RECORD: Darwin, C. R. [ny].04.17-24. Saxifraga sarmentosa. CUL-DAR209.3.303-304. Edited by John van Wyhe (Darwin Online, http://darwin-online.org.uk/)
REVISION HISTORY: Transcribed by Christine Chua and edited by John van Wyhe 12.2022. RN1
NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volume CUL-DAR209.3 contains materials for Darwin's book Movement in plants (1880).
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Saxifraga sarmentosa
A pot with thin runners more than a foot in length hanging down on all sides perpendicularly. — Two tied up almost vertically, but their upper ends bent down so slightly after several days that I think curvature due chiefly to mere weight— One 17 1/2 inches long still tied up but with distal part depending yet considerably above horizontal had thin stand strand [sketch] fixed to tip & its movement traced on vertical glass; certainly circumnutate a little, rising & moving during day to one side & then back, then again back & then upwards; but on next day moved very slightly from growth seem arrested by straw fixed to tip.—
A second runner 10 1/4 inches long & thicker was traced at same time on vertical glass; see Tracing (I) it moved first up & to left then to right & up & then parallel back & down; up again; to right & then back & parallel. so changed course 5 times completely Next morning (18th) was off vertical glass, as it had risen so much & this show how little affected by geotropism & it movement now traced on Horizontal glass in darkness See Tracing (II.) Magnified 2 1/5 times = 2.2 — & the movement in line of arrow implies rising The tip made rather more than 2 ellipses between 7° 45' a.m. & 11° P.m, & the point tra the extreme actual distance travelled by point was almost above .75 of inch or 3/4 of inch. It is not likely that under Nature the runners
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The thick runner's growth was delayed in passing through the forest of pins, & this part, now slightly sinuous thicker than the older part.
(Stones on ground)
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Saxifraga umbrosa Sarmentosa
In a state of nature the stolons freely grow along the ground, so that can hang-down & probably travel laterally on earth in midst of vegetation & the circumnutation of point much aid it in finding path.—
(Ap. 24th I put a forest of rather large pins several scores always close together in front of tip of thick runner it penetrated amongst them, but could not get in, the tip then rose & the part beyond or rather within the forest become bowed; the tip then found a passage high up between the pins & again descended & at last got through. At one time it bent at right angles to its former course; ultimately the stem was permanently slightly sinuous.— Two very thin lateral runners got through another forest of pins much more easily‒ I think no doubt circumnutation must aid such penetration, though I do not say this cause of circumnutation.
By scratching lines of damp-sand it was clear that the runner circumnutated whilst going on it.
Smoked glass
The forest of Pins
Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)
File last updated 24 January, 2023