RECORD: Darwin, C. R. & Francis Darwin. 1879.06.23-30. Geotropic Beans Caustic. CUL-DAR209.11.178-182. Edited by John van Wyhe (Darwin Online, http://darwin-online.org.uk/)
REVISION HISTORY: Transcribed by Christine Chua and edited by John van Wyhe 7.2023. RN1
NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volume CUL-DAR209.11 contains material for Darwin's book Movement in plants (1880).
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Geotropic Beans Caustic I
8.55 am Jun 23' 1879
Faba germinated in sawdust How many & employed when roots are from 3 to 5 cm in length. Each one marked at 10 mm with asphalt dot + root. Placed in fine ؟ Placed Horizontally sieved earth rather dry in a vessel perforated bottom & top; some were planted rather deep but most had only 2 cm of earth over them. Caustic with pointed bit of Ag No 3 I think I rather more than usual
Some Phaseolus multiflorus in the same way— Vessel stands on table in big room. Temp at noon 20° C
Examined at 6.35 pm. i.e after 9° 40'
Faba- caustic 16 mm, 18 mm 18, 15, 16, 20 — none of these show any geotropic bending
The non-caustic ones were harder to measure because of being bent —
|
mm 10-20 |
sharply bent at roughly 10 mm from tip only faintly bent— marked with a pin sharp bend sharp bend at roughly 10 mm from tip |
[in margin:] All not measured first day
Phaseolus. caustic marked at 10 mm. 15 mm, 19 mm. 14 mm none bent
after 9° 40' non-caustic Controls |
i ii iii |
14 mm. faintly bent —no mark — sharply bent 15 sharply bent.} increased to same extent |
Now put back in earth except 3 Fabas 2 of which which were v well bent & the 3rd was a dwarfed (? caustic) (? non caustic) oddly bent one not recorded above.
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Geotrop. Beans Caust.
June 24. 10 am — They were taken out about 9 & put in a plate of water.
∴ we may say after 24 hr
Faba caustic.
1. 27mm bent sideways not geotropic
2 28 just perceptibly geotropic
36 basal 21 straight & apical bent say 15 below horizon.
average 37 m.m 9 caustic
no mark — no geotropism
no mark Clear geotropism at 5 mm from tip
27 — basal 14 straight, apical 13 bent at say 30 below
30 22 straight (basal); 10 8? apical crook geotropic
32 20 straight 12 crooked but in a horizontal plane with perhaps some geotropism
35 22 straight, 13 about 30° below horizon
9 caustic
Faba non caustic 4 non
31 bent geotropically throughout whole length about 30 below
26 not well geotropic
37 — 10 basal straight 27 crooked well geotropic
27 5 basal straight 22 geotropic
4 non
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Geotropic Caustic Beans I
June 24th 10° a.m say 9° ie aft 24° After the earth
Phaseolus caustic
23 mm, basal 13 mm straight apical 10 about 10 below horizon
28 straight
22 straight
Non-caustic
25—basal 6 straight, apical 19 nearly vertical.
26—basal 5 straight; apical 21 abt 45° below
25. basal 8 straight apical 17 abt 45° below} Control
[4]
Faba (caustic) Geotropism
June 29. 11 45 am caustic some abo above & some below in both cases at the extreme tip (say 21 hr) [calculations not transcribed]
June 30 — 8. 35 am
Three cauterised on the upper side are sharply bent downwards
Three on the lower side only faintly bent as shown in sketches. In No 3 the black at the tip was very faint
They were all marked at 10 mm from tip but only the marks were hidden by dirt—: in most No 3 measured 26 mm—
The Three control beans to which nothing was done are fairly bent but no as well as they ought to be
Goebel picked out 2 of the 3 upper caustics as the best geotropic of the lot, & 2 of the under caustics as the least bent [sketch]
[4v]
[calculations not transcribed]
[182]
It is clear they are not well geotropic, Sachs thinks earth dry — Goebel made sections of Faba roots; the root cap is killed up to but not all of it ؟ when it dies away is is clear uninjured; the tissue of the root is normal except close within the root cap where a row of 6-8 cells is dark orange, G says killed. He expressed it by saying punctum veg was killed
some radicle were examined by an experience physiologists
Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)
File last updated 21 August, 2023