyia) galapagoensis, n. sp., (Santa Curz Island) (p. 1); Limonia (Geranomyia) tibialis (Loew), (Santa Cruz and San Cristobal islands) (p. 3); Erioptera (Trimicra) pilipes (Fabricius, var., (San Cristobal Island) (p. 4).]
ANONYMOUS
1901. Papers from the Hopkins Stanford Galapagos Expedition, 1898–1899. I. Introduction. Proceedings of the Washington Academy of Science, vol. 3, pp. 363–364.
ASAHINA, S.
1961. Dragonflies taken by Dr. Sekiguchi in the Galapagos Islands. Publication of the Entomological Laboratory, college of Agriculture, University of Osaka Sakai, no. 6, pp. 1–3, 1 fig.
BANKS, N.
1901. Papers from the Hopkins Standford Galapagos Expedition, 1898–1899. V. Entomological Results (5): Thysanura and Termitidae. Proceedings of the Washington Academy of Science, vol. 3, pp. 541–546. [Thysanura: Lepisma galapagoensis, n. sp. (p. 541, figs. 47–50), (Hood Island); L. insularis, n. sp. (p. 543, figs. 51–52), (Narborough and Albemarle islands); Termitidae: calotermes galapagoensis, n. sp. (p. 544, figs. 55–56), (Wenman Island); Calotermes pacificus, n. sp. (p. 545, fig. 57), (Albemarle and Narborough islands).]
1924. Neuroptera from the Williams Galapagos Expedition. Zoologica, vol. 5, pp. 117–180. [4 species in 3 families (Myrmeleonidae, Chrysopidae, Hemerobüdae): Myrmeleon perpilosus, n. sp. (Indefatigable Island); Chrysopa nigripilosa, n. sp. (South Seymour Island); Chrysopa galapagoensis, n. sp. (South Seymour Island); Megalomus darwini, n. sp. (Indefatigable Island).]
BANNING, G. H., and J. S. BARTH
1933. Hancock expedition of 1933 to the Galapagos Islands. Bulletin of the Zoological Society of San Diego, vol. 10, pp. 1–30. [Popular account of the expedition.]
BARBER, H.G.
1925. Hemiptera-Heteroptera from the Williams Galapagos Expedition. Zoologica, vol. 5, pp. 241–254. [8 new species in 6 families; total of 16 species listed—Harmostes disjunctus, n. sp. (Indefatigable Island); Heraeus pacificus, n. sp. (James Island); Creontiades juscosus, n. sp. (Indefatigable and James islands); Orthaea insularis, n. sp. (James, S. Seymour, Albemarle, and Indefatigable islands); Psallus insularis, n. sp. (James Island); Repipta annulipes, n. sp. (Indefatigable Island); Corythaica renormata, n. sp. (Daphne Major Island); Halobates robustus, n. sp., Indefatigable Island).]
1934. The Norwegian Zoological Expedition to the Galapagos Islands 1925, conducted by Alf Wolleback. XI Hemiptera-Heteroptera. Nyt Magazin for Naturvidenskaberne, vol. 74, pp. 281–289. [Reprinted as Meddelelser fra det Zoologiske Museum, Oslo, nr. 42, pp. 281–289 (1934).] [40 species recorded including Aknisus galapagensis, n. sp., (Chatham Island) (Neididae).]
BARLOW, NORA (editor)
1934 Charles Darwin's Diary of the Voyage of HMS "Beagle." Cambridge University Press, xxx + 442 pp. Map.

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1946. Charles Darwin and the Voyage of the Beagle, unpublished letters and notebooks, 279 pp. The Philosophical Library, New York.
BEEBE, WM.
1923. Williams Galapagos Expedition. Zoologica, vol. 5, pp. 1–20, 10 figs. 1 map.
1923. Notes on Galapagos Lepidoptera. Zoologica, vol. 5, pp. 51–59.
1924. Galapagos, World's End. C. P. Putnam's Sons, New York and London, xxi + 433 pp., illus.
1925. Logging the Arcturus. Bulletin of the New York Zoological Society, vol. 28, pp. 103–116.
BEEBE, WM., and RUTH ROSE
1925. The second half of the Arcturus Adventure. Bulletin of the New York Zoological Society, vol. 28, pp. 119–129, 4 col. plates.
BEQUAERT, J. C.
1933. The Templeton Crocker Expedition of the California Academy of Sciences, 1932. No. 11. The Hippoboscidae of the Galápagos Archipelago (Notes on the Hippoboscidae. 8), with an Appendix on the Tabanidae. Proceedings of the California Academy of Sciences, ser. 4, vol. 21, pp. 131–138. [3 genera: Olfersia, three species; Lynchia, two species; and Microlynchia, one species; no new species.]
1955. The Hippoboscidae or louse-flies (Diptera) of mammals and birds. Part II. Taxonomy, evolution and revision of American genera and species. Entomologica Americana, vol. 35, pp. 233–416, 38 figs. [Microlynchia galapagoensis, n. sp., (Indefatigable and Hood islands). (p. 384, figs.).]
BLAIR, K. G.
1928. Coleoptera (Heteromera, Teredilia, Malacodermata and Bruchidae) from the Galápagos Islands, collected on the St. George Expedition, 1924. The Annals and Magazine of Natural History, ser. 10, vol. 1, pp. 671–680.
1933. Further Coleoptera from the Galápagos Archipelago, The Annals and Magazine of Natural History, ser. 10, vol. 11, pp. 471–487. [New records and/or new species in Carabidae, Hydrophilidae, Histeridae, Coccinellidae, Languridae, Dermestidae, Scarabaeidae, Bostrychidae, Lycidae, Cantharidae, Cleridae, Elateridae, Tenebrionidae, Cistelidae, Oedemeridae, Cerambycidae, Chrysomelidae, Curculionidae, and Scolytidae.]
BOHEMAN, C. H.
1858–1859 Coleoptera. Species novas descripsit. in: Kongliga Svenska Fregatten Eugenies resa omkring Jorden… Zoologi I, Insecta, pp. 1–218, 2 pls. [7 new species from Galápagos: Feronia insularis, Copris lugubris, Heterocrepedius puberulus, Tessaroma morio, Phaleria manicata, and Eburia amabilis.]
BORGMEIER, T.
1960. Gefluegelte and ungefluegelte Phoriden aus der neotropischen Region, nebst Beschreibung von sieben neuen Gattungen (Diptera, Phorida). Studia Entomologica (new series), vol. 3, pp. 257–374, figs. [Dohrniphora cornuta (Bigot) (p. 277).]
BOWMAN, R. I.
1961. Morphological Differentiation and Adaptation in the Galápagos Finches. University of California Publications in Zoology, vol. 58, pp. iv + 302, 22 pls., 74 text figs. [Food habits of birds.]
BRAUER, F., and Y. E. VON BERGENSTAMM
1892. Die Zweiflugler des Kaiserlichen Museums zu Wien. v. Vorarbeiten zu einer Monographie der Muscaria schizometopa (Exclusive Anthomyidae). Pars II.

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Denkschriften der Kaiserlichen Akademic der Wissenschaften, vol. 58, pp. 305–446, Wien.
BREUNING, S.
1927. Monographic der Gattung Calosoma Web. (Carab.) Koleopterologische Rundschau, vol. 13, pp. 129–232. [Calosoma galapageum Hope (pp. 200–201); synonymy, distribution (Galápagos Islands and adjacent coast of South America).]
BRINCK, P.
1951. Anoplura from the Galápagos Islands. Nyt Magazin Naturvidenskaberne, vol. 88, pp. 246. [Polyplax spinulosarecorded from Epimys rattus alexandrinus on Floreana Island.]
BRUES, C. T.
1919. A new species of the Hymenopterous genus Scleroderma from the Galápagos Islands. Proceedings of the California Academy of Sciences, ser. 4, vol. 2, pp. 309–310.
1924. Triungulin larvae from the Williams Galápagos Expedition. Zoologica, vol. 5, pp. 125–136, figs. 28–32. [Larvae of a meloid bettle (presumably Cissites maculata (Swederer) from carpenter bee (Xylocopa) captured on South Seymour Island.]
BURNS, J. M.
1964. Evolution in skipper butterflies of the genus Erynnis. University of California Publications in Entomology, vol. 37, pp. 1–124, 1 pl. [p. 177, corrects Van Duzee's 1933 record of Thanaos persius to Erynnis funeralis Scudder and Burgees.]
BUTLER, A.G.
1877. Account of Zoological Collection made during the visit of H.M.S. "Peterel" to the Galapagos Islands, X. Lepidoptera, Orthoptera, Hemiptera. Proceedings of the Zoological Society of London, 1877, pp. 86–91.
CALVERT, P.P
1901–1908. Insecta. Neuroptera. Odonata. Biologia Centrali-Americana, pp. xxx+17–419, 9 pls., 1 map. [Anomalagrion hastatum(p. 130); Tramea darwini (p. 303); Pantala flavescens (p.307); Pantala hymenea(p. 309); Cannacria batesii (=Cannacria fumipennis) (p. 326).]
1947. The Odonate collections of the California Academy of Sciences from Baja California and Tepic, Mexico, of 1889–1894. Proceedings of the California Academy of Sciences, ser. 4, vol. 23, pp. 603–609.
1947. Odonata of Voyages under the auspices of the New York Zoological Society. Entomological News, vol. 58, pp. 227–230. [Records Pantala flavescens (Fabricius) (Indefatigable and South Seymour islands); Tramea cophysa darwini Kirby (Hood and Albemarle islands).]
1952. New taxonomic entities in Neotropical Aeshnas (Odonata: Aeshnidae). Entomological News, vol. 63, pp. 253–264. [Neureclipa galapagoensis (p. 258).]
1956. The Neotropical species of the "subgenus Aeshna" sensu Selysii 1883 (Odonata). Memoris of the American Entomological Society, vol. 15, pp. 1–251, 47 pls., 19 tables, maps 1–251, 47 pls., 19 tables, maps I–IV. [Aeshna (Neureclipa) galapageonsis, p. 137, pl. 17.]
COMPOS, R. FRANCISCO
1932. Estudio sobre una pequeña coleccion de Insectos de Puna Vieja y Archipielago de Galápagos. Revista del Colegio Nacional Vicente Rocafuerte. Ciño XIV, Núm. 46–47, pp. 9–14. [Lists 53 species from Puna Vieja and the Galápagos. The following are the Galápagos species: Sphingonotus fusco-irroratus Stål (Puna and Chatham islands); Calosoma howardi Linell (Chatham Island);

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Tropisternus lateralis Fabricius (Chatham Island); Dermestes carnivorusFabricius (Chatham Island); Pseuderyctes galapagoenis Linell (Chatham Island); Stomion galapagonese Waterhouse (Chatham Island); Gnathocerus cornulus Fabricius (Chatham Island); Acanthoderes galapageonsis Linell (Chatham Island); Phlegethontius rusticavar. galapagensis Holland (Santa Cruz Island); Celerio lineata Fabricius (Chatham Island).]
CANDEZE, E. C. A.
1859. Monographic des Elaterides. 2. Mémoires de la Société Royale des Sciences de Liége, vol. 14, pp. 1–543, pls. 1–7. [Physorhinus galapagoensis (p. 394).]
CARRIKER, M. A., JR.
1957. Notes on some of the Vernon L. Kellogg types of Mallophaga. Microentomology, vol. 22, pp. 95–110. [Corrects generic assignments of two Galápagos species.]
CARVALHO, J. C. M
1957–1960. A Catalogue of the Miridae of the World. Arquivos do Museu Nacional, Rio de Janeiro, Brazil. vol. 44, 158 pp. (part 1); vol. 44, 158 pp. (part 1); vol. 45, 216 pp. (part 2); vol. 47, 161 pp. (part 3); vol. 48, 384 pp. (part 4); vol. 51, 194 pp. (part 5). [Lists Galápagos species with new generic assignment for Capsus nigritulusWalker.]
CARVALHO, J. C. M., and E. WAGNER
1957. A world revision of the genus Trigontylus Fieber. Arquivos do Museu Nacional, Brazil, vol.43, pp. 121–155. [Miris lineatus Butler treated on p. 135; recorded from Seymour Island.]
CAUDELL, A. N.
1932. Insects of the order Orthoptera of the Pinchot Expedition of 1929. Proceedings of the United States National Museum, vol. 80, pp. 1–7. [8 species listed from the Galápagos Islands.]
CHAMPION, G. C.
1918. Notes on various South American Coleoptera collected by Charles Darwin during the voyage of the "Beagle," with descriptions of new genera and species. The Entomologist's Monthly Magazine, vol. 54, pp. 43–55.
1924. The insects of the Galapagos Islands. The Entomologist's Monthly Magazine, vol. 60, pp. 259–260. [Review of entomology in Beebe (1924) and the paper by Butler (1877b); generic placements by W. E. China.]
CHEESMAN, L. E.
1929. Hymenoptera collected by the "St. George" Expedition in Central America and the W. Indies. Transactions of the Entomological Society of London, vol. 77, pp. 141–154. [Xylocopa darwini: (Albemarle, Charles, Indefatigable, and James islands) (pp. 143–144, figs. 2a, b).]
CLARK, B.P.
1926. A revision of the Protoparce of the Galápagos Islands. Proceedings of the New England Zoology Club, vol. 9, pp. 67–71. [Recognizes: Protoparce calapagensis Holland; Protoparce nigrita Rothschild and Jordan; Protoparce rustica postscripta n. subsp. (Charles Island) (p. 70); and Protoparce leucoptera Rothschild and Jordan.]
CLARKE, J. F. GATES
1928. Catalogue of the Type specimens of Microlepidoptera in the British Museum (Natural History) described by Edward Meyrick. London, British Museum, vol. III, 600 pp. [Plateprota colobota, p.179. Crocidosema plebeiana, p. 319.]
CLAY, THERESA
1958. Revisions of Mallophaga genera. Degeeriella from the Falconiformes. Bulletin of the British Museum (Natural History), vol. 7, no. 4, pp. 123–207, 9 pls. [Synonymy of Degeeriella regalis, p. 186.]

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COCKERELL, T. D. A.
1926. Descriptions and records of bees.—CXI. The Annals and Magazine of Natural History, ser. 9, vol. 17, pp. 657–665. [Xylocopa darwini, n. sp., ♀, (Chatham Island) (pp. 659–660).]
1935. The Templeton Crocker Expedition of the California Academy of Sciences, 1932. no. 28. The carpenter bees of the Galápagos Islands. Proceedings of the California Academy of Sciences, ser. 4, vol. 21, pp. 379–382. [Discussion of Xylocopa darwini; history, variation.]
COLLENETTE, C. L.
1925. Sea-Girt Jungles. The experiences of a naturalist with the "St. George" Expedition. London. Hutchinson & Co., pp. xiv + 275, map, plates.
CONDÉ, B.
1960. Présence des Diploures Campodéidés sur les Iles Galapagos. Bulletin du Museum d'Histoire Naturelle, Paris, ser. 2, vol. 32, pp. 172–176, 1 fig. [Lepidocampajuradoi seclusa, n. subsp. (p. 172), (Santa Cruz Island).]
COQUILLETT, D. W.
1901. Papers from the Hopkins Stanford Galapagos Expedition, 1898–1899. II. Entomological results (2): Diptera. Proceedings of the Washington Academy of Science, vol. 3, pp. 371–379. [Records of 35 species; Ceratopogon galapagensis, n. sp. (p. 372), (Albemarle Island); Pareuxesta (n. gen.) latifasciata, n. sp. (p. 376), (Tower Island); Pareuxesta obscura, n. sp. (p. 377), (Albemarle Island); Pareuxesta hyalinata, n. sp. (p. 377), (Albemarle Island); Ephydra gilvipes, n. sp. (p. 377), (Albemarle Island); Canace snodgrassü, n. sp. (p. 378), (Albemarle and Narborough islands); Rhicnoessa costalis, n. sp. (p. 378), (Albemarle Island); Pseudolfersia diomedeae, n. sp. (p. 379), (Albemarle Island).]
CURIO, E.
1964. Über das Nächtigen von Nocticanace galapagensis Curran (Diptera, Canaceidae). Zeitschrift für Tierpsychologie. Band 21, Heft 7, pp. 794–797.
CURRAN, C. H.
1932. [The Norwegian Zoological Expedition to the Galápagos Islands 1925, conducted by Alf Wollebaek. IV.] Diptera (excl. of Tipulidae and Culicidae). Nyt Magazin Naturvidenskaberne, vol. 71, pp. 347–366, 1 fig. [Reprinted as Medelelser fra det Zoologiske Museum, Oslo, nr. 30, pp. 347–366, 1 fig. (1932).] [Summary list of previously known species with many new records; keys to: species of Villa (p. 350), genera of Dolichopodidae (p. 352), species of Hippelates (p. 356); descriptions of 7 new species.]
1933. The Templeton Crocker Expedition of the California Academy of Sciences, 1932. no. 12. Diptera. Proceedings of the California Academy of Sciences, ser. 4, vol. 21, pp. 147–172, 4 figs. [Many records and the following new species: Pipunculus galapagensis (Albemarle, Gardner, Charles, and James islands); Mesogramma crockeri (Charles, Chatham, Albemarle, and Indefatigable islands); Euxesta galapagensis (Hood Island); paroxyna crockeri (Albemarle, Charles, James, S. Seymour, Jervis, Duncan, and Indefatigable islands); Sphaerocera galapagensis (Chatham Island); Procanace galapagensis (Albemarle Island); Scatella galapagensis (Chatham Island); Xenocoenosia devia (Chatham and James islands); Viridinsula deceptor (N. Seymour, Albemarle, Hood, and Charles islands); Sarcophaga isla (Gardner and Hood islands); Sarcophaga williamsi (Charles Island); Galapagosia minuta (Indefatigable Island).]

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CURRIE, R. P.
1901. Papers from the Hopkins Stanford Galapagos Expedition, 1898–1899. Entomological Results (3): Odonata. Proceedings of the Washington Academy of Science, vol. 3, pp. 381–389. [7 species recorded: Aeschna galapagoensis, n. sp. (p. 382, fig. 29, 30), (Chatham Island); Cannacria fumipennis (p. 387, figs. 31–34), (Albemarle Island).]
DALLAS, W. S.
1851–1852. List of the Specimens of Hemipterous Insects in the collection of the British Museum, London, Part I, pp. 1–368, II pls.; Part II, pp. 369–592, 4 pls. [Anasa obscura, Stenocephalus insularis, Nysius? marginalis, new species, all collected by C. Darwin.]
DARWIN, C.
1839. Journal and remarks. In: Fitzory, Narrative of the surveying voyages of His Majesty's Ships Adventure and Beagle, between the years 1826 and 1836, describing their examination of the southern shores of South America and the Beagle's circumnavigation of the globe. vol. 3, pp. xiv + 615. H. Colburn, London.
DISTANT, W. L.
1900. Undescribed genera and species belonging to the Rhynchotal family Pentatomidae. Transactions of the Entomological Society of London, 1900, pp. 163–174.
1904. Rhynchotal Notes—XXI. The Annals and Magazine of Natural History, ser. 7, vol. 13, pp. 194–206. [Revision of Walker types and description of new genus Dagbertus, pp. 203–204.]
1909. VIII—Rhynchotal Notes—XLVIII. The Annals and Magazine of Natural History, ser. 8, vol. 4, pp. 73–87. [Galapagosana, n. gen., p. 73, for Issus rostrifer Butler.]
D'ORCHYMONT, A.
1937 Sphaeridiini broméliadicoles neuveaux (Coleoptera: Hydrophilidae, Sphaeridiinae). The Annals and Magazine of Natural History, ser. 10, vol. 20, pp. 127–140, 2 figs. [Galapagodacnum, n. gen., for Coelostoma darwini Blair (p. 134).]
DOS PASSOS, C. F.
1964. A synonymic list of the Nearctic Rhopalocera. Lepidopterists' Society, Memoirs, vol. 1, pp. 1–145.
DRAKE, C. J.
1954. Synonymic data and description of a new saldid (Hemiptera). Occasional Papers, Museum of Zoology, University of Michigan, no. 553. 5 pp. [Salda rubromaculata Heidemann, a synonym of Salda sphacelata Uhler.]
EDWARDS, R. L.
1961. Studies of the Philopteridae (Mallophaga) from birds of the order Procellariformes. 1. The genus Halipeurus Thompson. Journal of Parasitology, vol. 47, pp. 125–157. [Lipeurus limitatus treated as synonym of Halipeurus diversus (Kellogg); Halipeurus pelagicus recorded from Galapagos, with synonymy.]
EMERY, C.
1891. Voyage de M. Ch. Allnaud dans le territoire d'Assinie. Formicides. Annales de la société Entomologique de France, 1891, pp. 553–574. [Champsomyrmex bauri, n. sp. (p. 561), Galápagos Islands.]
1893. Notice sur quelques Fourmis des iles Galápagos. Annales de la Société Entomologique de France, vol. 62, pp. 89–92. [5 species recorded, two described as new.]
ESBEN-PETERSEN, P.
1920. Revision of some of the type specimens of Myrmeleonidae, described by Navas

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and placed in the Vienna Museum. Annales de la Société Entomologique de Belgique, vol. 60, pp. 190–196. [Dimares formosa Banks (= Dimares nummatus Navas) (p. 191).]
1934. The Norwegian Zoological Expedition to the Galápagos Islands 1925, conducted by Alf Wolleback. XII. Neuropterous insects from Galápagos Islands. Nyt Magazin Naturvidenskaberne, vol. 74, pp. 291–294.[Reprinted as Meddeleser fra det Zoologiske Museum, Oslo, nr. 43, pp. 291–294 (1934).] [Chrysopa wollebacki, n. sp. (p. 291, figs. 1, 2, (Floreana Island); myrmelionid larva (p. 293, fig. 3), (Floreana Island).]
EVANS, W. H.
1952. A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part II (groups B, C, D). Pyrginae. Section 1. British Museum, London. v + 178 pp., 16 pls. [Urbanus dorantes galapagensis (Williams), new combination (p. 93).]
EWING, H. E.
1924. Bird-infesting Mallophaga collected by the Williams Galapagos Expedition. Zoologica, vol. 5, pp. 81–84, fig. 12. [Records for 8 species, including Esthiopterum nannopteri, n. sp. (p. 82, fig. 12), from Nannoptera harrisi, and Esthiopterum potens var. minor, new (p. 84), from Sula nebouxii.]
FENNAH, R. G.
1954. The higher classification of the family Issidae (Homoptera: Fulgoroidea) with descriptions of new species. Transactions of the Royal Entomological Society of London, vol. 105, pp. 455–474. [Key to genera of Acanoloniinae including Galapagosana Distant and Euthiscia Van Duzee, p. 472.]
FERRIS, G. F.
1916. Notes on Anoplura and Mallophaga, from mammals, with descriptions of four new species and a new variety of Anoplura. Psyche, vol. 23, pp. 97–120. [Hoplopleura quadridentata Ferris, not Neumann, from the Galápagos Islands.]
1921. Contributions toward a monograph of the sucking lice. Part II. Stanford University Publication, University series, Biological Sciences, ser. 2, vol. 2, pp. 59–133, figs. 33–89. [Hoplopleura nesoryzomydis, n. sp., (Galápagos Islands) (p. 90), based upon specimens previously reported as Hoplopleura quadridentata by Ferris (1916).]
1923. Contributions toward a monograph of the sucking lice. Part IV. Stanford University Publication, University series, Biological Sciences, ser. 2, vol. 4, pp. 183–270, figs. 119–172. [Polyplax spinulosa recorded from the Galapagos Islands (p. 188).]
1941. Atlas of Scale Insects of North America—S III–362. [Aspidiotus smilacis Kuwana, not Comstock, a misidentification for Melanaspis obtusa Ferris, n. sp.]
1951. The Sucking Lice. Memoirs of the Pacific Coast Entomological Society, vol. 1, pp. ix + 320. [Galápagos species Hoplopleura nesoryzomydis, p. 138; Hoplopleura quadridentata (Neumann) a valid species, p. 142; Polyplax spinulosa (Burmeister), p. 211.]
FLUKE, C. L.
1942. Revision of the Neotropical Syrphini related to Syrphus (Diptera, Syrphidae). American Museum Novitates, no. 1201, pp. 1–24, figs. [Allograpta splendens (Thomson) (p. 17, fig. 40).]
1956. Catalogue of the family Syrphidae in the Neotropical Region (Diptera). Revista Brasileira de Entomologia, vol. 6, pp. 193–268. [Allograpta splendens (Thomson) (p. 210); Baccha clavata (Fabricius) (p. 240).]

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FOLSOM, J. W.
1924. Apterygota of the Williams Galápagos Expedition. Zoologica, vol. 5, pp. 67–76, pls. 3–5. [Three species recorded: Heterolepisma intermedia, n. sp. (pp. 67–70, figs. 1–12) (Indefatigable and South Seymour islands); Acrotelsa galapagoensis (Banks), (pp. 71–73, figs. 13–20) (Indefatigable, South Seymour, Daphne Major, and James islands); Lepidocyrtus intermixus, n. sp., (pp. 75–76, figs. 27–29) South Seymour Island.]
FROESCHNER, R. C.
1960. Cydnidae of the Western Hemisphere. Proceedings of the United States National Museum, vol. 111, pp. 337–680. [Redescription and generic placement of Geotomus murinus Van Duzee in Dallasiellus, p. 616.]
GARTH, J. S.
1933. IV. Insects. In: Banning, G. H., Hancock Expedition of 1933 to the Galapagos Islands. Bulletin of the Zoological Society of San Diego, no. 10, May 1933, pp. 18–20. [Charles Island butterflies—Leptotes marina, Catopsilis eubule, Dion vanillae, Danaus menippe (Monarch), and Eudamus galapagensis; Xylocopa (Charles Island) (nesting in "the agava"), Amantis (Barrington Island), Halobates (north of Albemarle Island), flat flies on birds.]
GIFFORD, E. W.
1919. Expedition of the California Academy of Sciences to the Galápagos Islands, 1905–1906. XIII. Field notes on the Land Birds of the Galápagos Islands and of Cocos Island, Costa Rica. Proceedings of the California Academy of Sciences, ser. 4, vol. 2, pt. II, no. 13, pp. 189–258. [Insects as food of the birds.]
HEBARD, M.
1920. Expedition of the California Academy of Sciences of Galápagos Islands, 1905–1906. XVII. Dermaptera and Orthoptera. Proceedings of the California Academy of Sciences, ser. 4, vol. 2, pp. 311–346, pl. t. figs. 1–11. [Reports 1 species of Dermaptera (adventive); 9 species of blattids, all but one, Anisopygia snodgrassii (McNeill), adventive; 1 endemic mantid, Galapagia solitaria Scudder; 8 species of acridids, 3 dividing into geographic races, 1 also recorded from Puna Island, Ecuador; 4 species of tettigoniids, 3 endemic; 4 species of gryllids, 3 endemic.] [Halmenus eschatus, n. sp. (Wenman Island).]
1934. The Norwegian Zoological Expedition to the Galápagos Islands 1925, conducted by Alf Wollebaek. X. Orthoptera of the Galápagos Islands. Nyt Magazin Naturvidenskaberne, vol. 74, pp. 279–280. [Reprinted as Meddelelser fra det Zoologiske Museum, Oslo, nr. 41, pp. 279–280 (1934).] [Records of 11 species from Floreana and/or Chatham Island; 17 species and 4 races regarded as native; of cockroaches, 8 are adventive, one native.]
HEIDEMANN, O.
1901. Papers from the Hopkings Stnford Galápagos Expedition, 1898–1899, I Entomological Results (1): Hemiptera. Proceddings of the Washington Academy of Science, vol. 3, pp. 364–370. [Records for 21 species of Heteroptera and Homoptera; Ghilianella galapagensis, n. sp. (p. 367), (Hood and Albemarle islands); Salda rubromaculata, n. sp. (p. 368) (Albemarle Island).]
HERRING, J. L.
1961. The genus Halobates (Hemiptera: Gerridae). Pacific Insects, vol. 3, no. 2, pp. 223–305. [Halobates micans, Galápagos, p. 227; Halobates sobrinus, Galápagos, p. 229; Halobates robustus (Santa Cruz and Isabela islands), p. 229 and 282–283.]
HINTON, H. E.
1936 New species of Ataenius allied to Ataenius cribrithorax Bates (Coleoptera, scara-

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(baeidae). The Annals and Magazine of Natural History, ser. 10, vol. 17, pp. 413–428. [Ataenius arrowi, n. sp. (p. 414), Galápagos Islands.]
HOLLAND, W. J.
1889. Lepidoptera. In: Howard, L. O., Scientific results of explorations by the U. S. Fish Commission Steamer Albatross. no. V—Annotated catalogue of the insects collected in 1887–'88. Proceedings of the United States National Museum, vol. 12, pp. 194–196.
HOPE, F. W.
1838. Descriptions of some species of Carabidae, collected by Charles Darwin, Esq., in his late voyage. Transactions of the Entomological Society of London, vol. 2, pp. 128–131. [Calosoma galapageium, n. sp., Galápagos Islands.]
HOPKINS, G. H. E.
1951. xxxvi.—Stray notes on Mallophaga.—xi. The Annals and Magazine of Natural History, ser. 12, vol. 4, pp. 375–377. [Lectotypes designated for Brü species of Kellogg and Kuwana.]
HOPKINS, G. H. E., AND T. CLAY
1952. A check list of the genera and species of Mallophaga. British Museum (Natural History), London, 362 pp.
HORN, W.
1915. Coleoptera, Adephaga, Fam. Carabidae, Subfam. Cicindelinae. Genera Insectorum, fasc. 82c, pp. 209–486, pls. 16–23. [Cicindela galapagoensis, n. sp. (p. 52, et seq.).]
1920. Cicindelinen-Studien aus dem schwedischen Reichsmuseum nebst einigen Bemerkungen neuer Arten, etc. Archives fuuml;r Zoologi, vol. 13, no. 11, pp. 1–21. [Cicindela galapagoensis Horn (p. 17).]
HOWARD, L. O.
1890. Scientific results of explorations by Steamer Albatross, No. 5. Annotated catalogue of the insects collected in 1887"'88. Proceedings of the United States National Museum, vol. 12, pp. 185–216 (1889).
HURD, P. D., JR.
1955. Observations on the nesting habits of some New World carpenter bees with remarks on their importance in the problem of species formation. Annals of the Entomolgical Society of America, vol. 51, pp. 365–375, 5 figs. [Reference to the Galápagos carpenter bee, X ylocopa darwini Cockerell, as host of parasitic beetle Cissites maculata (Swederus) (p. 372).]
1958. The carpenter bees of the eastern Pacific oceanic islands. Journal of the Kansas Entomological Society, vol. 31, pp. 249–255. [X ylocopa darwini, summary of literature.]
HUTCHINSON, G. E.
1931. On the occurrence of Trichocorixa Kirkaldy (Corixidae, Hemiptera-Heteroptera) in salt water and its zoogeographic significance. The American Naturalist, vol. 65, pp. 573–574. [Cites a personal note from Dr. H. B. Hungerford that Trichocorixa had been received from the Galápagos Islands.]
JOHN, H.
1959. Neue Spezies der Gattungen Notiophygus Gory und Fallia Sharp (Ins., Col., Discolomidae = Notiophygidae). Senckenbergiana Biologie, vol. 40, pp. 289–296, 2 pls., 2 figs. [Fallia galapagana, n. sp. (p. 295, fig. 2) (Santa Cruz Island); Fallia colorata, n. sp. (p. 295, pl. 35, figs. 2a–2e), (Galápagos Islands).]
JOHNSON, C. W.
1924. Diptera of the Williams Galápagos Expedition. Zoologica, vol. 5, pp. 85–92, figs. 13–14. [28 species recorded, 13 for first time; new species: Galapagomya

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longipes (Indefatigable Island); Asyndetus versicolor (S. Seymour Island); Odinia williamsi (S. Seymour Island).]
KÉLER, S. VON
1957. Die Mallophaga von Sturmvögeln und Ruderfüssern. 1. Harrisoniella Bedford und Perineus Thompson (Mallophaga) (2. Fortsetzung). Beiträge zur Entomologie, vol. 7, pp. 493–527, 30 figs. [Perineus oblongus, n. sp. (p. 523, figs. 27–28) (on Diomedea irrorata) (Galápagos Islands).]
KELLOGG, V. L.
1901. Insects and spiders of the Galápagos Islands. Psyche, vol. 9, pp. 173–175.
1903. Two new genera of Mallophaga. Biological Bulletin, vol. 5, pp. 85–91.
1906. A second collection of Mallophaga from birds of the Galápagos and Revillagigedo islands and neighboring waters. Transactions of the American Entomological Society, vol. 32, pp. 315–324. [Records of 41 spp. from Galápagos Islands, including Nirmus ridgewayi, n. sp. (p. 317) (Indefatigable Island). and Menopon beckii, n. sp. (p. 322) (Daphne Island).]
KELLOGG, V. L., and S. I. KUWANA
1902. Papers from the Hopkins Stanford Galápagos Expedition, 1898–1899. Entomological Results (8): Mallophaga from birds. Proceedings of the Washington Academy of Science, vol. 4, pp. 457–499, pls. 28–31. [43 species reported, many described as new.]
KERNBACH, K.
1962. Die Schwärmer einiger Galapagos-Inseln (Lep. Sphingidae). Opuscula Zoologica, vol. 63, pp. 1–19, abb. 1–12. [12 species recorded, 1 described as new; 6 named as new subspecies of mainland forms.]
KIRBY, HAROLD, JR.
1939. The Templeton Crocker Expedition of the California Academy of Sciences, 1932. no. 39. Two new flagellates from termites in the genera Coronympha Kirby, and Metacaronympha Kirby, new genus. Proceedings of the California Academy of Sciences, ser. 4, vol. 22, pp. 207–220, 4 pls.
KIRBY, W. F.
1889. A revision of the subfamily Libellulinae, with descriptions of new genera and species. Transactions of the Zoological Society of London, vol. 12, pp. 249–348, pls. 51–57. [Tramea darwini, n. sp. (p. 315, pl. 51, fig. 1).]
1890. A synonymic catalogue of Neuroptera Odonata or dragonflies with an appendix of fossil species. London. pp. 1–202.
KÖHLER, P.
1961. Noctuidarum miscellanea III. Anales de la Sociedad Cientifica Argentina, Buenos Aires, vol. 172, pp. 69–94, 25 figs. [Agrotis (Scotia) galapagosensis, n. sp.]
KUSCHEL, G.
1963. Composition and relationship of the terrestrial faunas of Easter, Juan Fernandez, Desventuradas, and Galapagos islands. Occasional Papers of the California Academy of Sciences, no. 44, pp. 79–95, 1fig., 11 tab. [Includes climatological data for San Cristóbal and Seymour islands and comparison of Coleoptera of Galápagos and Juan Fernandez by families.]
KUWANA, S. J.
1902. Coccidae from the Galapagos Islands. Journal of the New York Entomological Society, vol. 10, pp. 28–33, 2 pls. [Removed from herbarium specimens collected by the Snodgrass-Heller Expedition; includes: Orthezia galapagoensis, n. sp. (p. 28); Lecanium hesperidum var. pacificum, n. var. (p. 30).]
LACK, DAVID
1945. The Galápagos Finches (Geospizinae), a study in variation. Occasional Papers

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of the California Academy of Sciences, no. 21, 152 pp., 4 pls. [Insects as food.]
LESNE, P.
1910. Notes sur les Coléoptères Térédiles. 4. Les Bostrchides des Iles Galápagos. Bulletin du Muséum National d'Histoire Naturelle, vol. 16, pp. 183–186. [Schisloccros cornutus galapaganus, n. subsp. (p. 184).]
LIGHT, S. F.
1935. The Templeton Crocker Expedition of the California Academy of Sciences, 1932. no. 20. The termites. Proceedings of the California Academy of Sciences, ser. 4, vol. 21, pp. 233–258, pls. 9–10, 10 t.figs. [5 species recorded: Kalotermesimmigrans Snyder (Chatham, Indefatigable, James, Albemarle, Jarvis, Narborough, South Seymour, and Tower islands); Kalotermes pacificus Banks (Albermarle and James islands); Kalotermes darwini, n. sp. (Albemarle, Charles and James islands); Heterolermes orthognathus Light (Indefatigable Island).]
LINELL, M. L.
1898. On the coleopterous insects of the Galápagos Islands Proceedings of the United States National Museum, vol. 21, no. 1143, pp. 249–268.
LINNAVUORI, R.
1959. Revision of the Neotropical Deltocephalinae and some related subfamilies (Homoptera). Annales Zoologici Societatis Zoologicae Botanicae Fennicae "Vanamo," vol. 20, 370 pp., 144 figs. Finnland. [Generic placement of Dellocephalus, Scaphytopius, Amplicephalus, Exetranus, and synonymy under Balclutha (= Eugnalhodus).]
LINSLEY, E. G.
1965. Pollinating insects of the Galápagos Islands. In: Bowman, R. I. (Ed.). Proceeding of the symposia of the Galápagos International Scientific Project. University of California Press. Berkely-Los Angeles (in press).
LINELEY, E. G., C. M. RICK, AND S. G. STEPHENS
1965. Observations on the floral relationships of the Galápagos carpenter bee Panpacific Enlomologist (in press).
MACSWAIN, J. W.
1961. Primary larvae of Cissites associated with New World carpenter bees. The Panpacific Entornologist, vol. 37, pp. 191–194.
MAA, T. C.
1963. Genera and species of Hippoboscidae (Diptera): types, synonymy, habitats and natural groupings. Pacific Insects Monograph 6, pp. 1–186. [Olfersia diomedeae (p. 29).1]
MAIDL, F.
1912. Die Xylocopen (Holzbienen) des Wiener Hofmuseums. Ein Beitrag zu einer Monographie dieser Gattung. Annalen des Kaiserlichen-königlichen Naturhistorischen Hofmuseums, vol. 26, pp. 249–330, tag. 3–4.[Xylocopa recorded from the Galápagos Islands (p. 312) (as Xylocopa brasilianorum).]
MARTIN, R.
1908. Collection zoologiques Edm. de Selys Longchamps. Fasc. 18, Aeschnines. [Aeschna galapagoensis (p. 50, fig. 47).]
MARX, F.
1890. ?Arachnida. In: Howard, Scientific results of explorations by Steamer Albatross, no. 5. Annotated catalogue of the insects collected in 1887–88. Proceedings of the United States National Museum, vol. 12, pp. 207–211.
MCLACHLAN, R.
1877. Account of the zoological collection made during the visit of H.M.S. Peterel to

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the Galapagos Islands. Neuroptera. Proceedings of the Zoological Society of London, 1877, pp. 84–86. [pantala hymenaea (say) and Tramea sp. [from Galápagos expedition of H.M.S. Peterel.]
MCNEILL, J.
1901. Papers from the Hopkins Stanford Galápagos Expedition, 1898–1899. Entomological Results (4): Orthoptera. Proceedings of the Washington Academy of Science, vol. 3, pp. 487–506. [Distribution of 26 known species summarized and occurrence by island analyzed: Temnopteryx snodgrasii, n. sp. (p. 493) (Albemarle Island); Liparoscelis cooksoni Butler (p. 497, fig. 38); Liparoscelis ensifera, n. sp.(p. 498, fig. 39) (Hood Island); Liparoscelis paludicola, n. sp. (p. 499, fig. 40) (Albemarle Island); Liparoscelis pallidus, n. sp. (p. 500, fig. 41) (Barrington Island); Xiphidium exitiosum, n. sp. (p. 501, fig. 42); Nemobius speculi, n. sp. (p. 503, figs. 43–44) (Albermarle Island); Cycloptilum lepismoide, n. sp. (p. 505) (Albemarle, Narborough, and Abington islands).]
MEISE, W.
1933. The Norwegian Zoological Expedition to the Galápagos Islands 1925, conducted by Alf Wollebaek. VIII. Scropiones. Nyt Magain Naturvidenskaberne, vol. 74, pp. 25–43, figs. [Reprinted as Meddelelser fra det Zoologiske Museum, Oslo, nr. 39, pp. 25–43, figs. (1933).] [Rhopalurus testaceus exsul, n. subp. (p. 26), Floreana Island.]
MELICHAR, L.
1906. Monographie der Issiden (Homoptera). Abendlungen der Kaiserlichen-Königlichen Zoologisch-Botanische Gesellschaft Wien, Vol. 3, no. 4, pp. 1–327, 75 figs. [Issus varius Walker transferred to Thionia.]
METCALF, Z. P.
1943. General Catalogue of the Hemiptera. Fasc. IV, part 3, pp. 552 [Catalogues Ilburnia simulans (Walker) p. 317; Delphacodes substitua (Walker), p. 518, and Delphacodes vicaria (Walker), p. 522.]
1954. General Catalogue of the Homoptera. Fasc. IV, pt. 14, pp. vii + 54. [Genus Philatis catalogued, pp. 43–45.]
1958. General Catalogue of Homoptera. Fasc. IV, part 15, pp. vii + 561. [Galapagosana catalogued on pp. 289–290.]
MEYRICK, E.
1926. On Microlepidoptera from the Galápagos Islands and Rapa. Transactions of the Enotomological Society of London, vol. 74, pp. 269–278. [11 species listed; one probably introduced, 10 represent endemic species in American genera.]
MORRISON, H.
1924. The Coccidae of the Williams Galapagos Expedition. Zoologica, vol. 5, pp. 143–152, figs. 33–37. [6 identified species recorded, all endemic; new species: Margarodes similis (South Seymour and Indefatigable islands); Eriococcus Papillosus (Indefatigable Island); Phenacoccus parvus (Tower Island); Pseudococcus galapagoensis (Eden and Indefatigable islands); Pseudococcus insularis (South Seymour Island).]
MUIR, F.
1919. Notes on the Delphacidae in the British Museum Collection. The Canadian Entomologist, vol. 51. pp. 6–8. [Places Walker's Delphax simulans is Ilburnia.]
1919. On the Genus Ilburnia White (Homoptera, Delphacidae). Proceedings of the Hawaiian Entomological Society, vol. 4, pp. 48–50. [Notes on Ilburnia simulans (Walker).]

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MUTCHLER, A. J.
1925. Coleoptera of the Williams Galápagos Expedition, Zoologica, vol. 5, pp. 219–240.
1938. Coleoptera of the Galápagos Islands. American Museum Novitates, no. 981, pp. 1–19.
NAVAS, R. P. LONGIN
1912. Myrméléonides (Ins. Né.) nouveaux oú peu connus. Annales de la Société Scientifique de Bruxelles, vol. 36, pp. 203–248. [Dimares nummatus, n. sp. (p. 229, fig. 16). (S. Elena Island, Galápagos).]
NEEDHAM, J. G.
1902. New dragon-fly nymphs in the United States National Museum. Proceedings of the United States National Museum, vol. 27, pp. 685–720, 7 pls. [Aeschna galapagoensis Currie (nymph) (p. 695, pl. 40, fig. 3).]
OSBORN, H.
1889. Mallophaga. In: Howard, L. O., Scientific results of explorations by the U. S. Fish Commission Steamer Albatross. No. V—Annotated catalogue of the insects collected in 1887–'88. Proceedings of the United States National Museum, vol. 12, pp. 188–189.
1924. Homoptera of the Williams Galapagos Expedition. Zoologica, vol. 5, pp. 77–79. [Four species recorded: Jassus galapagoensis, n. sp. (pp. 77–78) (Indefatigable and James islands); philatis productus Stål (p.78) (Tower Island); Philatis cinerea, n. sp. (pp. 78–79) (Tower Island); and Philatis major, n. sp. (Indefatigable Island).]
PALCT, J.
1956. Biologie der primär flügellosen Insekten. Jena. [Stylifer galapagoensis (Banks), n. comb. (p. 27).]
1959. Über die Lepismatidae (Ins. Thysanura) von den Galapagos-Inseln. Sencken-bergiana Bilogie, vol. 40, pp. 171–172. [Heterolepisma insulare (Banks), Stylifera galapagoensis (Banks), synonymy, distribution.]
PAPP, C. S.
1954. New record for Calosoma galapageium Hope (Coleoptera: Carabidae). Bulletin of the Southern California Academy of Sciences, vol. 53, pp. 169–174, pls. 45–46. [Original descriptions of the 4 nominal Galápagos species of Calosoma reproduced; Calosoma galapageium recorded from Chatham Island.]
PARENT, O.
1934. Disptères Dolichopodides exotiques. Mémoires de la Société Nationale des Sciences Naturelles et Mathématiques de Cherbourge, vol. 41, pp. 257–312, 13 pls. [Vanduzeeia cheesmani, n. sp. (p. 258) (Galápagos Islands).]
PETROVITZ, R.
1961. Neue und verkannte Apodiinae aus allen Erdteilen (Col. Scarab.) II. Entomologische Arbeiten, vol. 12, pp. 344–356, 5 figs. [Ataenius aequatorialis, n. sp. (p. 355) (Santa Cruz Island); Ataenius dampieri, n. sp. (p. 356) (Floreana Island).]
RICHARDS, A. G., JR.
1941. The noctuid moths of the Galapagos from the collections of the Allan Hancock Foundation. Report of the Allan Hancock Pacific Expedition, vol. 5, pp. 233–251, 4 pls.
RICHARDSON, H.
1901. Papers from the Hopkins Stanford Galapagos Expedition, 1898–1899. VI. The Isopods. Proceedings of the Washington Academy of Science, vol. 3, pp. 365–368.

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RICK, C. M.
1963. Biosystematic studies on Galápagos tomatoes. Occasional Papers of the California Academy of Sciences, no. 44, pp. 59–77. [Includes records of flower visits of Xylocopa darwini Cockerell (p. 63).]
1965. Some plant-animal relationships on the Galápagos Islands.
RIS, F.
1913. Libelulinen. Collections zoologiques du Baron Edm. de Selys Longchamps, Fasc. 15, pp. 837–964, pl. 7; 16, pp. 965–1042, pl. 8.
ROESCHKE, H.
1900. Carabologische Notizen V. Ebntomologische Nachrichten, vol. 26, pp. 57–63. [Calosoma galapageium Hope: notes on the type (♂) in the British Museum.]
ROHWER, S. A.
1924. Hymenoptera collected by the Williams Galápagos Expedition. Zoologica, vol. 5, pp. 173–175. [Four species recorded, including Tachysphex galapagensis, n. sp. (Albermarle Island), and Aporinellus galapagensis, n. sp. (South Seymour Island).]
ROTHSCHILD, W., and K. JORDAN
1903. A revision of the lepidopterous family Sphingidae. Novitates Zoologicae, ix supplement, pp. cxxxv + 972, 67 pls.
RUCKES, H.
1957. Three new species of Thyania Stål(Heteroptera: Pentatomidae). The Pan-Pacific Entomologist, vol. 33, pp. 175–180, 8 figs. [Thyanta setigera, n. sp.]
SCHAUS, W.
1923. Galapagos Heterocera, with descriptions of new species. Zoologica, vol. 5, pp. 23–48.
SCUDDER, G. G. E.
1957. The systematic position of Dicranocephalus Hahn, 1826, and its allies. Proceedings of the Royal Entomological Society of London, ser. A, vol. 32, pp. 147–158. [Discusses anomalous distribution in Galápagos—the only locality in the Western Hemisphere.]
SCUDDER, S. H.
1893. Reports on the dredging operations off the west coast of Central America to the Galápagos, to the west coast of Mexico, and in the Gulf of California, in charge of Alexander Agassiz, carried on by the U. S. Fish Commission Steamer "Albatross" during 1891, Lieut. Commander Z. L. Tanner, U.S.N., Commanding VII. The Orthoptera of the Galapagos Islands. Bulletin of the Museum of Comparative Zoology, vol. 25, pp. 1–25, pls. I–III.
SIGNORET, V.
1859. Monographie du genre Corizus. Annales de la Société Entomologique de France, ser. 3, vol. 28, pp. 75–105. [Corizus lugens, p. 92, n. sp. also described in Stål 1859, Kongliga Svenska Fregatten Eugenies Resa Omkring Jorden.… Zoologic 1, Insecta, p. 240.]
SLEVIN, J. R.
1931. Log of the Schooner "Academy," on a voyage of scientific research to the Galapagos Islands, 1905–1906. Occasional Papers of the California Academy of Sciences, no. 17, 162 pp., 16 pls., 1 map.
SMITH, F.
1877. Hymenoptera and Diptera. In: Gunther, Account of the zoological collection made during the visit of H.M.S. "Peterel" to the Galápagos Islands, VIII. Proceedings of the Zoological Society of London, 1877, pp. 82–84

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SNODGRASS, R, E,
1902. Papers from the Hopkins Stanford Galapagos Expedition, 1898–1899. VIII. Entomological Results (7): Schistocerca, Sphingonotus, and Halmenus. Proceedings of the Washington Academy of Science, vol. 4, pp. 411–454, pls. XXVI–XXVII.
STACH, J.
1932. The Norwegian Zoological Expedition of the Galápagos Islands 1925, conducted by Alf Wolleback. III. Die Apterygoten aus den Galápagos-Inseln. Nyt Magazin Naturvidenskaberne, vol. 71, pp. 331–346, pls 1–4. [Reprinted as Meddelelser fra det Zoologiske Museum, Oslo, nr. pp. 331–346, pls. 1–4 (1932).] [Records of 3 species of Thysanura: Lepisma insularis (Fernandina and Isabela islands), Heterolepisma intermedia (Santa Cruz, South Seymour, and Floreana islands), Acrotelsa galapagoensis (Española, Santa Cruz, Daphne Major, South Seymour, San Salvador, and Floreana islands); 2 species of Collembola: Anurida maritima (Floreana Island), Lepidocyrtus intermixtus (South Seymour Island).]
STÅL, C.
1859. Hemiptera. Species novas descripsit. Kongliga Svenska Fregatten Eugenies Resa Omkring Jorden, under befäl af C. A. Virgin, åren 1851–3. Zoologi 1, Insecta, pp. 219–298, 2 pls. [Arma sordida, Rhaphigaster viridans, Rhopalus lugens, Cymus galapagensis, Mycterodus productus, n. spp.]
1860 Orthoptera species novas descripsit. In: Kongliga Svenska Fregatten Eugenies Resa Omkring Jorden…Zoologi 1, Insecta, pp. 299–350, 1 pl. [Acridium melanocerum, Oedipoda fusco-irrorata, n. spp.]
STEPHAN, J.
1937. Schmetterlingsleben auf den Galapagos-Inseln. Entomologische Rundschau, vol. 54, pp. 165–168.
STITZ, H.
1932. [The Norwegian Zoological Expedition to the Galapagos Islands 1925, conducted by Alf Wollebaek. V.] Formicidae. Nyt Magazin Naturvidenskaberne, vol. 71, pp. 367–372, 4 figs. [Reprinted as Meddelelser fra det Zoologiske Museum, Oslo, nr. 31, pp. 367–372, 4 figs. (1932).] [Records for 10 species, including Cylindromyrmex striatus tibidis, n. var. (p. 367) (Floreana Island); Monomorium floreanum, n. sp. (p. 368) (Floreana Island); Camponotus macilentus altinota, n. var. (p. 370) (Floreana Island); Camponotus macilentus wollebaeki, n. var. (p. 371) (Floreana Island).]
STONE, A., and W. W. WIRTH
1947. On the marine midges of the genus Clunio Haliday (Diptera, Tendipedidae). Proceedings of the Entomological Society of Washington, vol. 49, pp. 201–224, 3 pls. [Clunio schmitti, n. sp. (p. 217) (Narborough Island).]
THOMSON, C. G.
1869. Diptera species novas descripsit. In: Kongliga Svenska Fregatten Eugenies Resa Omkring Jorden…Zoologi 1, Insecta, pp. 443–614, 1 pl., 1868. [Tabanus vittiger, Chrysochlora fasciata, Anthrax lateralis, Anthrax curvirostia, Anthrax tincta, Anthrax brachialis, Syrphus splendens, Baccha facialis, Anchineura tibialis, Myobia dasycnemis, Lucilia quadrisignata, and Sarcophaga obtusifrons, all new species.]
TIMMERMAN, G.
1952. The species of the genus Quadraceps (Mallophaga) from the Larinae, with some remarks on the systematics and the phylogeny of the gulls. Part 1. The Annals

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and magazine of Natural History, ser. 12, vol. 5, pp. 209–222. 1 pl., 6 figs. [Quadraceps ernstmayri, n.sp. (on Creagrus furcatus).]
1954. CI.—Studien über Mallophagen aus den Sammlugen des Britischen Museums (Natural History), London. The Annals and Magazine of Natural History, ser. 12, vol. 7, pp. 829–841, 1 pl., 3 figs.
1962. Neue zangenläuse (Mallophaga, Philopteridae) von procellariiformen und charadriiformen wirten. Zeitschrift für Parisitenkunde, vol. 21, pp. 426–436, 5 figs. [Docophoroides irroratae, n. sp., p. 435, Galápagos Islands.]
1963. Ein interresssanter neuer Kletterfederling vom Galapagos-Albatros. Bonner Zoologische Beiträge, vol. 14, pp. 157–164, 3 figs, [Docophoroides levequei, n. sp.]
TODD, E. L.
1959. The fruit-piercing moths of the genus Gonodonta Hübner (Lepidoptera, Noctuidae). Technical Bulletin of the United states Department of Agriculture. no. 1201, pp. 1–52, 12 pls. [Gonodonta iarmata galapagensis n. subsp. (p. 20, figs.) (Santa Cruz, San Cristóbal, Floreana islands).]
TOKUNAGA, MASAAKI
1964. Insects of Micronesia. Diptera. Chronomidæ. Bernice P. Bishop Museum. Insects of Micronesia, vol. 12, no. 5, pp. 485–628. [Galapagomyia Johnson. A synonym of Thalassomyia Schiner (p. 539).]
TOWNSEND, C. H.
1930. The Astor Expedition to the Galapagos Islands. Bulletin of the New York Zoological Society, vol. 33, pp. 135–172, 2 col. pl.
TOWNSEND, C. H. T.
1917. Genera of the dipterous tribe Sarcophagini. Proceedings of the Biological Society of Washington, vol. 30, pp. 189–198. [Prosthetocirca cana, n. sp. (Narborough and Albemarle islands); Gigantotheca galapagensis, n. sp. (Albemarle Island); Sarothromyiops cinctus, n. sp. (Culpepper Island).]
1917. New Genera of Amobiinae (Diptera). Insecutor Inscitae Menstruus, vol. 5, pp. 157–165. [Opsophytopsis (n. gen.) insularis, n. sp. (p. 163), (Isabela Island).]
1917. New genera and species of American muscoid Diptera. Proceedings of the Biological Society of Washington, vol. 30, pp, 43–50.
TURNER, R. E.
1916. Notes on fossorial Hymenoptera, XXIV, on the genus Nitela Latreille. The Annals and Magazine of Natural History, ser. 8, vol. 18, pp. 343–345.
UHLER, P. R.
1889. Hemiptera. In: L. O. Howard, Scientific Results of Explorations by the U. S. Fish Commission Steamer Albatross. no. V.—Annotated Catalogue of the Insects collected in 1877-'88. Proceedings of the United States National Museum, vol. 12, p. 194. [Halobates wuellerstorfi var., (Chatham Island).]
VAN DUZEE, E. P.
1933. The Templeton Crocker Expedition of the California Academy of Sciences, 1932. no. 4. Characters of twenty-four new species of Hemiptera from the Galapagos Islands and the coast and islands of Central America and Mexico. Proceedings of the California Academy of Sciences, ser. 4, vol. 21, pp. 25–40.
1933. The Templeton Crocker Expedition of the California Academy of Sciences, 1932. no. 12. The diurnal Lepidoptera of the Expedition. Proceedings of the California Academy of Sciences, ser. 4, vol. 21, pp. 139-146. [Six butterflies and skippers recorded. No new species.]
1937. No. 33. Hemiptera of the Templeton Crocker Expedition to Polynesia in 1934–1935. Proceedings of theCalifornia Academy of Sciences, ser. 4, vol. 22, pp. 111–126. [Records of 13 previously described species and Europiella mella;

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Euthiscia crockeri; Agallia mera; Platymetopius retusus; and Jassus infestus; all new species and all from Santa Cruz Island.]
VAN DUZEE, M. C.
1933. The Templeton Crocker Expedition of the California Academy of Sciences, 1932. no. 7. Dolichopodidae and Phoridae. Proceedings of the California Academy of Sciences, ser. 4, vol. 21, pp. 65–74, 1 pl. [Key to species of Chrysotus; new species: Condylostylus dentaticauda (James Island); Chrysotus brevicornis (Indefatigable Island); Chrysotus breveispina (Indefatigable, Albemarle island), Chrysotus latifacies (Indefatigable Island); Dohrniphora willowsi (South Seymour Island); Dokrniphora crockeri (South Seymour Island).]
VAN DYKE, E. C.
1953. The Coleoptera of the Galápagos Island Occasional Papers of the California Academy of Sciences, no. 22, pp. 1–181. [Comprehensive treatment of known species; many described as new.]
VON HAGEN, V. W.
1940. Ecuador the Unknown. Oxford University Press, New York. 296 pp.
1949. Ecudor and the Galapagos Islands. University of Oklahoma Press, 290 pp.
WALKER, F.
1848–1849 List of the specimens of Dipterous Insects in the collection of the British Museum. London. Parts I–IV, 1172 pp. [Describes several species from Galápagos Islands.]
1851. List of the specimens of Homopterous insects in the collection of the British Museum. London. vol. 2, pp. 261–636; vol. 3, pp. 637–907.
1870. Catalogue of the specimens of Dermaptera Satatoria in the collection of the British Museum, vol. 4, pp. 605–809. [Acridium literosum, n. sp. (p. 620) (Galápagos Islangs).]
1873. Catalogue of the specimens of Hemiptera Heteroptera in the collection of the British Museum. London. Part IV, pp. 1–210. [Capsus spoliatus (Charles, James islands); Capsus nigritulus (Charles Island); Capsus quadrinotatus (James Island), var. (Charles Island); all collected by C. Darwin.]
WALLENGREN, H. D. J.
1861. Lepidoptera species novas descripsit. In: Kongliga Svenska Fregatten Eugenies Resa Omkring Jorden… Zoologi 1, Insecta. pp. 351–390, 2 pls. [Euchelia gallopagensis, n. sp.]
WARD, R. A.
1953. Brief Notes on the Mallophaga I. Entomological News, vol. 64, pp. 201–204. [Synonymy of Docophourus peristictus Kellogg and Kuwana under Saemundssonia platygaster (Denny).]
WARREN, W.
1904. New Thyrididae and Geometridae from the Oriental Region. Novitates Zoologicae, vol. 11, pp. 483–492. [Perixera impudens, n. sp., from Gardner, Galápagos Islands (p. 487).]
1905. New American Thyrididae, Uraniidae, and Geometridae [Lepidoptera]. Novitates Zoologicae, vol. 12, pp. 307–379. [Sericosema lignala (p. 362) (S.E. Albemarle Island).]
WATERHOUSE, C.
1877. Coleoptera. In: Gunther, Account of the zoological colletion made during the visit of the H.M.S. "Peterel" to the Galapagos Islands. VII. Proceedings of the Zoological Society of London, 1877, pp. 77–82. [Includes new records, new species, and a list of known Coleoptera from the various islands.]

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WATERHOUSE, G. R.
1845. Descriptions of coleopterous insects collected by Charles Darwin, Esq., in the Galapagos Islands. Annals of Natural History, vol. 16, pp. 19–41. [20 new species.]
WHEELER, M. R.
1961. Some Drosophilidae (Diptera) from the Galapagos Islands. Annals of the Entomological Society of America, vol. 54, pp. 750. [5 species recorded; new records: Drosophila simulans (Chatham, Baltra islands); Drosophila nebulosa (Chatham Island); Drosophila sp. (Santa Cruz Island); Gitona brasiliensis (Santa Cruz Island).]
WHEELER, W. M.
1919. The ants of the Galapagos Islands. Proceedings of the California Academy of Sciences, ser. 4, vol. 2, pp. 259–310.
1924. The Formicidae of the Harrison Williams Galapagos Expedition. Zoologica, vol. 5, pp. 101–122, figs. 19–27, 1 tab. [Records of 18 forms, 2 species and 1 variety described as new; Monomorium floricola, a tropicopolitan species, reported for the first time; also, a termite, Calotermes pacificus, recorded from South Seymour Island.]
1933. The Templeton Crocker Expedition of the California Academy of Sciences, 1932. no. 6. Formicidae of the Templeton Crocker Excedition. Proceedings of the California Academy of Sciences, ser. 4, vol. 21, pp. 57–64. [Notes and new species as follows: Crematogaster brevispinosa chathamensis (Chatham Island); Camponotus planus var. hephaestus (Albemarle Island).]
WILLIAMS, F. X.
1907. [Report of the expedition of the California Academy of Sciences to the Galapagos Islands.] Entomological News, vol. 18, pp. 260–261. [Insects taken include at least 5 species each of butterflies and sphinx moths, numerous noctuids, Microlepidoptera, and Orthoptera, few Diptera and Hemiptera, more than 150 species of Coleoptera.]
1911. Expedition of the California Academy of Sciences to the Galápagos Islands, 1905–1906. III. The butterflies and hawk-moths of the Galápagos Islands. Proceedings of the California Academy of Sciences, ser. 4, vol. 1, pp. 289–322, 2 pls., 1 col. [General introduction, field notes, and new species. Eudamis galapagensis, n. sp. (Chatham and Albemarle islands).]
1926. Expedition of the California Academy of Sciences to the Galápagos Islands, 1905–1906. The bees and aculeate wasps of the Galápagos Islands. Proceedings of the California Academy of Sciences, ser. 4, vol. 2, pp. 347–357. [Several records and the following new species: Photopsis galapagensis (South Seymour Island); Odynerus (Padrodynerus) galapagensis (Chatham Island); new species.]
WILLIAMS, R. C.
1926. Studies in the Neotropical Hesperoidea. Paper I. Transactions of the American Entomological Society, vol. 52, pp. 61–87, 3 pls., 28 figs. [Eudamus galapagensis Williams (p. 64, pl. 2, fig. 4).]
WILLISTON, S. W.
1889. Diptera. In: Howard, L. O., Scientific results of explorations by the U. S. Fish Commission Steamer Albatross. no. V.—Annotated catalogue of the insects collected in 1877–'88. Proceedings of the United States National Museum, vol. 12, pp. 197. [Culex.]
WIRTH, W. W.
1951. A revision of the dipterous family Canaceidae. Occasional Papers of the Bernice

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P. Bishop Museum, vol. 20, pp. 245–275, 4 figs. [Canace snodgrassii Coquillet (= macateei Malloch) (p. 260) (Narborough and Albemarle islands); Canace maritima, n. sp. (p. 263) (Bartholomew Island).]
1956. New species and records of biting midges ectoparasitic on insects (Diptera, Heleidae). Annals of the Entomological Society of America, vol. 49, pp. 356–364. [Synonym of Ceratopogon galapagensis (p. 357).]
WITLACZIL, E.
1896. Die Ausbeute des "Pisani" an Halobates während der Erdumseglung 1882–1885. Wiener Entomologische Zeitung, vol. 5, pp. 177–182, pp. 231–234. [Halobates splendens—Pacific Ocean near the Galápagos Islands; Halobates wüllerstorffi, somewhat east of Galápagos Islands, a feather with Halobates eggs, off southwest coast of the Galápagos Islands.]
WOLF, T.
1892. Geografía y Geología del Ecuador. Leipzig, F. A. Brockhaus. pp. xii + 671, 12 pls., 47 figs., 2 maps. [Galápagos, part V, pp. 469–493; Insects, very brief, p. 485.]
1895. Die Galapagos-Inseln. Verhandlungen der Gesellschaft für Erdkunde zu Berlin, vol. 22, pp. 246–265.

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