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Strawberry Stolon.

May 18th A long stolon (Right Hand) projected almost horizontally, having being supported far from tip on stick & had filament fixed to extreme growing tip which was a little upturned & movement traced on vertical glass — [sketch] Kept in complete darkness — Tracing given in Fig Tracing I. — Made 2 very narrow ellipses went twice up & twice down in zig-zag lines of considerable length in 12°— By next morning stood higher than on first morning. (Another & younger stolon (Left -Hand) had filament fixed to it & tried to observe in same manner, but this stood up about 45° above horizon & it soon circumnutated up above vertical glass & did not come on glass again till next morning.)

We thus see not geotropic (given on p. 2.)

May 19th The glass filament was detached & fixed to stolon itself behind growing point which was upturned & was again observed in same manner. Tracing 2. shows movement from 8° a.m. on 19th to 8° a.m on 21st ie during 48 Hrs — During 19th in incessant movement chiefly up & down in vertical plane — During 14° 30' it went 5 times down & 4 times up on the first of these it travelled, besides travelling 4 times to right or less in direction more or less horizontal. On the 20th its course was even more equally complicated (& direction & can hardly be followed on the

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Stolons of Strawberry.

Tracing) & but it moved. The two about 5 times up & 5 times but little laterally down. during this day of within 16° & Tracing considerably magnified, but During part of time a daytime tracing was made independently by square a cube on 20th applied to vertical plate, by which the movement of apex was traced not at all magnified, & this shows that apex during 9° moved over distance of. 45 of inch; & during 25° including night travelled .82 of inch, but this includes general fall. Comparing position on morning of 19th to that on morning of 21' it is clear that apex of stolon had fallen considerably, but whether due merely to weight or to geotropism, no means of deciding.

I here allude to a rather younger stolon. Movement of this one traced (fig. 3) on 19th from 8° a.m to 10° 30' P.m.

The movement was here chiefly lateral — During part of day (i.e from 3° P.m. to 10° 30') tracing made on true scale with Cube (see fig 3. A) & this shows that the apex moved more than 1.1 of inch from side, & earlier in day moved, quite twice as much. Therefore movement of

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Stolon of Strawberry

stolon almost like that of climbing plant. It was of course growing rapidly.— On the 20th movement somewhat less complex (see Fig. 4.) & chief movement in an almost vertical plane; was traced for 13° 30'— At same time traced with cube on true scale & actual movement shown to be in vertical plane 1.25 of inch, extreme distance 1.55 of inch, & at right angles or in horizontal plane .95 of inch.— During neither day did this younger stolon seem to be acted on Geotropism, or by ordinary bending from weight

Apheliotropic on surface of sand

May 21' 10° Placed several 4 stolons Pointing from light. In G and in front of a sand Ground of sticks & grass— [illeg] to crawl through.

May 27th The 4 stolons have crawled through them; where now freed of all sticks two of them are straight & 2 somewhat & permanently sinuous: I think that circumnutation of some was in avoiding obstacles, but where end of stolon came into contact it grew upwards, (the apex always having a tendency to grow upwards) & this surmounted obstacle. Services apparently not nearly sufficient to account for circumnutation. (Shall rear under Saxifraga to this pit.)