RECORD: Darwin, C. R. 1877.10.04-05. Oxalis acetosella / Draft of Descent vol. 1, folio 34 & 32. CUL-DAR209.14.67-68. (John van Wyhe ed., 2002-. The Complete Work of Charles Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Transcribed by Christine Chua and John van Wyhe, edited by John van Wyhe 7.2023. RN2

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volume CUL-DAR209.14 contains material for Darwin's book Movement in plants (1880). Draft is in the hand of Ebenezer Norman with corrections by Darwin. The text of the draft corresponds to Descent 1: 277-8.


[67]

1877 Oct. 4' Oxalis acetosella— youngish leaf oblique to light— somewhat hidden amongst other leaves— day high clouds

8º 5'

8º. 33    far down

8 47    so much down that I had to raise pot  in interval no doubt went down

8. 55'    new dot

9º 17    up ! } shaken

9º 35.    very short distance to light

10. 25    down— short distance— Then far down probably due to shaking X ↙

{ 10º 35    new dot high on glass & new leaf

10 54'    not moved

11º 24    risen greatly

11. 55    up & little from light highest point — (The old leaf has now risen much & come into sight & I have made blue dot

12º 11    down 12º 11    2d new dot down

12º 25    do      12º 25    do & from light

12 53'    up      12. 53    — to light

1.º 40'    down             1. 40'    tremendously down

2º    up 2º    up

2º 36    up & a little to light   2º 36    down

2º 55    a very little up.          

   up

3º 32.    down  3º. 33    to light

4º 47    4º 57'

5º 30    5º 30'

            53 [-] 25 [=] 28

55 [-] 46 [=] 9

7º    perfectly asleep with Bristle perpendicularly down       11º 55. to 12. 25' down = 30'

            12. 25 to 12. 53    up    28' shorter distinct

            12. 53 to 1. 46    down    53'

            1 46 to 2. 55— up    69'

5th 6º 45'    low down belong to upper figure            2. 55 to 3. 22 down    27'

7º 5'    vertically up    30 [+] 53 [+] 27 [=] 110 [÷] 3 [=] 37' aver

7º 30    still up 28 [+] 69 [=] 97 [÷] 2 [+] 48 average

7º 45    same spot       

9º 52º Oct 5'

(Used)

[67v]

36 34

Chap. 8

to inherit their superiority. In regard to structures acquired through ordinary or natural selection, and adapted for special ends, there is in most cases, as long as the conditions of life remain the same, a limit to the amount of advantageous modification in relation to some special end; but in regard to structures stat/adapted to make one male victorious over another either in fighting or in charming the female, there is hardly any limit to the amount of advantageous modification; so that as long as fitting variations arise the work of sexual selection will go on. This circumstance may partly account for the frequent and extraordinary amount of variability of presented by secondary sexual characters. Nevertheless, natural selection will determine that no secondary characters shall be acquired by the victorious males, which is injurious to them in any high degree, by expending too much of their vital powers, or exposing them to any great danger.

The development, however, of certain structures—of the horns, for instance, in certain stags,—has been carried to a wonderful extreme; and in some instances, as we shall afterwards hereafter see, to a degree which must be slightly injurious to the male, as far as the his general conditions of life are concerned. From this fact we learn that the advantages which the favoured males have derived from conquering

[68]

Oct 5th Oxalis acetosella Same leaf in same position

7º 52' —    Bright morning

8º 12'    down

8. 23    from light & a little down.

8 40'    down

9º 4    up

9. 17    up & crossed blue line from light

9' 35    crossed first line again & back to light

10º 2'    risen a good deal

10 29'    down up

10 55    high up

11º 29'    down much

11º 45'    down

12º 6    still down

12 32'    much up

12. 52—    rising

1º 24—    short way to light— (problem condensed)

1. 53'    a little down on an old thin line

2.º 15    to light & a little down

2º 42'    straight down

3º 2'    rose an atom & little from light

3º 30'    straight up.

4º 3'    to light & rising a little.

4. 35'    a little way up

5º. 2'    down some little way

5 15—    a little down— I have not connected this dot with last

5. 36—    dot in square not connected— leaf apparently not asleep.

7º    asleep. with bristle vertically down—

Oct 6 ⨀ 6º 57'    — say 7º

7º 50'

(Used)

[68v]

34 32

Chap. 8

attractive females. But this view is not probable from what we know of the eagerness of habit of most animals, for generally the male in general, this view does not seem probable most cases is, eager to pair with any female. It is a more probable view when both sexes are equally possess the same ornaments= al that these ornaments were have been acquired through sexual selection by one sex alone, generally the male sex, but were but, and have then been not only equally transmitted to both sexes. through both sexes, as is always the case, but equally developed in both. If, indeed, during a lengthened period the males of any species had were greatly to exceeded the female in number and then during another lengthened period the reserve had were to occurred, a double but not simultaneous process of sexual selection might easily have been carried on, but not during the same period by which both sexes could ultimately be modified.)

We shall hereafter see that there are many animals with the sexes resembling each other alike which probably have been modified through sexual selection, although they are not brilliantly cl coloured, not provided with any special ornaments. Their absence of bright tints or other ornaments may have be the resulted from of variations of the right kind not never having arisen, or from of the animals themselves having preferred, improbable as it this will at first appear, simple colours, such as plain


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Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

File last updated 21 December, 2025