RECORD: Darwin, C. R. & Francis Darwin. 1879.04.26-05. Phaseolus multiflorus. CUL-DAR209.11.138-143. Edited by John van Wyhe (Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Transcribed by Christine Chua and edited by John van Wyhe 7.2023. RN1

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volume CUL-DAR209.11 contains material for Darwin's book Movement in plants (1880).


4

Jar E

April 26th 1879 Phaseolus — Radicles Black Grease

all marked 10mm— grease tips 3 m.m— over water in dark. — grown in sawdust —on chimney-piece now placed on study Table.

Temp. 61°-62° F.

(1) Tip. black 4°. 50 (8° 20') no geotropism straight— length from dot 11 1/2 mm. (27' 8° 15' a.m— ie 24 hr. tip geotropic for length of only 4 mm— length from dot to apex 14 1/2 mm

(2) Tip Black 4° 5 length 15 1/2 mm— basal part [sketch] bowed down but was so originally not geotropic to certain extent (27' 8° 15 on same position as yesterday, length 28 m.m.

(3) 0 4° 50' about 50° from vertically geotropic (27 8. 15' less geotropic length 24 m. (All radicles too dry)

(4) Tip black 4° 50 bowed a little laterally, not geotropic at all, rather reversed, length 13 mm. (27' 8° 15 tip bowed laterally & downward, length. 18mm. slightly geotropic

5 0 4° 50' vertically geotropic (27th 8°. 15' less geotropic, length 24 1/2 mm

6 0 4 '50 about 40° from vertical (27' 8'. 15' less geotropic length 24 1/2 mm

(7) Tip black 4° 55' a trace of geotropism.— length 13 mm. (27th 8° 15 barely a trace of geotropism length 17 1/2, length of black tip itself 9 m.m

8 (0) 4° 55° about 20° from vertically geotropic (27' 8° 15 less geotropic, length 17 1/2 19. mm length of black tip 9 mm.—

The chief part of curvature rather variable from 4 to 6 mm from apex

[4v]

[calculations not transcribed]

Lessened increase might be expected as tips grow so rapidly.

(5

Jar D. Ap. 26.th. 1879 8° 55' a.m Phaseolus tips with Black grease all like Jar of same date But placed on Chimney Piece.

(1) (0) 4. 50 pm Temp 70° about (I suspect too hot & too dry)

Abt 45 below horizon length 14 mm {bent near to bean} length 14 mm

(27th 8° 30' like yesterday or less geotropic length 28 mm slightly geotropic

(2) Grease tip about 20 below: root nearly straight length 14mm

(27th 8° 30' rather more geotropic than (1)— length of Black tip itself 11 m length from dot 21 mm slightly geotropic

(3) Grease tip about 45 below, only black is 45 below, the rest of root is apogeot (slightly above horizon length 13 about

(27 .8. 30' not at all geotropic— length of black tip —10mm length from dot 20 mm

(4) Grease tip abt 20— 30 below (only black tip) rest of root above horizon 17?

(27th 8. 30 length of black tip. 13 mm length 28 mm.

(5) Grease tip. abt 15 below (tip); root strongly curved in horizontal plane

(27th? (8° 30 obliquely geotropic— more lateral than geotropic length of black tip 10 mm length from dot 19 mm

(6) 0 abt 45 below, base of root slightly below horizon 14?

(27th 8° 30 slightly geotropic length 25 mm

(7) 0. about 45 below, base of root slightly below horizon 14?

(27th 8° 30, slightly geotropic length 24 mm [sketch]

[5v]

[calculations not transcribed]

6

Jar E

May 1. 1879 8° 45' am

Phaseolus Radicles Tips Black grease

suspended horizontally over water in dark germinated in sawdust

(1) grease 1 mm in length 4. 30' slightly geotropic, not nearly so much as numbers (8) to (11) (2d 8° a.m only a trace of g. length 17 mm

(2) grease 1 m in length 4° 30' almost rectangularly geotropic —(2d 8' Geotropism decreased.

5 case with 3 m.

(3) grease 3mm in length 4° 30 only a trace of geotropism— length 12 mm (2nd 8° a.m. length 14 m.m, only slight geotropism)

(4) grease 3 mm in length 4° 30 moderately geotropic; nothing like 8° to 11.

(5) grease 3 mm in length 4° 35' only a trace of geotropism, length 11 mm (2d 8° a.m only a trace of a trace of G. length 13 1/2 m.m.

(6) grease 3 mm in length 4° 35 quite straight & horizontal — length 12 m.m. (2d 8° now curved in opposition to G.

(7) grease 3mm in length 4° 35 slight geotropic close to tip

(8) (0) 4° 35 rectangularly geotropic

(9) (0) 4° 35 do— length 13mm. & this represent apparent general growth

(10) (0) 4° 35 do

(11) (0) 4° 35 almost rectangularly geotropic} chief part of curvature 5-6mm from apex.} 2d 8° a.m all still strongly geotropic

(7

(Jar A

May 1. 1879 8° 20' a.m. Phaseolus Radicles Tips Black grease suspended Horizontally over water— in dark— germinated in sawdust.

(1) (0) 4° 25 slightly geotropic (2d 7° 45 a.m 22° from vertical, length 17 mm

2 mm

(2) grease 2m in length 4° 25 more geotropic than (1) much less than (8) & (9) (2d 7° 45 a.m hardly a trace of G. & curved a little laterally, length 15 mm.

(3) grease 2 mm in length 4° 25 like (2) (2d 7. 45° am hardly a trace of G. length 13 m.m.

(4) grease 2 m.m in length 4° 25 about as geotropic as (1) (2d 7° 45 — 35° from vertically geotropic

(5) grease 2 mm in length 4° 25 very slightly geotropic (2d 7° 45 61° from vertically geotropic

(6) grease 2 mm in length 4° 25 considerably geotropic, not as much as 8 & 9 (2d 7°. 45 strongly geotropic (& curved laterally) like 8, 9, & 1

(7) grease 1 mm in length 4. 25 like (6) (2d 7° 50' strongly geotropic, like 8, 9 & 1

(8) (0) 4° 25 rectangularly geotropic (2d 7° 50' rather less geotropic, but strongly so

(9) (0) 4° 25 almost rectangularly geotropic (2d rather less geotropic, but strongly so

4° 25 I conclude that 2 mm of grease only slightly lessen geotropism after 8° 10'. (2d 8° a.m, the 3 without caustic have increased more in length, judging by eye than the blacks


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Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

File last updated 20 August, 2023