RECORD: Darwin, C. R. [1863].08.26-27. Cobaea scandens. CUL-DAR157.2.5-6. Edited by John van Wyhe (The Complete Work of Charles Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Transcribed by Christine Chua and edited by John van Wyhe 5.2023. RN1

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin.

The volumes CUL-DAR157.1-2 contain notes, abstracts etc. for Darwin's long paper and later book Climbing plants (1865). It was also commercially available as a softbound offprint, F834, F834a. See R. B. Freeman's bibliographical introduction. Items CUL-DAR157.11-60 were in a folder marked "Twiners". Items CUL-DAR157.61-112 were in a folder marked "Leaf-climbers" and items CUL-DAR157.114-147 were in a folder marked "Tendrils".


(1

Aug. 26. Cobæa scandens

Tendril long at end of leaf.— straight with alternate branches— each branch several times divided, & branchlet terminated by little hooks. —

The le internode & leaf-stalk & tendril all revolved in straight line in hot-house, pointing to window (carrying young [internode] & leaf), inclined when [illeg] front light & almost perpendicular when passing over axis of leaf. I should think base of leaf revolved—

Revolution with watch in 1°. 415'!! I tried repeatedly rubbing main central branch & branches could not perceive at all sensitive—

There are kinks.— The tip of each branchlet is terminated flattened a little & ends in double head, very shape sharp horny hook, —will even stick to finger a little. These 2 hooks being horny cannot

[1v]

Hooks will catch stick, though so minute.

I counted on one tendril 36 double Hooks

It is very curious that yesterday though t. moved with watch so quick yet was in the least sensitive, & today extremely sensitive — I have seen same thing with Passiflora punctata, revolved but was not sensitive, now both — In Echinocystis sensibility remained after revolution ceased — not perfect arrangement.—

(2

Cobæa

move; but ends of branchlets twine round & round object in excessive complication & contract in spire— part sp twisted round any object becomes very stiff.—& strong after the first revolution — tendril & all oscillated backwards & forwards & did not proceed in single course.—

Aug 27 I find that touching main stem or main branches or branchlet sub-branch all in a few minutes curve greatly in any direction.

Except (as far as I have seen) internally [sketch]

Sides a a a touched do not close in; & wd not likely to be touched— by (No No)

By touching in various point at same time tendril assumes extraordinary shapes.

This is the most highly organized tendril, which I have seen — Bignonia unguis the lowest.—

(3

Cobæa

The branches have soon quicker powers of straightening after a curving from a a considerable touch—; they will do it in from 30' to 60'— when even after being much curved.—

Revolving The tendril revolves about 20 times in 25° or 26°.—

Tendril & internodes revolved to day in 1°. 15' & a second time in 1°. 20', but then I accidentally bent tendril during its revolution. — (This tendril moved at rate of 1°. 15', whilst I irritated & made many branches curl. —

The main stem of tendril is sensitive except towards base; the first & second branches do ditto; but the very short penultimate & ultimate bifurcation are not sensitive: their function is when a branch caught to wind round & round it & interlock with their double hooks & this they do most effectively

[sketch] — see p. 4 a = sensitive — main stem of tendril

[3v]

[calculations not transcribed] 27 Revolutions

(4

Cobæa

Another € in exactly 1°. 15' (this is third €, besides one of 1°. 20; when tendril was suddenly bent.

Every breath of lightest air blows tendrils about, wind must aid in climbing & hence the use of the hooks. —

The penultimate branchlet, I now find to converge & diverge on touch; but, I think, less readily than others. — If terminal are not irritable partake of nature of 2 horny claw which no doubt represent another represent another bifurcation.

Another € in 1°. 23

I now find ultimate minute branchlets converge when stick rubbed between them & more conspicuously diverge when rubbed externally. Hence all, except hooks are sensitive:—

The tendril is a tall branched tree with every part sensitive & bending towards an object which just touches it.—


Return to homepage

Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

File last updated 21 July, 2023