RECORD: Darwin, C. R. [1878]. Radicle of Bean. CUL-DAR209.5.6-7. (John van Wyhe ed., 2002-. The Complete Work of Charles Darwin Online, http://darwin-online.org.uk/)
REVISION HISTORY: Prepared and edited by John van Wyhe. RN1
NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volume CUL-DAR209.5 contains materials for Darwin's book Movement in plants (1880).
6
(A
Radicles of Bean with very small square about 1/20' of inch of stiff sand-paper almost like card & very thin paper almost like linen attached with shell-lac to opposite sides. (Cause of action must be slight resistance to growth for cannot be weight, & with still gum-water resistance very slight, & therefore I tried these experiments., to show sensitivity to difference.
Differential. 3 others
(62) Ap. 23d 8º a.m fixed bits of to straight radicle only just out of bean. 4º 30' P.m (i.e. 8 1/2 hour) decidedly turned from card 25th. 8º a.m. do, but only slightly. Used
(63) Ap. 23' 8º. a.m young radicle 4º 30' not turned from card & bit was had been parallel to longer axis of radicle, removed & refixed it. 24th 8º 15 bent from card, 4º. 30' straight 25th 8º a.m., curved in line of Sachs curvature— Doubtful case ✔
(64) card, fixed on 23d on morning of 24th.— at 4º 30' & 25' 8º a.m, tip bent upwards & not directly from thicker card, for the 2 met on under side & the bending was upwards from plane of junction (Used) ✔ [sketch] [sketch]
(65) Ap. 23'. 8º a.m young straight radicle 4º 30' not bent: 24th 8º a.m a little bent from card 4º 30' moderately bent from card. 25th 8º a.m. decidedly bent laterally from card.
(66) Ap 22d 8º a.m. 4º 40' just perceptibly bent from card: 23d 8º am. slightly but certainly curved from card 24 & 25' 25' 8º a.m. very decidedly bent from card
(67) Ap 22d 8º a.m. 4º 40' conspicuously bent from car thick card : 23d. 7º 45' am., very strongly curved curved from perpendicular about 45º.— 25' 8º a.m. do
(68) Ap. 23d. 8. a.m. fixed cards : 4º 30' not bent: 24' 8º am decidedly & considerably bent 25' 8º a.m greatly bent from thicker paper
7
(B
Thicker & thinner paper fixed to tips of Radicles
Beans
(69) Ap. 23d 8º a.m young radicle. 4º 30' not bent— 24th straight or not bent 24th 8º a.m. do. radicle point straight in line of gravity. The 2 cards or papers met at bottom thus & [sketch] on removing them, I fond that the thinner paper had far more shell lac forming quite a thick layer or cap.
(70) Ap 23d 8º am, very young radicle hardly out of bean, 4º 30' not, bent. 24th 8º a.m. 4º 30' slightly bent 25' 8º a.m moderately but decidedly bent from thicker card & from line of gravity
(Used)
(Nine specimens in these 2 pages.— thus doubtful case because card so fixed as to coincide with Sachs curvature .2d case where roots curved from line of junction of 2 cards & not more from thicker than thinner— 3d case no curvature for the thin paper was greatly thickened by shell lac.—)
(slip of paper pasted on:)
Differential action of Card & Paper.
(36)(37)(48) Ap 14' 8º 15' bits of stiff sand-paper & thin paper on opposite sides for differential action.
(48) Ap 14. 4º 30' P.m no effect.. 31— Ap 15th 8º 35 no effect
(36) — 4º 30 thin paper most effect (15'. 8º 40' very slightly curved from thick card) 16th 8º a.m. do) (17th do)
(37) — no effect (.15th. 8º 40' Curved from thicker card) 16th 8'. a.m. very decidedly bent from thicker card
— (17th. 8º am very considerably curved from thick card.)
Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)
File last updated 27 January, 2026