RECORD: Darwin, C. R. & Francis Darwin. 1879.06.23-30. Geotropic Beans Caustic. CUL-DAR209.11.178-182. (John van Wyhe ed., 2002-. The Complete Work of Charles Darwin Online, http://darwin-online.org.uk/)
REVISION HISTORY: Transcribed by Christine Chua and edited by John van Wyhe 7.2023. RN1
NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volume CUL-DAR209.11 contains material for Darwin's book Movement in plants (1880).
(1
Geotropic Beans Caustic I
8.55 am Jun 23' 1879
Faba germinated in sawdust How many & employed when roots are from 3 to 5 cm in length. Each one marked at 10 mm with asphalt dot + root. Placed in fine ؟ Placed Horizontally sieved earth rather dry in a vessel perforated bottom & top; some were planted rather deep but most had only 2 cm of earth over them. Caustic with pointed bit of Ag No 3 I think I rather more than usual
Some Phaseolus multiflorus in the same way— Vessel stands on table in big room. Temp at noon 20º C
Examined at 6.35 pm. i.e after 9º 40'
Faba- caustic 16 mm, 18 mm 18, 15, 16, 20 — none of these show any geotropic bending
The non-caustic ones were harder to measure because of being bent —
|
mm 10-20 |
sharply bent at roughly 10 mm from tip only faintly bent— marked with a pin sharp bend sharp bend at roughly 10 mm from tip |
[in margin:] All not measured first day
Phaseolus. caustic marked at 10 mm. 15 mm, 19 mm. 14 mm none bent
after 9º 40' non-caustic Controls |
i ii iii |
14 mm. faintly bent —no mark — sharply bent 15 sharply bent.} increased to same extent |
Now put back in earth except 3 Fabas 2 of which which were v well bent & the 3rd was a dwarfed (? caustic) (? non caustic) oddly bent one not recorded above.
(2
Geotrop. Beans Caust.
June 24. 10 am — They were taken out about 9 & put in a plate of water.
∴ we may say after 24 hr
Faba caustic.
1. 27mm bent sideways not geotropic
2 28 just perceptibly geotropic
36 basal 21 straight & apical bent say 15 below horizon.
average 37 m.m 9 caustic
no mark — no geotropism
no mark Clear geotropism at 5 mm from tip
27 — basal 14 straight, apical 13 bent at say 30 below
30 22 straight (basal); 10 8? apical crook geotropic
32 20 straight 12 crooked but in a horizontal plane with perhaps some geotropism
35 22 straight, 13 about 30º below horizon
9 caustic
Faba non caustic 4 non
31 bent geotropically throughout whole length about 30 below
26 not well geotropic
37 — 10 basal straight 27 crooked well geotropic
27 5 basal straight 22 geotropic
4 non
(3
Geotropic Caustic Beans I
June 24th 10º a.m say 9º ie aft 24º After the earth
Phaseolus caustic
23 mm, basal 13 mm straight apical 10 about 10 below horizon
28 straight
22 straight
Non-caustic
25—basal 6 straight, apical 19 nearly vertical.
26—basal 5 straight; apical 21 abt 45º below
25. basal 8 straight apical 17 abt 45º below} Control
[4]
Faba (caustic) Geotropism
June 29. 11 45 am caustic some abo above & some below in both cases at the extreme tip (say 21 hr) [calculations not transcribed]
June 30 — 8. 35 am
Three cauterised on the upper side are sharply bent downwards
Three on the lower side only faintly bent as shown in sketches. In No 3 the black at the tip was very faint
They were all marked at 10 mm from tip but only the marks were hidden by dirt—: in most No 3 measured 26 mm—
The Three control beans to which nothing was done are fairly bent but no as well as they ought to be
Goebel picked out 2 of the 3 upper caustics as the best geotropic of the lot, & 2 of the under caustics as the least bent [sketch]
[4v]
[calculations not transcribed]
[182]
It is clear they are not well geotropic, Sachs thinks earth dry — Goebel made sections of Faba roots; the root cap is killed up to but not all of it ؟ when it dies away is is clear uninjured; the tissue of the root is normal except close within the root cap where a row of 6-8 cells is dark orange, G says killed. He expressed it by saying punctum veg was killed
some radicle were examined by an experience physiologists
Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)
File last updated 21 August, 2023