RECORD: Darwin, C. R. 1878.09.19-20. Phalaris apogeotropism. CUL-DAR209.11.97. Edited by John van Wyhe (Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Transcribed by Christine Chua and edited by John van Wyhe 7.2023. RN1

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volume CUL-DAR209.11 contains material for Darwin's book Movement in plants (1880).


[97]

Sept 19th 1878 Phalaris apogeotropism

Two seedlings inclined at 40° below horizon, [sketch] with glass filaments attached & triangular marks, kept in darkness; were traced first on vertical glass & then on Horizontal glass.—

First dot 10°. 5'

Left-Hand seedling rose in almost straight line till 2° P.m. & then diverted from straight line (Height of seedling 1.3. inch. / & stood vertical at 2° 30' vertical ie in 4°. 30' (so had then bent 130°) & then diverted from straight line betw 3° & 4° circumnutated about same spot for rest of evening, the night & next morning. Hardly any gradation between cir the vertical rising through apogeotropism & circumnutation.

Right-Hand seedling 1.3 of inch part length of cotyledon which bent. (See Tracing)

Distance from soil to vertical glass 9.1 11.2 inches— distance to horizontal glass. 9.1, so the part tracing which shows circumnutation less magnified than on vertical glass. The degree of magnification Taking centre which can be calculated holds good for centre of circumnutation & for point where it is stated that Cot. stood horizontal.

First dot made at 9° 30' a.m; Sept 19th but the Cot had already risen considerably— It then rose in wonderfully straight line till 1°. 10' or better 1° 34' (ie for 5° during 4°), but then bent to one side & showed plain tendency to circumnutate. until — it subsequently at about 45° 15 stood quite vertical & had reached now circumnutated about same spot, During day rest of evening, the night & next morning, having bending however, bent a little laterally on the morning of the 20th

This diagram shows clearly a vertical rise from apogeotropism passing into through irregular circumnutation into regular circumnutation.— Perhaps put it vice versâ.


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Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

File last updated 19 August, 2023