RECORD: Darwin, C. R. [1878]. Gossypium herbaceum. CUL-DAR209.5.99-104. (John van Wyhe ed., 2002-. The Complete Work of Charles Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Prepared and edited by John van Wyhe. RN1

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volume CUL-DAR209.5 contains materials for Darwin's book Movement in plants (1880).


99

(p1

Gossypium herbaceum (from Thompson)

Jun 14 5º P.m 13 radicles, some long, some short pinned to cork lid & kept on chimney piece.— examined 15' 8º 30' a.m

(15 4º 40' P.m Cork dropped off & fell into water & washed off all the squares!!! but what then took place I know not)

(1) broken

(2) 15. 8. 30 not affected. ⨀

(3) bowed from card large piece, but to Sachs   ?

(4) not affected   O

(5) (2 pins) bowed laterally from card ✔

(6) bowed. a little laterally (4º. 45' bowed a good deal laterally, but card washed off ✔

(7) not affected O

(8) card dropped off (15 9º am put on big card)

99v

 (9) not affected (15 9º put on big card but not well  O

(10) (Big Pin) bowed to card   O

(11) slightly bowed from card (4º 45 considerably bowed) ✔

(12) straight (15' 9º put on big card but not well)   O

(13) bowed from card, but to Sachs— 4º 45 greatly bowed ? (15' 9º a.m. put on big square up to 23º.)

 6/13 appear to have been slightly affected after 15º 30' 

Result

100

(2

Another jar small Cotton

(11) (marked with pin) June 29th 9º a.m R straight fixed sq 4º 40' bent from squ. but still straight July 1'. 8º r. horizontal, whether due to sqr I cannot say   ?

(12) 29' 9º a.m r. straight R— — fixed sq — 4º. 40' no effect) (30. 8.20' do looks unhealthy.) 0 July 1. dying

(13) 29' Big Pin 9º a.m R strait fixed sqr 4º 40' no effect — (30th. 8º 30 quite straight. O

(14) 29' Biggish pin 9º a.m R straight fixed sqr 4º 40 no effect. (30th 8º 20 straight. but sq has slipped off O

(15) 29º Biggish pn 4º 45 fixed sqr straight radicle short —(30th 8º 20 tip curved to sq. a little.   O

(16) 29' small pin. 4º 45 fixed sqr. radicle straight not short & short —(30' 8º 20 no effect O

(17) 29' small pin 4º 45 fixed squ — radicle straight — (30. 8º 20'' a little bent from sq. &; 30'' 4º 30' distinctly bent from sqr. (July 1' 8º a m .at right angle ✔

 July 1  st 8º a the whole lot yellow & dreadfully unhealthy I cannot tell why

100v

(2A

Cotton Herbaceous

(1) (28th 8º 15' sq fixed 5º P.m curved from sqr but slipped off refixed (29th. 8º 45 again slipped off & radicle look brown throw away) ?

(2) 28' 8º 15 sq. fixed— 5º Pm little curved from sq. (29th 8º  45' am— apex at right angles. but sqr loose 4º 45 sq dropped off  ✔

(3) 28' 8º 15 sqr fixed 5º P.m straight — (29th 8º 45 little curved from sqr— put fresh spe. & refixed fixed new sqr 29th 4º 30 no effect) (30th 8º 10' inclined from sqr. but in direction of Sachs & terminal portion straight.

(July 1' 7º 30' a.m The radicle is now quite Horizontal which must be attributed to the sq. on the lower side) ✔

(4) 28' 8º 15 sqr fixed 5º Pm straight (29th 8º 45 no effect, but then there is shell-lac all round)

(29' 9º put fresh spec. radicle straight 4º 30' no effect) (30th 8º 10' sq. dropped off, refixed one O

(July 1st 7º 40' a.m — decidedly bent to one side & to S. in accordance with position of sq.)   ✔

(5) 28' 5º 10' fixed sqr quite straight — 29th 8º 45 greatly curved from sq but chiefly toward S &  a little to one side  ?

(6) 28' 5º. 10 fixed sq quite straight but very short (29' 8º 45 quite straight but sqr on summit nearly at right angles to radicle, 4º 30 quite straight O

(8) 29th  9º a.m. r. straight. fixed sq 4º 30' no effect (30th 8º 15 sq. dropped off, refixed sq.) July 1. 7º 40' no effect x) but all lower part of r. brown O

(9) 29' 9º a.m r. straight fixed sqr 4º 30' no effect (30th 8º 15 perhaps a little curved from sqr). July 1' 7º 40' terminal portion decidedly bent from sq.) ✔

over

101

(3

Herbaceous Cotton— Radicles

June 30th 8º 40' a.m. fixed squ. of sand-paper & glass on 13 radicles all straight, excepting 1 (with free oblique pin by base) & this has basal part curved— day cooler temp. Temp on study Table 20º to 20 1/2 C.

(These were looked at 4º 30' P.m & only no 4' affected at all)

(1) (1 Big Pin) July 1 8º a.m. no effect   O

(2) 30' 4º 40' fixed bit of glass   July 1' 8º am. no effect yellow & unhealthy O

(3) July 1. 8º am squ dropped off unhealthy

(4) cinder. 30th 4º 35' a little curved from do ✔ slightly July 1' 8º cinder dropped off, straight

(5) (2 Big Pins) July 1' 8º a m no effect   O

 (6) July 1' 8º a m no effect  O

(7) July 1' 8' a.m no effect   O

(8) July 1 8º a.m no effect   O

over

101v

3A

Herbaceous Cotton

(9.) July 1' 8º a.m unhealthy no effect O

(10) (3 Big Pins) July 1. 8º a m no effect ? but as much curved from S. may have curled ?

(11) July 1' 5º am yellow unhealthy— much curved from S. & card ✔

(12) July 1. 8º a m sq dropped off straight   O

(13) July 1' 8º a.m no effect. O (July 1' 5º. '30 P.m tip now curved to sq.

Five Six of these were left viz (1) 10 (5), (6) (7) (10) & ( n 13) at 5º. 30' P.m. on July 1'. & I am nearly sure hung down more or less perpendicularly & now 5 of them are extended almost horizontally, being bent close  to the seed itself & not at the tip— they are rigid but do not look healthy— This is an extraordinary fact.— Have they been attracted to the damp cork, but if so why has not tip bent? July 3d 8º 30' a.m the 6th radicle has now turned up so as to be horizontal— The other are limpid — I have little doubt that this movement is precursor of death, but why it occurs I know not.—

102

(4

A   Cotton— July 5' 5º P.m. sq. fixed— in Kitchen

(July 6th 11º a.m. Kept on Study Table— roots do not look well) 4º 50' no effect roots look very bad.

(1) short st geotropic (July 6th 11º no effect. on any)

July 7' 7º 45' a .m no effect, all look unhealthy) Thrown away

(2) short straight geotrop

In Kitchen on July 5th general temp was from 23º to 26º C. & this seemed m too hot, but then heat was not applied below, & so radicles may have become too dry.)

(3) like above

(4) like above

(On July 6th on my study Table Temp. from 18 1/2 to 19 1/2, but the radicles did not recover.)

(5) straight inclined over seed

6) straight geotropic

(7) inclined little over seed & to right.

(over

103

(p 5

Cotton

(9) 7th 8º— no effect sqr. slipped — refixed. (5º no effect)

(10) 2 Big Pins 7th 8º curved to S. fixed sq (5º no effect

(11) 7' 8º lower part crooked fixed sqr (5º completely Hooked but sq. dropped off

(12) 7' 8º fixed sq (5º no effect)

(13) 7' 8º fixed sq (5º no effect)

(14) (Central) 7th 8º fixed sq (5º no effect)

July 7' 5º P.m the bases of all radicles yellowish-brown.

All the above radicles straight & geotropic except 2 or 3 somewhat inclined to S. & one a little crooked in basal part above alluded to

(July 8th 8º. 30' I can see no evidence of sqr. having acted.—)

103v

(p5

Jar B      Cotton   July 6th 11º 30' sqr fixed on; Kept on Study Table —(Temp July 6th during day 18 1/2 to 19 1/2. C.)

(1) 5º P.m no effect except, perhaps in one case. ☞ July 7th 8º a.m put on Sand Bath

(1) 7th 8º no effect 8º fixed sqr 5º P.m curved to S ?

(2) 7th 8º a.m a little curved from sqr (this is one noticed at 5º P.m last night) 5º P.m. much deflected but sq. dropped off ✔

(3) 7th 8º fixed sq (5º much curved to S. but sq. dropped off ?

(4) 7th 8º a. fixed sq (5º no effect   O

(5) (Big Pin) 7th 8º no effect (5º curved to S.   O

(6) 7th 8º no effect (5º do)   O

(7) 7' 8º no effect sq. slipped refixed sq (5º no effect.) O

(8) 7th 8º no effect (5º. do   O

(9)

  I had better try another lot on sand-bath with evaporation & not very hot

104

(p 6

July 7th Cotton

5º P.m fixed squ on 6 radicles which had gone in water all day & were nearly all fairly straight— & placed on warmest part of sand-Bath

July 8th 8º. 15 all 6 look yellowish & unhealthy — sq rubbed off several.— I can see no evidence of sq. having acted— 4 of them strongly inclined towards S.


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Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

File last updated 27 January, 2026