RECORD: Darwin, C. R. 1856.05.15. In Dr Grays manual there are 689 genera with 2004 species. CUL-DAR15.2.21,21a-21b. Edited by John van Wyhe (Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Transcribed by Christine Chua and edited by John van Wyhe 3.2023. RN1

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volume CUL-DAR15.2 contains calculations and tables for Darwin's 'big book' Natural Selection F1583.

Gray, Asa. 1856. Manual of the botany of the northern United States: second edition; including Virginia, Kentucky, and all east of the Mississippi: arranged according to the natural system. New York: George P. Putnam.


[21]

In Dr Grays Manual there are 689 genera with 2004 species subtracting the 369 genera with single species, we have 689 [-] 369 [=] 320

2004 2004

2004 [-] 369 [=] 1635 [÷] 320 [=] 5.11

So that the genera excluding the genera with single species have 5.11 species per genus. —

But Dr gray has not marked in his M.S list Salix: with 22 sp & as Carex has so very many viz 126 (with many of these close) these two may be left out. Then we shall have 318/1638 [-] 148 [=] 1487 1 /4.67

(If we further leave out aster with 35 sp. & solidago with 31 species

(no good reason to do this) [calculations not transcribed]

Now Asa Gray has marked for me 115 genera with 733 close species (N. B he has marked more than half the species (with Carex Salix & species of single genera omitted)); therefore, but the species of Salix & carex here omitted, now these 115 genera have on average (115/733 (6.37) whe but the other genera, ( see above) with which this number is comparable, have on average 4.67; hence the genera with "close species" have 1.7 on average more species. — How funnily the same as H. Watson viz 1.68 see Back

(If aster with 35 sp & solidago with 31 sp be omitted then the genera with close species have only 5.90 species on average, & 4.49 is standard of comparison, but I see no good reason to omit these genera.)

In the Manual, subtracting the genera salix & carex, & the 369 genera with single species, & 121 genera with 2 species, there is left 197 genera having more than 2 species in each. These species amount 1245. Therefore each genus has on average 6.32 species; But the 32 genera marked in Dr Grays list, as having 3 or more species "close" have on average 11.28 species: so that same result brought out very strong. (even if aster & solidago be omitted the average is 9.50 so much larger than 6.32.)

[21v]

In Dr Gray Manual these 532 genera with 1431 species now subtracting the 369 genera with single species, we have

532 [-] 369 [=] 163

1431 [-] 369 [=] 1062 [÷] 163

[21]

Genera with 3 or more "close species", abstracted from Dr. Grays M.S. List

no' species in Genus

Ranunculus 16

Viola 17

Silene 12

Cerastium 5

Polygala 13

Œnothera 7

Viburnum 9

Hedyotis 5

Liatris 5

Eupatorium 13

Aster 35

Solidago 31

Helianthus 16

Artemisia 6

Nabalus 9

Dipleuranthus 3

Gerardia 8

17 210

 

no' species in Genus

Onosmodium 3

Cuscuta 7

Polygonum 20

Callitriche 4

Quercus 18

Abies 5

Corallorhiza 4

Platanthera 16

Juncus 20

Cyperus 15

Suipus 14

Agrostis 6

Poa 10

Festuca 6

Triticum 3

15 151 [calculations not transcribed]

17

32

So that these genera have on average 11.28 species per genus.

[21a]

Another & fairer way of working is this. The genera which have not close species are (318 - 165 =) 203 & these include (1487 - 733=) 754 species, so that 203/754 (3.71

The genera with close species having 6.37

May 15/56


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Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

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