RECORD: Darwin, C. R. [1863].08.26-27. Cobaea scandens. CUL-DAR157.2.5-6. Edited by John van Wyhe (Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Transcribed by Christine Chua and edited by John van Wyhe 5.2023. RN1

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin.

The volumes CUL-DAR157.1-2 contain notes, abstracts etc. for Darwin's long paper and later book Climbing plants (1865). It was also commercially available as a softbound offprint, F834, F834a. See R. B. Freeman's bibliographical introduction. Items CUL-DAR157.11-60 were in a folder marked "Twiners". Items CUL-DAR157.61-112 were in a folder marked "Leaf-climbers" and items CUL-DAR157.114-147 were in a folder marked "Tendrils". Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin.


(1

Aug. 26. Cobæa scandens

Tendril long at end of leaf.— straight with alternate branches— each branch several times divided, & branchlet terminated by little hooks. —

The le internode & leaf-stalk & tendril all revolved in straight line in hot-house, pointing to window (carrying young [internode] & leaf), inclined when [illeg] front light & almost perpendicular when passing over axis of leaf. I should think base of leaf revolved—

Revolution with watch in 1°. 415'!! I tried repeatedly rubbing main central branch & branches could not perceive at all sensitive—

There are kinks.— The tip of each branchlet is terminated flattened a little & ends in double head, very shape sharp horny hook, —will even stick to finger a little. These 2 hooks being horny cannot

[1v]

Hooks will catch stick, though so minute.

I counted on one tendril 36 double Hooks

It is very curious that yesterday though t. moved with watch so quick yet was in the least sensitive, & today extremely sensitive — I have seen same thing with Passiflora punctata, revolved but was not sensitive, now both — In Echinocystis sensibility remained after revolution ceased — not perfect arrangement.—

(2

Cobæa

move; but ends of branchlets twine round & round object in excessive complication & contract in spire— part sp twisted round any object becomes very stiff.—& strong after the first revolution — tendril & all oscillated backwards & forwards & did not proceed in single course.—

Aug 27 I find that touching main stem or main branches or branchlet sub-branch all in a few minutes curve greatly in any direction.

Except (as far as I have seen) internally [sketch]

Sides a a a touched do not close in; & wd not likely to be touched— by (No No)

By touching in various point at same time tendril assumes extraordinary shapes.

This is the most highly organized tendril, which I have seen — Bignonia unguis the lowest.—

(3

Cobæa

The branches have soon quicker powers of straightening after a curving from a a considerable touch—; they will do it in from 30' to 60'— when even after being much curved.—

Revolving The tendril revolves about 20 times in 25° or 26°.—

Tendril & internodes revolved to day in 1°. 15' & a second time in 1°. 20', but then I accidentally bent tendril during its revolution. — (This tendril moved at rate of 1°. 15', whilst I irritated & made many branches curl. —

The main stem of tendril is sensitive except towards base; the first & second branches do ditto; but the very short penultimate & ultimate bifurcation are not sensitive: their function is when a branch caught to wind round & round it & interlock with their double hooks & this they do most effectively

[sketch] — see p. 4 a = sensitive — main stem of tendril

[3v]

[calculations not transcribed] 27 Revolutions

(4

Cobæa

Another € in exactly 1°. 15' (this is third €, besides one of 1°. 20; when tendril was suddenly bent.

Every breath of lightest air blows tendrils about, wind must aid in climbing & hence the use of the hooks. —

The penultimate branchlet, I now find to converge & diverge on touch; but, I think, less readily than others. — If terminal are not irritable partake of nature of 2 horny claw which no doubt represent another represent another bifurcation.

Another € in 1°. 23

I now find ultimate minute branchlets converge when stick rubbed between them & more conspicuously diverge when rubbed externally. Hence all, except hooks are sensitive:—

The tendril is a tall branched tree with every part sensitive & bending towards an object which just touches it.—


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Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

File last updated 21 July, 2023