[120]

Oxalis corniculata. (Coppery var.)

Jun 12° 9° 20' P.m. when no doubt properly asleep.

1 seedling quite young with Cots at right angles

1 with Cots Horizontal!!

2 (with true leaves just beginning to from) with Cots at right angles, & this seemed general position

1 seedling (& other) with 2 true leaves small but well-formed with Cots horizontal—

So may conclude that Cots rise about 45° but do not go to sleep.—

Jun 13' 9° 30' a.m seedlings left floating in water all night cots now horizontal or even a little deflected downwards—

[121]

1878

Oxalis corniculata (coppery var)

& floribunda (cotyledons)

June 14th

Some seedlings come up yesterday & 2 today at noon had Cots. horizontal, & 4 other younger ones had them open at about rt. angles.

At 4° P.m the 2 had then closed at about rt angles — To night at 10° 30' P.m— all 6 seedlings have cots. at about rt. angles — so certainly do not shut at night whilst young.—

[in margin:] so must have risen 48°

15' 7° a.m. all the cotyledons are now horizontal, so it is certain that whilst young they rise about 45° above horizon at night.—

Jun 17th 10° 15' P.m I have looked again carefully with protractor & I cannot say more accurately than that on average the 2 Cots from together an angle of 90°.—

19th same state

[121v]

Jun 26'

Ox. floribunda little or no difference in length of petiole of young seedling & old with true leaves, yet we know Cots go on bowing at night. — This movement must be due to pulvinus. —

I have looked at O. floribunda, the pulvinus Pulvinus consists of small square cells (like those in O. corniculata) but placed close to blade, & apparently chiefly on the under side of petiole, where it joins the blade.— I have looked again it here forms a distinct & considerable protuberance. Are not more & smaller cells required to give it straight when it exudes water? The water goes into intracellular spaces, according to Pfeffer & if so it is mere assumption on my part that there is turgescence of cells, which subsequently favour growth

[121b]

(O. corniculata)

June 25th The upper part of petioles approach each other, but the blade also 11 days old bends upwards where joined on to top of petiole.

The petioles of older seedlings are longer than those of younger seedlings so there is growth.

Jun 25. Cots of seeding bearing 1 true & fully open leaf & a 2d leaf partly developed are as much raised as the Cots. of young seedlings ie about 45°.—

26' I have been looking again carefully at petioles of Cots & in middle between bases of Cots. & base of plumule there is narrow zone of longitudinally shortened cells like this.

Cots were at right angles & were flattened & pressed under covering glass, & these cells on upper lower side had longitudinally a shrunk & wrinkled appearance compared with those on upper side which seemed tense, as might be expected, as Cots were at right angles.

Are not the small cells confined to upper side of petiole?

[122]

Oct 4. 1878 [illeg] Oxalis corniculata

Young seedling with cotyledon only 75 long & 60 broad in divisions of micrometer, in length. There is no trace of pulvinus; but in another rather sudden change young seedlings with cot 97 long & 70 wide in length, the pulvinus was well developed. So it is in seedlings with a fully developed true leaf, which rose at night in usual manner about 45°. — The pulvinus consists of transverse zone of narrow (on long axis of petiole) cells, & they occur just at point where petiole bends.— In the older seedlings petiole increases in length chiefly above pulvinus.

(Used)

Do the young Cot. rise at night.—? For it will be very odd.

[123]

Oct. 1878.

Oxalis corniculata

Seedling born or opened Cots on 16th, on 11 23d had cots raised 33° — on 28th raised 30°, & now has little true leaf— Frank has described pulvinus.— (Used) 8 days old

Some seedlings born on 21s. had on 22d ie after 1 day Cots raised about 23° or 24° — à It is evident that Cots are raised slightly at first on Oct 28', raised 30°. Frank will describe pulvinus — Nov. 2d ie after 13 day some of them with Cots still raised 20° or 25°

Two seedlings born on 22d (Pins) on 28th had Cots about 40° — on Nov. 2d ie. after 12 days 10° P.m raised about 37°.

Nov. 4th 10° 15' P.m seems as much raised as yesterday. ie. after 14 days

N.B I had better say Cots raised about 35° to 40°, when fully elevated

[123v]

In older Cots still below Horizon

[123b]

November 3d

One of above seedlings which opened Cots. on Oct 21' on Nov. 3d ie after 14 days at 2° 10' P.m had each cots 19° below Horizon; when 10° 30' P.m the Cots stood 28° above Horizon, so had risen 47°.— 14 days old another seedling born on Oct 22d with cots at 2° 10' P.m 25° below horizon, at 10° 30' P.m, had then 38° above horizon, so had risen 25° + 38° = 63° — 13 days old—

(On Nov. 4th 10° P.M & 5th & 6th the cots seemed equally raised judging by eye.) ie. after 14 days

Nov 7' 10° 15 as much raised as before — good-sized leaves now rise from between Cots i.e. 17 days old

Nov. 8th 10° P.m do — 9th 10° P.m do. — 10th do

(11th 10° 15 do. 2 true leaves formed.) (12th do) (13th do)

(14th still somewhat raised) 15' 10° a.m. cots. horizontal 10° 15 considerably raised)

(16th Cots still somewhat raised at night) 18th 10° 30' P.m. raised) ie after 11 + 18 day or 29 days!!

I may say not actually measured after first fortnight