RECORD: Darwin, C. R. [ny].07.10-13. Smithia pfundii / Draft of Cross and self fertilisation, folio 705. CUL-DAR209.3.23. (John van Wyhe ed., 2002-. The Complete Work of Charles Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Transcribed by Christine Chua and John van Wyhe, edited by John van Wyhe 11.2022. RN2

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volume CUL-DAR209.3 contains materials on Circumnutation of leaves and hyponasty for Darwin's book Movement in plants (1880).

Draft is in the hand of Ebenezer Norman with corrections by Darwin. The text of the draft corresponds to Cross and self fertilisation, pp. 408-09.


[23]

Smithia pfundii

—apex

glass

 

Left Hand Transverse filament           Right Hand —

shoot at rt angles         long filament shoot depending beneath right angle

July 10 8º 50' first dot            July 10th 8º 50' first

10º to left & up           10º up

11. 35 to left   11 . 35 up & to right

1º to left & little up    1º to right

2º up    2º down

3º to right        3º up again

4º to right        4º far up ( this means shoot depressed at apex

5º same course            5 very far up

6 to right & little down           6º so high cannot be traced do — near frame

7. 15 almost same spot            7º. 15 close to frame, still up.—

9º up & to left             9. mark on frame.

10. 40 up & to right    10º 40 a little down

 

11th 6º 35       11th 6º 35

8º little up       8º a good way down.

9. 7 do & to right        9. 7 far down

10 to right       10' far down

11º little to right         11' little down

12 up & little to right 12

1º nearly same course             1º do a little to left

2º to right & little down         2º up down

3— same course         3º straight down

4º to right & up — (whole shoot is rising which account for twisting to right. & zigzags.       X 4º  down— this due to shoot rising now at rt angles

5. 10 to right & down             5º 10 down

6. 10 down      6. 10 down

7. 15 to right & down             7º 15 do

9. 14. up        9 14. to left & down

10. 50 down parallel   10 50 down & to R

Height of Plant 1.ft 5in           Height of Plant 1.5 ft inch

ft 1. 10 inches

            Distance 9½ inchs — I cannot measure shoot)

12th 6º. 35 ⨀ 12th 6º. 35' up & back

8º to left.        8º down

9º 3' to right & back a little (straightening)   9. 3 far down & little to left (straightening rapidly)

10. 5 up & to right     10. 5 do.

11. 10 a little up.       11. 10' far do

11. 55 down    11' 55 down & to left (Straw may have caught)

1º do    1º considerably up.

2 a little to right & uppish      2º up

3 considerably up & to left     3' to left & little down

4 — down up & to left         4º down

5 down            5. 4 down & to right

6 to left & down         6 . much down & to R.

7. 15 little way to left             7. 15 20 far to right : great lateral circumnutation

9. 5 far to left & down

            10. 35 very far down: almost guess-work

13. 6º 40 far down due to slewing of fibre     13' 6º 40 up again

            8 10 down & to right

(Distance of shoot 6 inchs fr vertical glass)

[23v]

94 705

Chap. E 10

of insects having the power to carrying pollen to another and sometimes distant plant much more securely than can the wind.

In the above two classes taken together there are 38 anemophilous and 36 entomophilous genera; whereas in the great mass of hermaphrodite plants the proportion of one anemophilous to entomophilous species genera is very extremely small.

The cause of this remarkable difference may be attributed to anemophilous plants being having retained to a a primordial condition, in which the sexes were separated and 205 (fertilisation was effected by means of the wind, in a greater degree, than entomophilous plants. That the earliest and lowest plants had their sexes separated; as is still the case to a large extent with the more lowly organised forms, is the opinion of a high authority of Nägeli.* (*Entesthung und Begriff der Naturalist. Art. 1865. p. 22)

It is indeed difficult to avoid this conclusion, if we admit the extremely probable view that the conjugation of the Algæ and of many infusoria some of the simplest animals, is the first step towards sexual reproduction; and if we further bear


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Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

File last updated 4 February, 2026