RECORD: Darwin, C. R. [ny].10.21-.10.29. Marchantia / Draft of Forms of flowers / Proof sheet of Cross and self fertilisation, p. 134, published p. 265. CUL-DAR209.3.233-235. (John van Wyhe ed., 2002-. The Complete Work of Charles Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Transcribed by Christine Chua and John van Wyhe, edited by John van Wyhe 12.2022. RN2

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volume CUL-DAR209.3 contains materials for Darwin's book Movement in plants (1880).


233

(1

Oct 18th 1879. Marchantia— N-E window — slightly inclined frond with 2 Sights. — covered with Bell-glass [watched] in intervals [sketch]

vertical glass 

1      

8° 20' 9. 35' a.m. (?)

10. 30   down

11. 45   little down to the R.

2°  5    to R & up little

4 . 20'   about same spot

10 . 20'  ⨀  marks fallen down.

19th 7° 43 am    risen greatly apparently

19th 9. 15' new mark —(moved pot)

 

[diagram label]

mark close behind

 

    10 ·20   to right & down— slewed pot & new dot → ⨀

    12    5    to right— Horizontal

    12 . 53   down & to left

     2°    down little & to R.

     2 50'    to right.

     4°     little down

     7°. 18   down

    10'. 10' P.m   to right & little up

20th — 7°. a.m

(Epinasty?)   

       10 45    horizontally back to left

      almost under rim of Pot

10. 45   new dot pushed down peg without moving position of Frond

11. 45   down & to left

   1. 55   to Right Horizontal

   4° down & to Left

   7. 20   down.

    8. 55   down.

  10 30   to Right Horizontal

21st 7° a.m.

     9°   same spot

   10 20 down & to R

   12° down & to L.

     2° down & to R

     4 · 30 .    do .

7°20 P.m   down to Left.

10. 25   vertically down

22d   7° a.m.

⎼⎼   12°     down

⎼⎼   4. 20' down

⎼⎼   10. 45 Pot to Left Horizon

23d· 6· 50 am.

   9° 10   down

   12°   up   !!

   1· 30 atom down

   4°   down & parallel as at 9°·10'

   5 . 30   down & to left

    7 · 20   down

  10 · 30' P.m   do

[233v]

363

Ch VIII Concluding R

proportion of them than of other ordinary plants bury their capsules young ovaries in the ground;─ an action, which it may be presumed, serves for to protect them from being devoured by birds or other enemies though this gain. But this advantage is accompanied with by the loss of the power of wide dissemination. No less than eight of the genera in the list at the be beginning of this chapter include species which act in this manner, namely, several kinds of Viola, Oxalis, Vandellia, Linaria, Commelina, & at least three genera of Leguminosæ. The seeds also of Leersia, though not buried, are concealed in the most perfect manner within the sheaths of the leaves. The facilities

[The text of the draft corresponds to pp. 338-9.]

[bottom of page excised]

234

(2

24th Octobr. 1879      Marchontia

Plant kept damp   

under Bell-glass.  

7° a.m.

8· 15 down

8° 15· new dot ⨀ pushed peg deep.

     into ground & slewed pot a mere trifle

10. 20'  down & to R.

12°     —  up.

2°     down.

4     down & to Left

6  filament fallen off. (cd it have been falling for days?)

Oct 25 8° 45' am. — New glass — new filament well fixed. on old frond placed tracing — (New Tracing) nearly in same position to light

10. 15   risen greatly— This had best be the new start

11. 50'  down

2°   to   right & little up.

2°  50   back & to Left

4°   to   right & down

5. 5' to  L left & down

6. 5   to Right & down

7. 20·   straight up

9°.   up‒ higher than ever .

10· 45   still up same course

26th  7° a.m.

   8°· 25   to Right — Horizontal

   9°· 17    to Right     do

10 . 30    up & to left

12°   vertically down

1. 48'   to Right & little down

2.  50'   up.

4°    up & to left

 5°    back & parallel

  6     to left & down

  7° 10 down & to right on old line

   8 45  to left, little up & parallel

   10. 30' P.m a little back & down & parallel

27th — 6 45' a.m.

    7° 45' considerably up

Oct 19' 9° 15 a m   

to

24th 6° P.m.

[234v]

(a) text: lead on p. 338

The most irregular fact about the present species is that the long-styled plants produce long-styled cleistogamic flowers, & the mid-styled as well as the short-styled cleistogamic flowers of the corresponding forms; so that there are three kinds of cleistogamic & three kinds of perfect flowers produced by this one species! Most of the heterostyled species of Oxalis are more or less sterile, many absolutely so, when if illegitimately fertilised with their own-form pollen. It is therefore almost certain probable that the pollen of the cleistogamic flowers must have has been modified in power, so as to act on their own stigmas, for they yield an abundance of seeds.

We may perhaps account for the cleistogamic flowers consisting of the three forms, through the

235

 (3

Oct 27'   Marchontia

 

8°. a.m.   new tracing ↘   viewed more horizontally so not so much distorted & magnified

Diameter of Disc .4 transversely by .3 

Magnification — about 36 times   

Length of filament .7 of inch       

Distance from dot to glass   8.5 inches  

Distance from end of filament to dot   .25 inch

9°  –   down

10°    up & to Right

11° 10'   up & little to left

12°    down & to left—

1°      up.

2. 5    down

3°      up.

4°   down & good way to the Left

5°   to left

6°   to right back & parallel

7° 18·   to right same line crossed down line

8. 23   . to left & back

9. 30   . up & to right

10. 35 P.m. same course to right —(Filament dropped off. during night)

 

Oct 28th   10°. 34' a.m. fixed filament with [sketch]  to same frond, at right' angle to margin inclined upwards & made dot.— Pot moved so that abut 45° fr Light

      (New tracing on different method)

How much magnified?

assume that whole breadth of .3 inch moves

11° 10'   an atom up.

12   10'   an atom to left side

  1°  30'     nearly same spot.

  3°   same spot. about

  4°   risen a little

     moved very little not enough magnified

Oct 29'   8° a.m to test whether still in motion put black dot behind again

8° 45 · down

9° 25   down & to left

10· 30'  back & up— little

12   to left

1°   down

2°    little up

4°   in middle of same small arc.—

5°   35      do.    do

10. 30     do     do

Oct 30'   8° am nearly same general spot

(say when I examined again on 29th   

days subsequently— was still circumnutating

but over very small space)

11 inches

.33 dot to end of line

magnified 33 times    

Disc partially joined on to another one

[235v]

[Proof sheet of Cross and self fertilisation, p. 134, published p. 265.]


Return to homepage

Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

File last updated 20 December, 2025