RECORD: Darwin, C. R. 1878.06.18-19. Oxalis valdiviana / Draft of Cross and self fertilisation. CUL-DAR209.4.320. (John van Wyhe ed., 2002-. The Complete Work of Charles Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Transcribed by Christine Chua and John van Wyhe, edited by John van Wyhe 9.2022. RN2

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volume CUL-DAR209.4 contains materials for Darwin's book Movement in plants (1880).

Draft of Cross and self fertilisation in the hand of Ebenezer Norman with corrections by Darwin.


[320]

Oxalis Valdiviana

8th

8° 15 am

9°  risen greatly

9 10'   changed glass filament same Cot.

10°   down

10. 30   down

11° & 11° 30' so low down as almost to touch sand not marked

12. 5 still down, but I think rising

12. 40   up

1. 37   up little

2. 42   down out of sight, but marked by chance

3. 5 far out of sigh.    do

4°    do

5. rising, but not yet in sight   Measure length of Cotyledons

6. 5 well up     & of rosea

9' greatly up, but still on glass     (Used)          

10. 35 

19' 6° 40' ⨀ Cot Horizontal

  8° 10 so far down as to be out of sight

Changed position of Pot— 8° 35 new dot, cannot actually see but near plane.

  9° 20 risen 

19th

  10 3' do      5° 45 same course

10: 35 up         7. 10 down & to right; △ prevents

11 very little up            leaf shutting— other seedlings asleep

11 27' fallen a little    7° 22 up & little to right long-ways

12 down — Cotyledon here horizontal          8, 7' high up— Hypo— stand point free

12. 43' far down, marked        8 46 still higher & to left

1. 30 further down        (may be hypocotyl which is

2. 12 up              moving

3. 5 do 20th 6° 35' ⨀

3. 30 up             8. 5

4. up & to right left (cause of line bending to left is that Hypocotyl be thus bent.)  

4° 30 same course      

5' a little up    

[320v]

[text excised]

Kerner has shown in his interesting essay,*(b) (*(b) Die Schultzmittel des Pollen, 1873) by the movements of the petals or of the whole flower during cold and wet weather. In order to compensate the loss of pollen in so many ways, the anthers produce a far larger amount than is necessary for the fertilisation of the same flower. I know this from my own experiments on Ipomœa, given in the introduction; and it is still more plainly 189 shown by the astonishingly small quantity produced by cleistogamic flowers, which

[Cross and self fertilisation, p. 376: "With many plants this latter evil is guarded against, as far as is possible, by the anthers opening only during dry weather,* —by the position and form of some or all of the petals,—by the presence of hairs, etc., and as Kerner has shown in his interesting essay,† by the movements of the petals or of the whole flower during cold and wet weather. In order to compensate the loss of pollen in so many ways, the anthers produce a far larger amount than is necessary for the fertilisation of the same flower. I know this from my own experiments on Ipomoea, given in the Introduction; and it is still more plainly shown by the astonishingly small quantity produced by cleistogene flowers, which lose none of their pollen, in comparison with that produced by the open flowers borne by the same plants; and yet this small quantity suffices for the fertilisation of all their numerous seeds."]


Return to homepage

Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

File last updated 19 December, 2025