RECORD: Darwin, C. R. & Francis Darwin. 1878.07.22-29. Trifolium subterraneum/ Draft of Cross and self fertilisation (several fragments). CUL-DAR209.11.232-236. (John van Wyhe ed., 2002-. The Complete Work of Charles Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Transcribed by Christine Chua and edited by John van Wyhe 7.2023. RN2

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volume CUL-DAR209.11 contains material for Darwin's book Movement in plants (1880).


[232]

July 22/ subterraneum Trifolium

Young— Flower head — not fully open (?) or possibly just begun to fade??

standing vertical — stem tied

Roof

11° first dot
12. 25 far to Left
12. 56 — back
2° to me — .very little
3° to left.    decidedly zig-zag
4° to left & to me
5 10 on vertical part of Frame just on vertical glass ?
7. 16 20' to me & to right. too great a crowd of m. Vertical glass
7° 20 down & to right
8. 37' much to right & down
10. 50 nearly same course

(The peduncle did not become more than rectangular with shoot) & flower only at ∟' angle —

(N. B the lifting up of stick & shoot prevented the peduncle from becoming perpendicularly down)

[232v]

[right side of page excised. The text is here combined with that of the excised half in CUL-DAR209.11.233v]

753 (610

Ch XII

Gen Con

cross-fertilisation are transmitted in the case of by plants to the next generation;

and judging from the varieties of the common pea to

many succeeding generations. But this may merely

be that the crossed plants of the first generation are

extremelyvigorous, and that they, transmit their organs like any other character to

their successors.

(Notwithstanding the evil which many plants suffer

from self-fertilisation they can be thus propagated by this means under favourable

conditions for many generations as shown by [illeg] some of my experiments,

and more especially of by the survival during at last

half-a century of the same varieties of the common

pea & sweet-pea, as well probably as of several other

exotic plants which are never or most rarely cross-

-fertilised in this country. But all these plants, as far

as they have been tried, profit greatly by a cross with

a fresh stock. Some few plants, for instance Ophrys

apifera have almost certainly been propagated in a

state of nature for rendered of thousands of

generations without having been once intercrossed

[Cross and self fertilisation, p. 439.]

[233]

subterraneum z

Head withered inclined 36° below Horizon stem tied.

left Hand nearer base of peduncle      oldest Head Right Hand on Head

11°. 5' 11° 5

12. 27. down   12. 27' down & to R.

12. 57 up & to R.        12. 57 up Back & parallel

2° down           2° down

3° up. !            3° down [sketch]

4° little up       4° almost same spot

5 10' a little to Left     5 10' back or up & parallel

6°. little to right          6 far up to left due to slewing of filament

7°. 15 too great a row of marks          7. 15. down again

            8° 40 up & to left 4 time down & 3 time up

            9 30 up & to left same course

            10. 55' vertically down

23d 6. 55'        6°. 55

8° 10' to left & down 8° 10 .   do

9° 12 back to right      9° 10 to left

(These 2 parts do 

            not circumnutate together)

10°. 10'            10° 10' down & left

            (Used)

no new marking more

25 a 9° am

(sunk below diagram as much as diagram

cal

26th ?

filament interfered with action.

[233v]

[see page 232v above]

[234]

First Flower

On Roof

July 22d 4° 35 P. m    

[sketch]

5. 8 a little to left       

6° to left & to me       

7 20  to me & to right            

8 40 to me & to left   

9 27 to me & to right } is bending down & zig zagg in

10. 53 — to me & little to left            24th. 6° 40 ......

23d 6° 55         8° 7 to me & to left

8°. 5 from me 8. 30 a little to left

8° 40 to right   9. 30 to me

9. 15 to left     10. 25 to R & from me

10. 17 nearly same course short ways            11. 50 a little to left

11. 8. to right 1° direct to me.

12. 25 same course     2° little to L

1. 30    do        3 to me

2. 55    nearly same spot — an atom 4' to left

4°.    do;    do 5 back to right

5°    do.    do (watered sand)   6 from me

6° — a little to right   7 15 to me

7° 15 some way to right          25th 6° 45' ......

9° — do    do  

10. 40 from me           

(All used)

[234v]

"a result which is probably effected by mixing the slightly-different physiological units of slightly-different individuals."* (a) But we must not allow this or any other highly generalized view, or the more probable analogy of chemical affinity, to conceal from us our ignorance. We do not know what is the nature or degree of the differentiation in the sexual elements which is favourable for union, and what as in the union case of distinct species, is injurious. We cannot say say why the individuals of certain species profit greatly, and others very little by being crossed. There are some few species which have been self-fertilised for a vast number of generations, and yet are so vigorous that they can compete successfully with a host of surrounding plants. Highly self-fertile varieties

[Cross and self fertilisation, p. 458.]

[235]

First Flower Roof

26th. 6° 40' ⨀

8° from me.

9°. 10' I think gone back at a bit

10 . 40'— to left & from me

12° to me & to right

1°. 45' to left

3° little to me

4. 35 to right

6 a little to right

9' same course, short way

(used)

Tracing IV A

[235v]

act in this manner both with

, parts of Europe, in the U.

& therefore probably in all p

which actually depend on the entire

=ation, the result of holes being bitten in sterile

are capable of fertilizing

there can be no cross-

The extent to which hum

is surprising: a remarka

near Bournemouth, wh

with Erica tetralix grow

walk and every now & then

I had got a handful

[Cross and self fertilisation, p. 426.]

[236]

Monday 29' T. subterraneum (First flower)

[236v]

737 (23

Ch XI

lyx. In these several cases the perforations

ide, but in Antirrhinum majus little

e made on the lower side, close to the

rance which represents a nectary, and

n front of & close to the spot where the

(But the most remarkable case of skill

un to me, is that of of the manner in which perforation of the flowers of Lathyrus sylvestris,

my son Francis* (*Natures Jan 8. 1874

r in this plant is enclose within a tube,

[Cross and self fertilisation, p. 428.]


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Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

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