RECORD: Darwin, C. R. [1880]. Beans / Force of Radicle. CUL-DAR209.5.74-75. (John van Wyhe ed., 2002-. The Complete Work of Charles Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Prepared and edited by John van Wyhe. RN1

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volume CUL-DAR209.5 contains materials for Darwin's book Movement in plants (1880).


74

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Bean Force of Radicle

Penetrating wet tracing Paper

Stra Match-stick thus [sketch] with needle-Hole Marc 17th 7° P.m

March 18th 8° a.m split more open I think & had lifted up bean not sufficiently fastened down; fastened it better. Slipped out of Hole, twice over— too small a hole.

17th 7° P.m. tip placed on thinnest tin-foil.). 18' 8° a.m had glided over it— changed position of tin foil & put little water on it.

20' no power whatever of penetrating tin-foil (B) 

☞ First trials with Smoked glass

(18th 8° 10' 11.? Smoked glass almost flat: 20th afternoon radicle has ceased to grow. 21' 10°. a.m. removed glass— radicle has grown out almost horizontally & has left long straight track on glass.; with central clear space of nearly uniform breadth, & margin on each side with brownish film — The black edge festooned with laterl bays, the result of laterl nutation. In 2 place the tip has moved a little upwards. From breadth of clean track & breadth of apex it is clear that See Frank diagram & measurement of apex of root & the track on the glass

18th   9° a.m smaller bit of smoked glass inclined. at angle of 30° below horizon 19th 8° 30' am radicle curled round so that apex travelled after travelling down glass travelled up towards under side of bean, & black line left on radicle & then on opposite side. In only a few parts of the track is the glass absolutely clean, & hardly in any across whole space a yellowish film being left, showing with what little pressure radicle travelled over glass & here & there little bridges of black matter cross the track: of these irregular (a)

(This deserves closer examination— different widths, showing different degrees of 30th pressure. & indented each side unequally, but the little scallops on each side I doubt whether due to circumnutation.)

Kept dark Sticks all soaked in water

18th a bigger & thicker bit of wood with rather larger Hole March 19' 9° a.m certainly much cracked— on both sides of Hole — put red dot where crack ended. (March 20' 8° 30 the bean has forced itself up from beneath the wires, instead of penetrating hole deeply) March 21 9° a.m, the end of radicle has now grown through hole & penetrated earth & the split has travelled beyond the red spot: at first only the end beyond hole was split — March 24th 8a.m  has travelled beyond the red mark

18th thicker stick.

tied firmly →

—March 30th

March 19' 9° a.m. crack much open.— [illeg]

20' 8°. 30' crack much more open— radicle seems much swollen above stick

21' 10° I am sure crack more open — made scratch where it now ceases: 24th 8° a m crack has travelled further beyond scratch.

74v

(a) bridges I counted 4.; this can be accounted for only by circumnutation upwards— Margins of both sides very irregular, almost zig-zag— observe width of track    of inch. The radicle, as I saw, did not rest by apex on glass, but obliquely on side of apex

(B) March 20th. I find that stick A same thickness & pointed like end of radicle, with weight of 1/4 of oz beside card & thin stick dents the tin-foil & [sketch] weigh of 1 1/2 oz burst through it. So radicle when coming down oblique on surface exerts very little force, but when point of radicle secured in hole & growth continues will lift up bean though held down by crossed wire Frank finds by micromter that the tin-foil is between .02 & .03 m.m. or between .00079 & .0012 of an inch.—

75

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Bean  Force of Radicle

March 18th 11° 10' a.m. Bean in a Soap-Stand

3' of inch

Not split.

.3 of inc

8°. 30' P.m slipped out; made Hole larger with next-sized bradall.—

— 19th 9° a.m slipped out of Hole— bored hole larger with 2d bradall & put root in again the wood is a little cracked, but not the ends —(March 21 9° radicle wonderfully contorted— wood not split.— March 27th wood l .7 of inch long & very nearly .1 of inch square Diameter of Hole .05— Root growth enlarged above & below hole.— Wood appe not split to ends— 

Mach 19th 9° 30' a.m. Fixed end of radicle in Hole in split cylinder in brass Pincers March 21' 8° a.m. The apex after remaining in some time, somehow got out, Hole too small. Failure .7 long & very nearly .1 squ.

Mach 27th 9° a.m tin cap on long— radicle

75v

[diagram]

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Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

File last updated 27 January, 2026