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A162
Book:
Seward, A. C. ed. 1909. Darwin and modern science. Essays in commemoration of the centenary of the birth of Charles Darwin and of the fiftieth anniversary of the publication of The origin of species. Cambridge: Cambridge University Press.
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subjects which their teachers were investigating. During this period nevertheless one distinct advance was made, that with which Weismann's name is prominently connected. In Darwin's genetic scheme the hereditary transmission of parental experience and its consequences played a considerable role. Exactly how great that role was supposed to be, he with his habitual caution refrained from specifying, for the sufficient reason that he did not know. Nevertheless much of the process of Evolution
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A162
Book:
Seward, A. C. ed. 1909. Darwin and modern science. Essays in commemoration of the centenary of the birth of Charles Darwin and of the fiftieth anniversary of the publication of The origin of species. Cambridge: Cambridge University Press.
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the temperature of our atmosphere decides that no liquid carbon shall be found on the face of the earth: but we do not suppose that the form of the diamond has been gradually achieved by a process of Selection. So again, as the course of descent branches in the successive generations, Selection determines along which branch Evolution shall proceed, but it does not decide what novelties that branch shall bring forth. La Nature contient le fonds de toutes ces vari t s, mais le hazard ou l'art les
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A162
Book:
Seward, A. C. ed. 1909. Darwin and modern science. Essays in commemoration of the centenary of the birth of Charles Darwin and of the fiftieth anniversary of the publication of The origin of species. Cambridge: Cambridge University Press.
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plane back, and so looked rather less prominent. The abundance of Adaptation, we all grant, is an immense, almost an unsurpassable difficulty in all non-Lamarckian views of Evolution; but if the steps by which that adaptation arose were fortuitous, to imagine them insensible is assuredly no help. In one most important respect indeed, as has often been observed, it is a multiplication of troubles. For the smaller the steps, the less could Natural Selection act upon them. Definite variations—and of
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A162
Book:
Seward, A. C. ed. 1909. Darwin and modern science. Essays in commemoration of the centenary of the birth of Charles Darwin and of the fiftieth anniversary of the publication of The origin of species. Cambridge: Cambridge University Press.
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chapter may be very briefly summarised. In it Darwin shows that the intellectual and moral faculties are perfected through natural selection. He inquires how it can come about that a tribe at a low level of evolution attains to a higher, although the best and bravest among them often pay for their fidelity and courage with their lives without leaving any descendants. In this case it is the sentiment of glory, praise and blame, the admiration of others, which bring about the increase of the better
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A162
Book:
Seward, A. C. ed. 1909. Darwin and modern science. Essays in commemoration of the centenary of the birth of Charles Darwin and of the fiftieth anniversary of the publication of The origin of species. Cambridge: Cambridge University Press.
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discovery, which he illustrated by a humorous sketch1. But there are still many followers in Darwin's footsteps in England. Keane has worked at the special genealogical tree of the Primates; Keith has inquired which of the anthropoid apes has the greatest number of characters in common with man; Morris concerns himself with the evolution of man in general, especially with his acquisition of the erect position. The recent discoveries of Pithecanthropus and Homo primigenius are being vigorously discussed
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A162
Book:
Seward, A. C. ed. 1909. Darwin and modern science. Essays in commemoration of the centenary of the birth of Charles Darwin and of the fiftieth anniversary of the publication of The origin of species. Cambridge: Cambridge University Press.
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indicate a common descent, and cannot be explained as phenomena of convergence. I believe I have shown in the above sketch that a theory which derives man directly from lower forms without regarding apes as transition-types leads ad absurdum. The close structural relationship between man and monkeys can only be understood if both are brought into the same line of evolution. To trace man's line of descent directly back to the old Eocene mammals, alongside of, but with no relation to these very
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A162
Book:
Seward, A. C. ed. 1909. Darwin and modern science. Essays in commemoration of the centenary of the birth of Charles Darwin and of the fiftieth anniversary of the publication of The origin of species. Cambridge: Cambridge University Press.
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cenogenesis. As early as 1874 I had emphasised, in the first chapter of my Evolution of Man, the importance of discriminating carefully between these two sets of phenomena: In the evolutionary appreciation of the facts of embryology we must take particular care to distinguish sharply and clearly between the primary, palingenetic evolutionary processes and the secondary, cenogenetic processes. The palingenetic phenomena, or embryonic recapitulations, are due to heredity, to the transmission of
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A162
Book:
Seward, A. C. ed. 1909. Darwin and modern science. Essays in commemoration of the centenary of the birth of Charles Darwin and of the fiftieth anniversary of the publication of The origin of species. Cambridge: Cambridge University Press.
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existed long before Darwinian evolution was thought of and will endure without any reference to philosophic speculations. She is a mistress in whose face are beauties and in whose arms are delights elsewhere unattainable. She is and always has been pursued for her own sake without any reference to philosophy, science, or utility. Darwin's own views of the bearing of the facts of embryology upon questions of wide scientific interest are perfectly clear. He writes1: On the other hand it is highly
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A162
Book:
Seward, A. C. ed. 1909. Darwin and modern science. Essays in commemoration of the centenary of the birth of Charles Darwin and of the fiftieth anniversary of the publication of The origin of species. Cambridge: Cambridge University Press.
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they ever discharged a branchial function. Until such evidence is forthcoming, it is beside the point to say that it cannot be disputed that they are evidence of a piscine ancestry. It must, therefore, be admitted that one outcome of the progress of embryological and palaeontological research for the last 50 years 1 The Evolution Theory, by A. Weismann, English Translation, Vol. II. p. 176, London, 1904. [page] 17
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A162
Book:
Seward, A. C. ed. 1909. Darwin and modern science. Essays in commemoration of the centenary of the birth of Charles Darwin and of the fiftieth anniversary of the publication of The origin of species. Cambridge: Cambridge University Press.
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Fayoum show us that the region sought for is Africa, and that the elephants form just such a series of gradual modifications as we have found among other hoofed animals. The 1 C. W. Andrews, On the Evolution of the Proboscidea, Phil. Trans. Roy. Soc. London, Vol. 196, 1904, p. 99. [page] 19
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A162
Book:
Seward, A. C. ed. 1909. Darwin and modern science. Essays in commemoration of the centenary of the birth of Charles Darwin and of the fiftieth anniversary of the publication of The origin of species. Cambridge: Cambridge University Press.
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lines of inquiry has established the theory of evolution upon a foundation of ever growing solidity. It is Darwin's imperishable glory that he prescribed the lines along which all the biological sciences were to advance to conquests not dreamed of when he wrote. [page 200
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A162
Book:
Seward, A. C. ed. 1909. Darwin and modern science. Essays in commemoration of the centenary of the birth of Charles Darwin and of the fiftieth anniversary of the publication of The origin of species. Cambridge: Cambridge University Press.
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governed by Natural Selection, just as much as the specialisation of the rostellum in an Orchid, or of the pappus in a Composite. Did space allow, other examples might be added. We may venture to maintain that the glimpses which the fossil record allows us into early stages in the evolution of organs now of high systematic importance, by no means justify the belief in any essential distinction between morphological and adaptive characters. Another point, closely connected with Darwin's theory
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A162
Book:
Seward, A. C. ed. 1909. Darwin and modern science. Essays in commemoration of the centenary of the birth of Charles Darwin and of the fiftieth anniversary of the publication of The origin of species. Cambridge: Cambridge University Press.
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accordance with the environment. In the examples mentioned the modified character in the simple varieties (or a number of characters in elementary species) appears more or less suddenly and is constant in the above sense. The result is what de Vries has 1 Bateson, Reports to the Evolution Committee of the Royal Society, London, 1902; cf. also Lotsy, Vorlesungen, Vol. I. p. 234. D. 16 [page] 24
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A162
Book:
Seward, A. C. ed. 1909. Darwin and modern science. Essays in commemoration of the centenary of the birth of Charles Darwin and of the fiftieth anniversary of the publication of The origin of species. Cambridge: Cambridge University Press.
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produced. But perhaps the view here put forward in regard to the production of form throws new light on this puzzling problem. 1 Mutationstheorie, Vol. I. pp. 412 et seq. 2 Korschinsky, Heterogenesis und Evolution, Flora, 1901. 3 L. de Vilmorin, Notices sur l'am lioration des plantes, Paris, 1886, p. 36. 4 Klebs, K nstliche Metamorphosen, Stuttgart, 1906, p. 152. [page] 24
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A162
Book:
Seward, A. C. ed. 1909. Darwin and modern science. Essays in commemoration of the centenary of the birth of Charles Darwin and of the fiftieth anniversary of the publication of The origin of species. Cambridge: Cambridge University Press.
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successive generations of the same species in these regions than in the temperate or tropical regions. The writer is inclined to believe that these results have some bearing upon a problem which plays an important role in theories of evolution, namely, the cause of natural death. It has been stated that the processes of differentiation and development lead also to the natural death of the individual. If we express this in chemical terms it means that the chemical processes which underlie development
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A162
Book:
Seward, A. C. ed. 1909. Darwin and modern science. Essays in commemoration of the centenary of the birth of Charles Darwin and of the fiftieth anniversary of the publication of The origin of species. Cambridge: Cambridge University Press.
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therefore of interest to compare, in a brief introductory section, the older with the newer teleological views. The distinctive feature of Natural Selection as contrasted with other attempts to explain the process of Evolution is the part played by the struggle for existence. All naturalists in all ages must have known something of the operations of Nature red in tooth and claw ; but it was left for this great theory to suggest that vast extermination is a necessary condition of progress, and even
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A162
Book:
Seward, A. C. ed. 1909. Darwin and modern science. Essays in commemoration of the centenary of the birth of Charles Darwin and of the fiftieth anniversary of the publication of The origin of species. Cambridge: Cambridge University Press.
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older naturalists who thought and spoke with Burchell of the intention of Nature and the adaptation of beings to each other, and to the situations in which they are found, could have conceived the possibility of evolution, they must have been led, as Darwin was, by the same considerations to Natural Selection. This was impossible for them, because the philosophy which they followed contemplated the phenomena of adaptation as part of a static immutable system. Darwin, convinced that the system
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A162
Book:
Seward, A. C. ed. 1909. Darwin and modern science. Essays in commemoration of the centenary of the birth of Charles Darwin and of the fiftieth anniversary of the publication of The origin of species. Cambridge: Cambridge University Press.
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movements. This suggests that the perfect imitation in shape, as well as in movement, seen in many species was started in forms of an appropriate size and colour by the mimicry of movement alone. Up to the present time Burchell is the only naturalist who has observed an example which still exhibits this ancestral stage in the evolution of mimetic likeness. Following the teachings of his day, Burchell was driven to believe that it was part of the fixed and inexorable scheme of things that these
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A162
Book:
Seward, A. C. ed. 1909. Darwin and modern science. Essays in commemoration of the centenary of the birth of Charles Darwin and of the fiftieth anniversary of the publication of The origin of species. Cambridge: Cambridge University Press.
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Bates in Life and Letters, II. p. 392. 4 See Poulton, Essays on Evolution, 1908, pp. 65, 85-88. 5 New Ser. Vol. III. 1863, p. 219. 6 Ed. 1872, pp. 375-378. 7 Ed. 1874, pp. 323-325. 8 More Letters, I. p. 176. [page] 28
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A162
Book:
Seward, A. C. ed. 1909. Darwin and modern science. Essays in commemoration of the centenary of the birth of Charles Darwin and of the fiftieth anniversary of the publication of The origin of species. Cambridge: Cambridge University Press.
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example. In Hypolimnas itself the females mimic Danainae with patterns very different from those preserved by the non-mimetic males: in the sub-genus Euralia, both sexes resemble the black and white Ethiopian Danaines with patterns not very dissimilar from that which we infer to have existed in the non-mimetic ancestor. (7) Although a melanic form or other large variation may be of the utmost importance in facilitating the start of a mimetic likeness, it is impossible to explain the evolution
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