RECORD: Darwin, C. R. 1863.08.09-12. Tropaeolum Canariense. CUL-DAR157.1.89-90. Edited by John van Wyhe (Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Transcribed by Christine Chua and edited by John van Wyhe 4.2023. RN1

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here.

The volumes CUL-DAR157.1-2 contain notes, abstracts etc. for Darwin's long paper and later book Climbing plants (1865). It was also commercially available as a softbound offprint, F834, F834a. See R. B. Freeman's bibliographical introduction. Items CUL-DAR157.1.11-60 were in a folder marked "Twiners". Items CUL-DAR157.1.61-112 were in a folder marked "Leaf-climbers" and items CUL-DAR157.1.114-147 were in a folder marked "Tendrils". Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin.


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Aug. 9th 63/ Tropæolum Canariense — young plant 7 inches high—

2 lowest internodes do not move: above these 4 at unequal internodes (not counting a minute one on summit, moved reverse of watch.

The plant first swept a circle with shoot bent, & then one with shoot more upright, & then a li one with shoot still more bent, & so formed a spiral on glass, which interacted itself.

First circle (ie spire coming to meridian on which it started) was performed in 1°. 56' — a second smaller circle in 1°. 44'— a third circle in 1°. 45; this was largest sweep of all, ie shoot much bent during whole course; but it came back to exact point, where 3 circles previously it had started at 11°. 13' in morning. It performed the 3 circles in 5° 25'.— 1°. 48'

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Aug 10th I found after night that footstalk of young leaf had wound round stick in direction of reverse of watch — Took away stick & in 6°, peduncle which was greatly hooked, quite straightened itself. Accordingly I tied internode above & below a young leaf; it moved greatly toward light; but cd. detect no other movement. —

Aug. 11' In greenhouse I can see 2 leaves on same stem wound round stick in opposite direction — in one case peduncle completely hooked— In study a young leaf which was completely hooked yesterday has freed itself & is almost straight; 2 or 3 older leaves a have their peduncles bent & are marked with red to see if get straight.

Lettington says climb up stems, by leaves & not by twining.

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Does not climb Tropæolum Canariense (Peregrinum)

Aug 11th I have now ascertained that quite young leaves & even leaves 1 1/4 inch in diameter have whole peduncle sensitive & if touched on any side in from 1/4 to 1° curl up to that side, & thus clasp stick if stationary):

Leaf Leaf does not move.— As yet not seen leaf move move that to form a hook, ie. upper & basal ends parallel. — I doubt if this plant winds?? Movement to bring peduncle into contact — In greenhouse a leaf which had clasped stick has slipped off & straightened itself: say occasionally happen with changed direction of growth.—

The bent footstalk become more rigid than ordinary footstalks. Occasionally two twine.— Begin by saying everyone who has looked at this plant will have seen it climb by footstalks.

[2v]

T. elegans clasps by leave in same manner but in less degree & flower capsule peduncle do the same, — The peduncle of leaves are very slightly sensitive to touch.—

Shoot apparently does not rotate.

Aug. 12th I can see no movement, in tip of shoot.


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Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

File last updated 19 July, 2023