RECORD: Darwin, C. R. & Francis Darwin. 1878.07.31-08.01. Mimosa pudica / Draft of Cross and self fertilisation, folio 759. CUL-DAR209.2.138-139. (John van Wyhe ed., 2002-. The Complete Work of Charles Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Transcribed by Christine Chua and John van Wyhe, edited by John van Wyhe 5.2023. RN2

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volume CUL-DAR209.2 contains materials on circumnutation of leaves and stems for Darwin's book Movement in plants (1880).

The text of the draft corresponds to Cross and self fertilisation, pp. 442 & 443.


[138]

July 31 Mimosa pudica

Upper Yong leaf          Lower old & hardy sensitive leaf

8°   10  8° 10'

9. 16 down & little to right      9. 15'  down & little to right

10. 17 down     10. 17' down & to left

11. 34 up & to left

 11. 34' up & left

12  13 same course Temp 82° F.         12. 13 same course

12. 52 right up             12. 52  same spot

2° up     2° up

3   above glass             3° almost same spot

4°. 8' greatly fallen      4 .8' still up

4. 34° to right & little up         4.° 34' same course

5  9 down         5' 9  down

6°   much down            6..   down

8. 25 down ( resting on old immovable leaf.)

10. 5  fallen & to left  10.    fallen 5. 

Aug 1' 7° 10' far above glass  Aug 1' 7° 10'  much risen during night

8° 50'   coming on glass            8° 50  to left & down

9°  come on glass        9° co

10 23 risen & again off glass  10. 23  to right & down

 70°F. early morning             12°  5  to right & very little down

 76°. 10°. 30' a.m                

2° 5 still off glass         2.  5   nearly same spot

             4. 12 nearly same spot

(Used)

[138v]

(616

759

Ch XII

Gen: Con:

the two species of Lobelia & of the individuals of Nicotiana. The species belonging to distinct genera in the same family certainly differ in this respect. as may is be seen in my tables. The effects of cross-and self-fertilisation may be confined either to the growth or fertility of the offspring, but generally extends to both qualities. There does not seem to exist any close correspondence relation between the degree of adaptation of in flowers for cross-fertilisation such as for to the visits of insects of certain groups of insects, and the degree to which these offspring profit by this process we may easily err on this head, as cross fertilisation; but there there are two caused effectually ensuring means which ensuring cross-fertilisation which are not internally distinguishable perceptible, namely self-sel-sterility and the prepotent fertilising influence of pollen from another individual. Lastly it has been shown in a former chapter that there is no close correspondence between the number of seeds produced either by the parent-plants, or by their crossed  self-fertilised offspring, when they these are fertilised by pollen from another plant & & or by their own pollen, and the height, vigour or fertility of the offspring raised from from, such

[139]

Mimosa pudica

Augt. 2d

8° 30'

10. 10' down —

11  30   up & to right —

12. 12.   down. —

12. 38'   up — (to base of arm of arrow

1.  45   up

2  55 to right &   little up  

4  2' to left &     2.55       little up  

4. 30 down & an atom to right —

5 . 25 far down (leaflets asleep)

5  55    down

8  20'

3 down

2 up

 

Aug. 3d 7°. 7'

7° 25' down a little —

8. 37. down on nocturnal line

9. 35 risen greatly —

10. 35 to right & atom down —

11. 43 considerably down

12. 10 up again —  

12. 55 down —

1. 44 up —

2. 55 up considerably

4. 7   to right

4. 30 risen good way —

5 53'   up again!! —

7. 12     down —  

5 down

4 up

12 Hr

removed Plant

 

7° . 25' dot (cross show where wd have stood if not shaken.)

now Bell glass

8 25 up recovered shook probably

9. 3   up

9. 35 do

10. 40' very high up

12 50 (night) do do

rose

3. 55 down

6.15 a.m do

6. 40' risen a little

7  46   down

8 . 18   do

9 . 3  do

10 little way down

10 36'   up

11   down

down with

 2 rises   

Leaf with Petiole to end of Pulvn

   4 inches long   

[139v]

[right side of page excised]

Ch XII

Gen Con

seeds. This want of corr

from on the number of seeds des production number of seeds produced be

freely which reach reaching the ovules, and this a

reaction between the pollen an

tissues; whereas the growth ar

the offspring will be chiefly determined not

the ovules, but by the nature & perfection of the

their contents of the pollen-grains

→ (one line open)

(Two other A second A pollen important conclusion from my observation is

fertilisation do not result follow from

mere union of the sexual element of

their having been subjected

,or to their having varied

somewhat different condition

that in either case their sexual element

differentiation in their sexual

the inquiry from self-fertile

such differentiation in the sexual element. These

substantiated established by my exper

the Ipomœa and of the Mimulus

ilised for the last seven previous

all the time under the same


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Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

File last updated 20 December, 2025