RECORD: Darwin, C. R. [ny].10.21-.10.29. Marchantia / Draft of Forms of flowers / Proof sheet of Cross and self fertilisation, p. 134, published p. 265. CUL-DAR209.3.233-235. (John van Wyhe ed., 2002-. The Complete Work of Charles Darwin Online, http://darwin-online.org.uk/)
REVISION HISTORY: Transcribed by Christine Chua and John van Wyhe, edited by John van Wyhe 12.2022. RN2
NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volume CUL-DAR209.3 contains materials for Darwin's book Movement in plants (1880).
233
(1
Oct 18th 1879. Marchantia— N-E window — slightly inclined frond with 2 Sights. — covered with Bell-glass [watched] in intervals [sketch]
vertical glass
1
8º 20' 9. 35' a.m. (?)
10. 30 down
11. 45 little down to the R.
2º 5 to R & up little
4 . 20' about same spot
10 . 20' ⨀ marks fallen down.
19th 7º 43 am risen greatly apparently
19th 9. 15' new mark —(moved pot)
[diagram label]
mark close behind
10 ·20 to right & down— slewed pot & new dot → ⨀
12 5 to right— Horizontal
12 . 53 down & to left
2º down little & to R.
2 50' to right.
4º little down
7º. 18 down
10'. 10' P.m to right & little up
20th — 7º. a.m
(Epinasty?)
10 45 horizontally back to left
almost under rim of Pot
10. 45 new dot pushed down peg without moving position of Frond
11. 45 down & to left
1. 55 to Right Horizontal
4º down & to Left
7. 20 down.
8. 55 down.
10 30 to Right Horizontal
21st 7º a.m.
9º same spot
10 20 down & to R
12º down & to L.
2º down & to R
4 · 30 . do .
7º20 P.m down to Left.
10. 25 vertically down
22d 7º a.m.
⎼⎼ 12º down
⎼⎼ 4. 20' down
⎼⎼ 10. 45 Pot to Left Horizon
23d· 6· 50 am.
9º 10 down
12º up !!
1· 30 atom down
4º down & parallel as at 9º·10'
5 . 30 down & to left
7 · 20 down
10 · 30' P.m do[233v]
363
Ch VIII Concluding R
proportion of them than of other ordinary plants bury their capsules young ovaries in the ground;─ an action, which it may be presumed, serves for to protect them from being devoured by birds or other enemies though this gain. But this advantage is accompanied with by the loss of the power of wide dissemination. No less than eight of the genera in the list at the be beginning of this chapter include species which act in this manner, namely, several kinds of Viola, Oxalis, Vandellia, Linaria, Commelina, & at least three genera of Leguminosæ. The seeds also of Leersia, though not buried, are concealed in the most perfect manner within the sheaths of the leaves. The facilities
[The text of the draft corresponds to pp. 338-9.]
[bottom of page excised]
234
(2
24th Octobr. 1879 Marchontia
Plant kept damp
under Bell-glass.
7º a.m.
8· 15 down
8º 15· new dot ⨀ pushed peg deep.
into ground & slewed pot a mere trifle
10. 20' down & to R.
12º — up.
2º down.
4 down & to Left
6 filament fallen off. (cd it have been falling for days?)
Oct 25 8º 45' am. — New glass — new filament well fixed. on old frond placed tracing — (New Tracing) nearly in same position to light
10. 15 risen greatly— This had best be the new start
11. 50' down
2º to right & little up.
2º 50 back & to Left
4º to right & down
5. 5' to L left & down
6. 5 to Right & down
7. 20· straight up
9º. up‒ higher than ever .
10· 45 still up same course
26th 7º a.m.
8º· 25 to Right — Horizontal
9º· 17 to Right do
10 . 30 up & to left
12º vertically down
1. 48' to Right & little down
2. 50' up.
4º up & to left
5º back & parallel
6 to left & down
7º 10 down & to right on old line
8 45 to left, little up & parallel
10. 30' P.m a little back & down & parallel
27th — 6 45' a.m.
7º 45' considerably up
Oct 19' 9º 15 a m
to
24th 6º P.m.
[234v]
(a) text: lead on p. 338
The most irregular fact about the present species is that the long-styled plants produce long-styled cleistogamic flowers, & the mid-styled as well as the short-styled cleistogamic flowers of the corresponding forms; so that there are three kinds of cleistogamic & three kinds of perfect flowers produced by this one species! Most of the heterostyled species of Oxalis are more or less sterile, many absolutely so, when if illegitimately fertilised with their own-form pollen. It is therefore almost certain probable that the pollen of the cleistogamic flowers must have has been modified in power, so as to act on their own stigmas, for they yield an abundance of seeds.
We may perhaps account for the cleistogamic flowers consisting of the three forms, through the
235
(3
Oct 27' Marchontia
8º. a.m. new tracing ↘ viewed more horizontally so not so much distorted & magnified
Diameter of Disc .4 transversely by .3
Magnification — about 36 times
Length of filament .7 of inch
Distance from dot to glass 8.5 inches
Distance from end of filament to dot .25 inch
9º – down
10º up & to Right
11º 10' up & little to left
12º down & to left—
1º up.
2. 5 down
3º up.
4º down & good way to the Left
5º to left
6º to right back & parallel
7º 18· to right same line crossed down line
8. 23 . to left & back
9. 30 . up & to right
10. 35 P.m. same course to right —(Filament dropped off. during night)
Oct 28th 10º. 34' a.m. fixed filament with [sketch] to same frond, at right' angle to margin inclined upwards & made dot.— Pot moved so that about 45º fr Light
(New tracing on different method)
How much magnified?
assume that whole breadth of .3 inch moves
11º 10' an atom up.
12 10' an atom to left side
1º 30' nearly same spot.
3º same spot. about
4º risen a little
moved very little not enough magnified
Oct 29' 8º a.m to test whether still in motion put black dot behind again
8º 45 · down
9º 25 down & to left
10· 30' back & up— little
12 to left
1º down
2º little up
4º in middle of same small arc.—
5º 35 do. do
10. 30 do do
Oct 30' 8º am nearly same general spot
(say when I examined again on 29th
days subsequently— was still circumnutating
but over very small space)
11 inches
.33 dot to end of line
magnified 33 times
Disc partially joined on to another one
[235v]
[Proof sheet of Cross and self fertilisation, p. 134, published p. 265.]
Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)
File last updated 8 January, 2026