RECORD: Darwin, C. R. [1880]. Beans / Force of Radicle. CUL-DAR209.5.74-75. (John van Wyhe ed., 2002-. The Complete Work of Charles Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Prepared and edited by John van Wyhe. RN1

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volume CUL-DAR209.5 contains materials for Darwin's book Movement in plants (1880).


74

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Bean Force of Radicle

Penetrating wet tracing Paper

Stra Match-stick thus [sketch] with needle-Hole Marc 17th 7º P.m

March 18th 8º a.m split more open I think & had lifted up bean not sufficiently fastened down; fastened it better. Slipped out of Hole, twice over— too small a hole.

17th 7º P.m. tip placed on thinnest tin-foil.). 18' 8º a.m had glided over it— changed position of tin foil & put little water on it.

20' no power whatever of penetrating tin-foil (B) 

☞ First trials with Smoked glass

(18th 8º 10' 11.? Smoked glass almost flat: 20th afternoon radicle has ceased to grow. 21' 10º. a.m. removed glass— radicle has grown out almost horizontally & has left long straight track on glass.; with central clear space of nearly uniform breadth, & margin on each side with brownish film — The black edge festooned with laterl bays, the result of laterl nutation. In 2 place the tip has moved a little upwards. From breadth of clean track & breadth of apex it is clear that See Frank diagram & measurement of apex of root & the track on the glass

18th   9º a.m smaller bit of smoked glass inclined. at angle of 30º below horizon 19th 8º 30' am radicle curled round so that apex travelled after travelling down glass travelled up towards under side of bean, & black line left on radicle & then on opposite side. In only a few parts of the track is the glass absolutely clean, & hardly in any across whole space a yellowish film being left, showing with what little pressure radicle travelled over glass & here & there little bridges of black matter cross the track: of these irregular (a)

(This deserves closer examination— different widths, showing different degrees of 30th pressure. & indented each side unequally, but the little scallops on each side I doubt whether due to circumnutation.)

Kept dark Sticks all soaked in water

18th a bigger & thicker bit of wood with rather larger Hole March 19' 9º a.m certainly much cracked— on both sides of Hole — put red dot where crack ended. (March 20' 8º 30 the bean has forced itself up from beneath the wires, instead of penetrating hole deeply) March 21 9º a.m, the end of radicle has now grown through hole & penetrated earth & the split has travelled beyond the red spot: at first only the end beyond hole was split — March 24th 8a.m  has travelled beyond the red mark

18th thicker stick.

tied firmly →

—March 30th

March 19' 9º a.m. crack much open.— [illeg]

20' 8º. 30' crack much more open— radicle seems much swollen above stick

21' 10º I am sure crack more open — made scratch where it now ceases: 24th 8º a m crack has travelled further beyond scratch.

74v

(a) bridges I counted 4.; this can be accounted for only by circumnutation upwards— Margins of both sides very irregular, almost zig-zag— observe width of track    of inch. The radicle, as I saw, did not rest by apex on glass, but obliquely on side of apex

(B) March 20th. I find that stick A same thickness & pointed like end of radicle, with weight of 1/4 of oz beside card & thin stick dents the tin-foil & [sketch] weigh of 1 1/2 oz burst through it. So radicle when coming down oblique on surface exerts very little force, but when point of radicle secured in hole & growth continues will lift up bean though held down by crossed wire Frank finds by micromter that the tin-foil is between .02 & .03 m.m. or between .00079 & .0012 of an inch.—

75

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Bean  Force of Radicle

March 18th 11º 10' a.m. Bean in a Soap-Stand

3' of inch

Not split.

.3 of inc

8º. 30' P.m slipped out; made Hole larger with next-sized bradall.—

— 19th 9º a.m slipped out of Hole— bored hole larger with 2d bradall & put root in again the wood is a little cracked, but not the ends —(March 21 9º radicle wonderfully contorted— wood not split.— March 27th wood l .7 of inch long & very nearly .1 of inch square Diameter of Hole .05— Root growth enlarged above & below hole.— Wood appe not split to ends— 

Mach 19th 9º 30' a.m. Fixed end of radicle in Hole in split cylinder in brass Pincers March 21' 8º a.m. The apex after remaining in some time, somehow got out, Hole too small. Failure .7 long & very nearly .1 squ.

Mach 27th 9º a.m tin cap on long— radicle

75v

[diagram]

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Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

File last updated 27 January, 2026