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A331
Book:
Poulton, Edward Bagnall. 1909. Charles Darwin and the Origin of species: addresses, etc., in America and England in the year of the two anniversaries. London: Longmans, Green, and Co.
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the group of species hitherto regarded as ant-like, and he adds, 'It is most interesting that Burchell should have noticed the resemblance to an ant in its movements. This suggests that the perfect imitation in shape, as well as in movement, seen in many species was started in forms of an appropriate size and colour by the mimicry of movement alone.' Up to the present time Burchell is the only naturalist who has observed an example which still exhibits this ancestral stage in the evolution of
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A331
Book:
Poulton, Edward Bagnall. 1909. Charles Darwin and the Origin of species: addresses, etc., in America and England in the year of the two anniversaries. London: Longmans, Green, and Co.
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, ii. 392. 4 See Poulton, Essays on Evolution, 1908, 65, 85-8. [page] 126 VALUE OF COLOU
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A331
Book:
Poulton, Edward Bagnall. 1909. Charles Darwin and the Origin of species: addresses, etc., in America and England in the year of the two anniversaries. London: Longmans, Green, and Co.
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interesting and important of subjects in relation to this theory as well as to evolution. In Mimicry we investigate the effect of environment in its simplest form: we trace the effects of the pattern of a single species upon that of another far removed from it in the scale of classification. When there is reason to believe that the model is an invader from another region and has only recently become an element in the environment 1 More Letters, i. 175. 2 Life and Letters, ii. 245. See also pp
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A331
Book:
Poulton, Edward Bagnall. 1909. Charles Darwin and the Origin of species: addresses, etc., in America and England in the year of the two anniversaries. London: Longmans, Green, and Co.
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one so simple that we might have expected it to act as a' unit character', so small a fraction of the pattern that we could hardly speak of its sudden disappearance as 'discontinuous' evolution—that even this behaves differently on the two surfaces of the wing, while the individuals from which it has disappeared are [page] 168 MIMICRY IN N. AMERICAN BUTTERFLIE
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A331
Book:
Poulton, Edward Bagnall. 1909. Charles Darwin and the Origin of species: addresses, etc., in America and England in the year of the two anniversaries. London: Longmans, Green, and Co.
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of wine.'2 CONCLUSIONS It will probably be convenient to sum up rather fully the chief conclusions contained in the foregoing address. 1. The study of Mimicry possesses special advantages for an understanding of the history and causes of evolution. 1 The letter is addressed: 'The Revd. F. W. Hope, 56, Upper Seymour Street. 'At the head Mr. Hope had written' D', and the date' 1837' . The red-stamped postmark gives the date' Ju. 22, 1837'. Darwin's own address (36, Great Marlborough Street) does
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A331
Book:
Poulton, Edward Bagnall. 1909. Charles Darwin and the Origin of species: addresses, etc., in America and England in the year of the two anniversaries. London: Longmans, Green, and Co.
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-brown apex of the fore wing in Limenitis lorquini would be demonstrated by a glance at its average condition in specimens from the different localities as we pass from north to south. Furthermore, we might reasonably hope that a similar series collected after an interval not greatly prolonged would exhibit differences in average composition—the actual measurable evidence of the evolution of a character in a species in the natural state. Even though such evidence be left for our successors to
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A331
Book:
Poulton, Edward Bagnall. 1909. Charles Darwin and the Origin of species: addresses, etc., in America and England in the year of the two anniversaries. London: Longmans, Green, and Co.
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far the species of the same genus differ in the ocelli. As I know from your Orchid Drawings how skilful an artist you are, perhaps it would not give you much more trouble to sketch any variable ocelli than to describe them.—I am very much obliged to you for so kindly assisting 1 For a further account of this and other uses of these markings, together with references to the Original memoirs, see' eye-spots' in index of Essays on Evolution (1908), 424. [page] 233 EYE-SPOTS ON BUTTERFLIES' WINGS: 186
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A331
Book:
Poulton, Edward Bagnall. 1909. Charles Darwin and the Origin of species: addresses, etc., in America and England in the year of the two anniversaries. London: Longmans, Green, and Co.
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females.3 My friend Mr. Harry Eltringham has recently pointed out to me a passage, marked by much confusion of thought, in Hewitson's Exotic Butterflies, which might be read as an anticipation 1 See' dardanus' in index of Essays on Evolution (1908), 414; also Plate XXIII in Trans. Ent. Soc. Lond. (1908 J, 427-45. 2 Trans. Ent. Soc. Lond. (1874!, 137. 3 E. M. M. (Oct., 1874), 113. 4 London, 1862-66, III: text of plate' Nymphalidæ. Diadema iii.: (pages unnumbered). [page] 238 DARWIN'S LETTERS TO R
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A331
Book:
Poulton, Edward Bagnall. 1909. Charles Darwin and the Origin of species: addresses, etc., in America and England in the year of the two anniversaries. London: Longmans, Green, and Co.
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possible to give the strongest cases opposed to me, and often such conjectures as occur to me.'2 1856, July 19.—'it is absolutely necessary that I should discuss single and double creations, as a very crucial point on the general Origin of species, and I must confess, with the aid of all sorts of visionary hypotheses, a very hostile one.'3 The above-quoted sentences sum up very briefly Darwin's conclusion that evolution as he conceived of it implied that each species had appeared once only in a
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A331
Book:
Poulton, Edward Bagnall. 1909. Charles Darwin and the Origin of species: addresses, etc., in America and England in the year of the two anniversaries. London: Longmans, Green, and Co.
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time refuting the opinion-not uncommon even at the present day-that a terrestrial species such as man may exist on Mars or on some other body outside the earth. For Darwin shows in the following letter that, in order to produce the same species twice over, the same material must have been subject to the same selection at every stage, right back to the unknown starting-point of organic evolution. 'As far as I can judge, the improbability is extreme that the same well-characterised species should
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A331
Book:
Poulton, Edward Bagnall. 1909. Charles Darwin and the Origin of species: addresses, etc., in America and England in the year of the two anniversaries. London: Longmans, Green, and Co.
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assistance. The extent of this unintentional, but very serious, misrepresentation of an authority by his exponent, can be most clearly shown by printing together passages by de Vries and Bateson from 1 'Of the inheritance of mutations there is no doubt. Of the transmission of fluctuations there is no very strong evidence. It is therefore reasonable to regard the mutation as the main, if not the only, basis of evolution.'(p. 72.) 2 Mendel's Principles of Heredity, Cambridge (1909), 287. 3 Species
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A331
Book:
Poulton, Edward Bagnall. 1909. Charles Darwin and the Origin of species: addresses, etc., in America and England in the year of the two anniversaries. London: Longmans, Green, and Co.
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he has termed fluctuating.'2 In order to ensure an accurate recognition it will be safest to quote de Vries's words. (1) In the celebrated Mutationstheorie (Leipzig, 1901, I.) de Vries states that, in advocating the use of the term' fluctuation', he is merely adopting a word often used by Darwin himself.3 Thus, 1 Variation, Heredity and Evolution, London, 1909, 2nd Ed., 155. See also passage (1) quoted from Mr. Lock on p. 270. 2 Mendelism, R. C. Punnett, 2nd Ed., Cambr. (1907), 70. 3 An
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A331
Book:
Poulton, Edward Bagnall. 1909. Charles Darwin and the Origin of species: addresses, etc., in America and England in the year of the two anniversaries. London: Longmans, Green, and Co.
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hypothesis of the direct influence of environment as a motive cause of evolution:— 'In regard to thorns and spines I suppose that stunted and (illegible) hardened processes were primarily left by the abortion of various appendages, but 1 must believe that their extreme sharpness and hardness is the result of fluctuating variability and the survival of the fittest . 'In a letter to G. H. Lewes, Aug. 7, 1868. More Letters, i. 308. 1 De Vries is here referring to p. 29, where he distinguishes the two
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A331
Book:
Poulton, Edward Bagnall. 1909. Charles Darwin and the Origin of species: addresses, etc., in America and England in the year of the two anniversaries. London: Longmans, Green, and Co.
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. * 'From his own experiments, de Vries has come to the conclusion that, when selection is really efficient, the full possible effect of this process is exhausted in quite a small 1 Fifty Years of Darwinism, New York (1909), 173-4. 2 Variation, Heredity and Evolution. London, 1909. Second Ed. [page] 271 OTHER WRITERS ON 'FLUCTUATIONS
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A331
Book:
Poulton, Edward Bagnall. 1909. Charles Darwin and the Origin of species: addresses, etc., in America and England in the year of the two anniversaries. London: Longmans, Green, and Co.
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'fluctuations' are assumed by him to be transmissible by heredity, and that this assumption is an essential element in the author's definition of his technical term. When we remember that they are just the 'Individual differences' of Darwin, and that de Vries's belief in their powerlessness for continued evolution is based on Francis Galton's well-known law of recession, it is really waste of time to inquire whether they are transmissible. But such positive statements to the contrary have been made by
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being, modified as time goes on. This is the fate of all scientific theories; none are stationary, none are final. The development of Science is a continuous process of evolution, like the world of phenomena itself. It has however some few landmarks which stand out exceptional and prominent. None of these is greater or will be more enduring in the history of thought than the theory associated with the name of Charles Darwin. I cannot attempt further to weigh or estimate the influence and the
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A36
Periodical contribution:
Darwin Centenary Number. Christ's College Magazine. vol. XXIII, Easter Term, 1909.
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Law that has regulated the Introduction of New Species (Ann. and Mag. Nat. Hist., 1855). The Law is Every species has come into existence coincident both in space and time with a pre-existing closely allied species. Mr Wallace remarks (My Life, Vol. 1. p. 335) This clearly pointed to some kind of evolution but the how was still a secret. Mr Huxley has said of this powerful essay : On reading it afresh I have been astonished to recollect how small was the impression it made. (Life and Letters of
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A36
Periodical contribution:
Darwin Centenary Number. Christ's College Magazine. vol. XXIII, Easter Term, 1909.
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migratory bird might carry them great distances. 2 In the Origin however he made it plain that Man was included in his scheme of evolution. [page] 22
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A36
Periodical contribution:
Darwin Centenary Number. Christ's College Magazine. vol. XXIII, Easter Term, 1909.
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of the means by which evolution has taken place in the organic world, the theory of Natural Selection which he published contemporaneously with Mr A. R. Wallace in 1858, and set forth with a wealth of illustration and argument in The Origin of Species in the following year. Now although at the present time we are all evolutionists we hear it frequently said that Darwinism has had its day, and that Natural Selection is insufficient to account for the facts. It is true that very many of the
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A36
Periodical contribution:
Darwin Centenary Number. Christ's College Magazine. vol. XXIII, Easter Term, 1909.
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England during the last half-century. Here we see evolution taking place in Nature, but as to the usefulness of the dark forms we can only guess. But it is possible that while the change of colour is useless in itself, it is associated with increased constitutional vigour or some other advantage of the kind, and that it is the associated improvement which has caused the varieties to increase, not the colour-change itself. This leads us on to the third, and perhaps the most frequent objection in
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