RECORD: Darwin, C. R. & Emma Darwin. 1864.03.01-31. Clematis viticella venosa. CUL-DAR157.1.70. Edited by John van Wyhe (Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Transcribed by Christine Chua and edited by John van Wyhe 4.2023. RN1

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here.

The volumes CUL-DAR157.1-2 contain notes, abstracts etc. for Darwin's long paper and later book Climbing plants (1865). It was also commercially available as a softbound offprint, F834, F834a. See R. B. Freeman's bibliographical introduction. Items CUL-DAR157.1.11-60 were in a folder marked "Twiners". Items CUL-DAR157.1.61-112 were in a folder marked "Leaf-climbers" and items CUL-DAR157.1.114-147 were in a folder marked "Tendrils". Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin.


[70]

March 1' 64 _Clematis viticella venosa

String put on terminal peduncle caused slight curvature in 24° & caused it to become rectangular in 4 days & + 18°.— (Confirmed roughly by previous trial)

In 7 days had turned back parallel to peduncle— In another cases had clasped string against first leaflet in 4 days—

String, instead of cotton as in C. flammula—

Certainly end of peduncle far more sensitive than any other part which were rarely acted on by loops of string — a basal leaf & its leaflets (ie 3' divisions) & main peduncle close by this leaflet) have all bend round stick —

March 14th a slight rub rub 4 or 5 times with twig caused in 1°. 15' perhaps some two terminal peduncles to curve & much of continued for several hours — Ternate peduncles all sensitive

[70v]

The leaf angularly bends downwards as in drawing, thus forms a hook; hence sensitiveness of end portion— action of wind & growth. Stick put to basal leaflet peduncle was not sensibly curved till 3° 30'. Stick taken away peduncle continued to moved during the night.

When once bent has a either from a touch or a stick has apparently no power to straighten itself. & always continues to move long after stimulus — Can certainly straighten again very slowly, in course of about 24°.—

I allowed under surface of terminal peduncle to catch pin it was perceptibly curved in 45' & in 70' was bent from right angle to parallel to leaf, so moved 90° in this time.

(3

A long inclined branch, upper shoot with close leaves 5 1/2 long & with lower internode 7 1/2 inch long, placed in front of window during 2 days, — second day cloudy & uniform, continually moved across the window from N. to S. (& so not caused by light) & made 3 traverses, (or possibly ellipses) in 8°. — (no doubt ellipses) i.e. on average each in 2°. 40'. The traverses were curved & of unequal length.— By this movement apex moved 4 5/8 of inch from side to side & top of lower internode, which bore leaves 1 1/8 of inch. — Thus younger leaves are moved & I have seen them thus catch stick; but yet very little power thus.— wind & growth far more potent — I saw similar movements in Greenhous Hard to understand cause unless a creeper. —

[3v]

Does not twine up a straight stick

March 31' it is true that young leaves are quite hooked.—

The String which so slowly caused the end peduncle to curve & hardly acted on lateral peduncle was 12 inches long & 50 inches weighed 8.2 gr. so that 12 inch weigh, 1.968 gr ie nearly 2 grains ―

Here peduncle hardly sensible— exclusively in C. glandulosa— & probably in C. Montana — not in each division of C. calycina— Most in hook at end of viticella — More specific action & travelling to end in sensitiveness

12 inch

12 in

9 1/2


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Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

File last updated 19 July, 2023