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F1574a
Pamphlet:
de Beer, Gavin ed. 1960. Darwin's notebooks on transmutation of species. Part I. First notebook [B] (July 1837-February 1838). Bulletin of the British Museum (Natural History). Historical Series 2 (2) (January): 23-73.
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work . The clock had been provided with a mainspring. There is another point on which the First Notebook throws light, for it contains a splendid discussion of the principle of branching and sub-branching of the evolutionary tree (pp. 21, 22), and this shows that Darwin had already grasped fully the principle of divergence. This emerges also from his query whether Geoffroy-Saint-Hilaire visualized evolution as having taken place in straight or in branching lines (p. 113). The question naturally
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F1574d
Pamphlet:
de Beer, Gavin ed. 1960. de Beer, G. ed. 1960. Darwin's notebooks on transmutation of species. Part IV, Fourth notebook [E] (October 1838-10 July 1839). Bulletin of the British Museum (Natural History). Historical Series 2 (5) (September): 151-183.
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meet, they act precisely like two species of animals, they fight, eat each other, bring diseases to each other c, but then comes the most deadly struggle, namely which have the best fitted organization, or instincts (i.e. intellect in man) to gain the day (IV 63). It is not difficult in this passage to recognize experiences which Darwin underwent during the voyage of the Beagle. With regard to the evolution of man and the question whether his ancestors were bimanous or quadruped, Darwin had
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F3484
Book contribution:
Darwin, C. R. 1909. [Letter to F. W. Hope, 1837, 19 letters to R. Trimen, 1863-71]. In E. B. Poulton ed. Charles Darwin and the Origin of species: addresses, etc., in America and England in the year of the two anniversaries. London: Longmans, Green, and Co.
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far the species of the same genus differ in the ocelli. As I know from your Orchid Drawings how skilful an artist you are, perhaps it would not give you much more trouble to sketch any variable ocelli than to describe them.—I am very much obliged to you for so kindly assisting 1 For a further account of this and other uses of these markings, together with references to the Original memoirs, see' eye-spots' in index of Essays on Evolution (1908), 424. [page] 23
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F3484
Book contribution:
Darwin, C. R. 1909. [Letter to F. W. Hope, 1837, 19 letters to R. Trimen, 1863-71]. In E. B. Poulton ed. Charles Darwin and the Origin of species: addresses, etc., in America and England in the year of the two anniversaries. London: Longmans, Green, and Co.
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females.3 My friend Mr. Harry Eltringham has recently pointed out to me a passage, marked by much confusion of thought, in Hewitson's Exotic Butterflies, which might be read as an anticipation 1 See' dardanus' in index of Essays on Evolution (1908), 414; also Plate XXIII in Trans. Ent. Soc. Lond. (1908 J, 427-45. 2 Trans. Ent. Soc. Lond. (1874), 137. 3 E. M. M. (Oct., 1874), 113. 4 London, 1862-66, III: text of plate' Nymphalidæ. Diadema iii.: (pages unnumbered). [page] 23
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F1574a
Pamphlet:
de Beer, Gavin ed. 1960. Darwin's notebooks on transmutation of species. Part I. First notebook [B] (July 1837-February 1838). Bulletin of the British Museum (Natural History). Historical Series 2 (2) (January): 23-73.
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simplification. cf. also Jean Rostand. L' tat pr sent du transformisme, Paris 1931, p. 13. 2 Charles Lyell. Principles of Geology, London 1832. vol. 2, p. II which contains the first use in English of the term evolution in its present accepted sense. 3 Eiseley. Charles Darwin, Edward Blyth, and the theory of natural selection. Proc. Amer. Phil. Soc. vol. 103, 1959, p. 108. 4 Sir Ronald Fisher. Retrospect of the criticisms of the theory of natural selection , Evolution as a Process edited by Julian
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F1575
Periodical contribution:
Barrett, P. H. ed. 1960. A transcription of Darwin's first notebook [B] on 'Transmutation of species'. Bulletin of the Museum of Comparative Zoology, Harvard 122: [245]-296, for 1959-1960 (April).
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Pacific existed, might have been monsoons, when they ceased, importation ceased, and changes commenced; or intermediate land existed, or they may represent some large country long separated. On this idea of propagation [i.e., evolution] of species we can see why a form peculiar to continents, all bred in from one [page] 251 BARRETT: DARWIN'S FIRST EVOLUTIONARY NOTEBOO
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F1574a
Pamphlet:
de Beer, Gavin ed. 1960. Darwin's notebooks on transmutation of species. Part I. First notebook [B] (July 1837-February 1838). Bulletin of the British Museum (Natural History). Historical Series 2 (2) (January): 23-73.
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will be profitable to consider the legacies of Darwin's immediate predecessors and the contributions of his contemporaries that were known to him. First comes Erasmus Darwin.4 Erasmus Darwin believed in the transmutation of species and evolution because of the observed changes undergone by organisms during their life-history, the changes brought about by domestication and resulting from hybridization, and monstrous 1 William Sharp MacLeay. Horae entomologicae, London 1819-21. The Quinarian System
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F1574a
Pamphlet:
de Beer, Gavin ed. 1960. Darwin's notebooks on transmutation of species. Part I. First notebook [B] (July 1837-February 1838). Bulletin of the British Museum (Natural History). Historical Series 2 (2) (January): 23-73.
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fundamental difference between the Darwinian view of fortuitous variation (which has been experimentally demonstrated by Sir Ronald Fisher as correct), and all other attempts to explain evolution as due to adaptively directed mutation. It is at the base of the argument about design. If variation were designed, as Darwin wrote1 to Asa Gray, 26th November 1860, you would have to believe that the tail of the Fantail was led to vary in the number and direction of its feathers in order to gratify
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F1574a
Pamphlet:
de Beer, Gavin ed. 1960. Darwin's notebooks on transmutation of species. Part I. First notebook [B] (July 1837-February 1838). Bulletin of the British Museum (Natural History). Historical Series 2 (2) (January): 23-73.
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to Lyell about the succession of forms, Darwin forgot that he had himself published a paper on this subject in 1837. In 1860 when writing4 to Baden Powell he excused himself for not having given a list of his predecessors who rejected special creation, saying that he had attempted no history of the subject; yet later on the same day he remembered that he had a year or two previously drafted a Historical Sketch for his large work on evolution, in which Powell's name was mentioned with honour. All
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F1574a
Pamphlet:
de Beer, Gavin ed. 1960. Darwin's notebooks on transmutation of species. Part I. First notebook [B] (July 1837-February 1838). Bulletin of the British Museum (Natural History). Historical Series 2 (2) (January): 23-73.
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well worth while to study profoundly the origin and history of every terrestrial mammalia, especially moderately large ones. 1 Jean Baptiste de Lamarck. See Introduction. 2 Darwin means that arguments against the formation of species are absurd. The argument about the evolution of the otter through intermediate forms is developed in the Essay of 1844, p. 152. 3 Leonard Jenyns, afterwards Blomefield. Probably personal communication. 4 Thomas Bell. A History of British Quadrupeds, London 1837; pp
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F1574b
Pamphlet:
de Beer, Gavin ed. 1960. Darwin's notebooks on transmutation of species. Part II. Second notebook [C] (February to July 1838). Bulletin of the British Museum (Natural History). Historical Series 2 (3) (May): 75-118.
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varieties of many ages standing will not readily breed together (II 30). This may be compared with T. H. Huxley's caveat that Darwin had not proved his point until he could show that natural selection had led to sterility between divergent products of evolution. Darwin realized this diffi- 1cf. Developments in the study of animal communications by P. Marler, in Darwin's Biological Works, edited by P. R. Bell, Cambridge 1959, pp. 150-206. [page] 7
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F1574b
Pamphlet:
de Beer, Gavin ed. 1960. Darwin's notebooks on transmutation of species. Part II. Second notebook [C] (February to July 1838). Bulletin of the British Museum (Natural History). Historical Series 2 (3) (May): 75-118.
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realized that if species had not been separated and specially created, there must have been a mechanism of evolution. Before he hit upon the principle of natural selection his views on this problem could not be very precise, but the notion of competition appears in the following passage, albeit without defined penalties for the losers: Once grant that species and genus may pass into each other, grant that one instinct to be acquired (if the medullary point in ovum has such organization as to
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F1574c
Pamphlet:
de Beer, Gavin ed. 1960. Darwin's notebooks on transmutation of species. Part III. Third notebook [D] (July 15 to October 2nd 1838). Bulletin of the British Museum (Natural History). Historical Series 2 (4) (July):119-150.
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these principles and used this argument for the express purpose of discrediting the theory of evolution by natural selection. Other critics, more recent, have accepted Darwin's conclusions but claimed that he was mistaken in thinking that he had arrived at them inductively and even reproached him for speculating in his Notebooks, as if it was reprehensible. Darwin himself always maintained that he had used induction and wrote1 in his Autobiography that he worked on the true Baconian principles
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F1574c
Pamphlet:
de Beer, Gavin ed. 1960. Darwin's notebooks on transmutation of species. Part III. Third notebook [D] (July 15 to October 2nd 1838). Bulletin of the British Museum (Natural History). Historical Series 2 (4) (July):119-150.
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reasons which led Darwin to abandon the view of the immutability of species and to accept the hypothesis of transmutation or evolution. It is noteworthy that the three facts which started Darwin on his train of thought do not figure in this exposition, and this is a measure of the amount of consideration which Darwin had given to the problem between the time when he was in the Beagle and July 1837 when he opened his First Notebook on Transmutation of Species. As the problem was one of differences
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F1574c
Pamphlet:
de Beer, Gavin ed. 1960. Darwin's notebooks on transmutation of species. Part III. Third notebook [D] (July 15 to October 2nd 1838). Bulletin of the British Museum (Natural History). Historical Series 2 (4) (July):119-150.
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he was in the possession of the key of natural selection, Darwin worked by a combination of induction and deduction to construct the full theory of evolution by natural selection, as Sir Julian Huxley2 and I3 have shown. But this belongs to a stage in the development of Darwin's thought later than is represented by the first three Notebooks on Transmutation of Species, and is found in the Fourth Notebook, the Sketch of 1842, and the Essay of 1844. The remainder of the Third Notebook is largely
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F1574c
Pamphlet:
de Beer, Gavin ed. 1960. Darwin's notebooks on transmutation of species. Part III. Third notebook [D] (July 15 to October 2nd 1838). Bulletin of the British Museum (Natural History). Historical Series 2 (4) (July):119-150.
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page 189: The close resemblance of the Bird to the Reptile in its skeleton is well exemplified in the young Ostrich, 4Isaac Disraeli. Curiosities of Literature. 5of. Darwin's Sketch of 1842 and his Essay of 1844; in Evolution by Natural Selection with a Foreword by Sir Gavin de Beer, Cambridge 1958, pp. 83 and 249. [page] 13
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F1574d
Pamphlet:
de Beer, Gavin ed. 1960. de Beer, G. ed. 1960. Darwin's notebooks on transmutation of species. Part IV, Fourth notebook [E] (October 1838-10 July 1839). Bulletin of the British Museum (Natural History). Historical Series 2 (5) (September): 151-183.
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those who solved the problem of evolution and natural selection are by no means complete, but it is already possible to see two curious patterns in the history of thought. The first relates to the manner in which arguments have simply been turned upside down as a result of progress of knowledge. The foremost example of this reversal of direction is the fate of Paley's arguments aimed at proving that the adaptations of plants and animals to their environment show evidence of purposive design.2 First
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F1574d
Pamphlet:
de Beer, Gavin ed. 1960. de Beer, G. ed. 1960. Darwin's notebooks on transmutation of species. Part IV, Fourth notebook [E] (October 1838-10 July 1839). Bulletin of the British Museum (Natural History). Historical Series 2 (5) (September): 151-183.
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ingredients which he required, both to establish the fact of evolution and to show that natural selection provided the explanation of how species become modified, was unknown to Lyell who missed the great chance, partly by failing to test the imaginary link between catastrophism and progressionism, and partly because his mind was orientated away from transmutation of species for reasons of theological orthodoxy. How close Lyell came to the facts without recognizing them may be seen in the second
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F1574d
Pamphlet:
de Beer, Gavin ed. 1960. de Beer, G. ed. 1960. Darwin's notebooks on transmutation of species. Part IV, Fourth notebook [E] (October 1838-10 July 1839). Bulletin of the British Museum (Natural History). Historical Series 2 (5) (September): 151-183.
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though he had available to him all the basic ingredients out of which Fisher constructed his synthesis, Bateson was blinded by the clean-cut results of such Mendelian crosses as were known to him, appearing to have arisen ready-made without selection, and this prejudiced him against Darwinian selection. He was eventually driven to the untenable view that evolution had been stopped down at the start, and had occurred through the successive removal of inhibitory factors.1 The lesson to be
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F1647
Periodical contribution:
Darwin, C. R. 1837. On certain areas of elevation and subsidence in the Pacific and Indian oceans, as deduced from the study of coral formations. [Read 31 May] Proceedings of the Geological Society of London 2: 552-554.
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than a hemisphere, divided into symmetrical areas, which within a limited period of time have undergone certain known movements, we obtain some insight into the system by which the crust of the globe is modified during the endless cycle of changes. 1 This is Darwin's first published reference to his interest in the origin of species. This public statement just three months after becoming an evolutionist, shows he was not afraid to talk about evolution and did not keep this part of his work secret
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