à (1

Oxalis carnosa

The main peduncle, bears 3 or 4 long sub-peduncle with yellow flowers directed upwards, long straw fixed within calyx, & 2 triangularly sights fixed to straw, traced on Horizontal glass (Tracing Used) feebly feebly illuminated from above in my study. See tracing. Distance from glass to base of sub-peduncle 8 inches— length of sub-peduncle which alone makes most. 1.65. I believe this will show how much tracing magnified. There can be no doubt about circumnutation.— These flowers were observed, because each sub-peduncle bends vertically down, after the flowers have withered. On the morning of the 14th the flowers observed had withered— Filament fixed within calyx — Peduncle observed from 9° a.m on 13th to 9° 20' a.m on 15th— [sketch]

Used under circumnutation of Flower-stems

Light

N.B. the leaves of of this plant after having been during 2 days illuminated very feebly in my study from vertically above showed hardly a trace of going to sleep.— (over)

[63v]

[calculations not transcribed]

Mem. Pfeffer movements of Flowers

[illeg] — Also Kerner

Left during the 15th in front of N.E. window & at night leaves went to sleep, so that little more illumination during day induce sleep.— 16th morning leaves awake.—

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Oxalis carnosa

circumnutation of old Flower-peduncle

The same lately withered flower which was Observed 13-15 morning — again observed also bearing 1 developed pod, in same manner with same straw morning 16th —to morning of 17th i.e. after interval of 24° — Tracing shows still circumnutation; but main peduncle secured to stick, so whole movement in sub-peduncle & this is evidently chief source of movement. (See Tracing)

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Oxalis carnosa.

The same old flower-peduncle described pot observed 13h— to morning of 15th— after 24° p.2  16th to morning morning of 17' April still circumnutating (main peduncle still secured to stick) by the evening of Ap. 19' aft 36 hr had sub-peduncle projecting a little beneath a horizontal line, with same straw-mark attached to calyx.— It was observed on vertical glass till morning of Ap. 22d & Tracing A show course till about 6° P.m. on 20' descended in nearly straight line, but afterwards apparently zig-zagged. The

dots, however, cannot be fully trusted for straw now pointed so much down that the marks difficult to observe, & tracing necessarily exaggerated. The movement had become slower. (If seed-pod circumnutates when almost vertically down, then the zig-zag line may be trusted but not otherwise).

Ap 22d 9° a.m. The straw was fixed on same flower- sub-peduncle transversely to top of calyx, so that the movement cd be more easily observed. The sub-peduncle now stood at angle of beneath horizon nearly perpendicularly downwards in the plane passing through stem of plant; but in a plane at right angles to this only 60° below horizon, but it did not move sensibly all day, nor next early next morning April. 23d: Afterwards on in the course of the 23d it kept almost in same spot, but fell a little during it fell a little & during the following night night &

On Ap. 22d 9° am. A straw was fixed transversely above base of calyx of another sub-peduncle which had already bent down to the perpendicular parallel with hang perpendicularly down in a plane passing through stem of plant; when viewed in one position, but travelled beyond this line when but in the plane at right angles to this not quite

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Oxalis carnosa

viewed in another position, that it what had occurred will be better thus expressed the sub-peduncle was originally inclined upwards, it must have sunk till horizontal, then sunk to perpendicularly downwards & then travelled to one side. This latter movement was proceeding all the 23d & the line traced was slightly zigzag; during the night it rose & continued rising

perpendicular. Its movement was observed from 9° a.m on 22d to 9° a.m on 24 & during the whole of this time it nutated in this plane; or may be said to have nutated laterally. The movement is shown in tracing on vertical glass. fig (z)

The men meaning of movement going on on ie. growth going on after the deflexion of the sub-peduncle & whilst pod maturing is that it has to turn up. before seeds shed. In Tracing (z) enough zig-zag to indicate some circumnutation— certainly circumnutation laterally

[calculations not transcribed]

(Ap. 24th red worsted I see one sub-peduncle now slightly bowed which is almost ready to shed seed; as pod stands perpendicularly upwards there must have been movement also at the joint— The sub-peduncle is always straight during the primary depression-movement—)

Ap. 23d. When the sub-peduncle is perpendicularly downwards, the 5 long & unequal sepals are becoming are closed over the ovarium, the walls of which are very thin. When mature, the ovarium opens to scatter its seeds & the sepals are then widely reflexed, & as I believe before the scattering the sub-peduncle becomes bowed in middle part so as to elevate per vertically upward the ovarium & then scatter seed as far as possible. The bending now is not at the joint of sub-peduncle as hitherto.— (over)

[4v]

Ap. 24 Doubtful point how much the joint & how much whole sub-peduncle bends, when turning up to scatter seeds & whether it then circumnutates.

5

Oxalis Carnosa

(Light excluded except dull from above)

Two young Plants, not very vigorous— the main flower-peduncles in 2 cases was pulled down till they were horizontal & the flo flower sub-peduncles, after flowers had withered, moved towards main flower-peduncle forming angle of about 40° with it, & they showed no tendency to move towards centre of earth. Therefore their position due to Epinasty; for they held the same position relatively to main peduncle as if this had remained upright. (We may naturally suspect that epinasty & hyponasty the inherited effect of geotropism or apogeotropism, or of heliotropism or apoheliotropism.) [sketch]

(A sub-peduncle had filament affixed to it, after it had moved towards main peduncle & its movement traced on vertical & horizontal glass— see Tracing, but it never become quite upright, for it then withered & died, after the pod had opened, which contained contained only a few & poor seeds — The rising was due to bending upwards of middle of sub-peduncle.— & may certainly be attributed to apogeotropism. The tracing not worth giving, much magnified, yet the movement was zig-zag, with two little circumnutations in the course of two days— Certainly modified circumnutation.—