RECORD: Darwin, C. R. Wheat (application of grease). CUL-DAR209.5.179-180. (John van Wyhe ed., 2002-. The Complete Work of Charles Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Prepared and edited by John van Wyhe. RN1

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volume CUL-DAR209.5 contains materials for Darwin's book Movement in plants (1880).


179

(14

Wheat Grease

Sieve 30° above, Temp Nov 28. 10 am 4.8C

Nov 27. 3 pm

Greased about 1mm from to 1.5 mm —1 to 1.5mm— on 3 roots from 1.5 to 2.5 long and growing vertically down. After 19 1/4 hrs 20 minutes 2/3 curved to wet they have grown so as to 5mm to 6mm long, one has remained vertical the rest other two have gone close to wet

19 (+) 24 (=) 43 

After 43 1/4 42 hrs from the beginning the one still vertical one has grown to 7.5 mm and moved 15° towards the wet.

Nov 28. 10. 20 am greased 1—1.5mm on 5 roots from 2 to 3 mm projecting except one which is 10 mm.

Before greasing the angles are:—

Before greasing          When examined after 22 3/4 hrs

1 — 20° (inside vertical         2 35 inside ie has moved 15° towards wet

2 — 10° outside          10 inside:— has moved 20 10° towards wet

(10 mm.) 3 — 10° inside        flat against sieve: it has moved 50° towards

4 — 2° 0° inside flat against:— has moved 40° towards

5 — 10 outside           flat against:— has moved 70° toward

This examination was made after 22 3/4 hrs when the roots were 5 & 6 mm long

Control

Nov 30

— None of the very numerous wheat roots which were not greased have failed to go flat against wet surface: ie all non-greased gone flat

The bending place is distant from tip not less than 4 mm or more than 6.5mm

Another trial gives— not less than 2.5 ........ 5 mm

⸻⸻⸻⸻⸻ 1.5 ...... 4.5 mm

Another one bent which had grown 10mm out from the sieve without bending did subsequently bend; so that the bending place may be as much as 10 mm from the tip.

180

15 (2

Wheat — Grease.

The sieve experiment was tried at the beginng of Oct when the temperature was 12° or 13°C in the daytime, & then I made a note to say that so many non-greased ones failed to reach go the towards the wet that no conclusions could be drawn. In the earlier experiments I kept the air of the room damper & gradually left it off from finding that the roots bent to the wet better & grew quite well in somewhat drier air, so that it is just possible it the drier air may have had something to do with the success at lower temperatures. tho' I very much doubt it.

[sketch]


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Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

File last updated 27 January, 2026