RECORD: Darwin, C. R. 1879.07.12. Cause of Sachs' curvature. CUL-DAR209.6.16-17. Edited by John van Wyhe (Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Transcribed by Christine Chua and edited by John van Wyhe 12.2022. RN1

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volume CUL-DAR209.6 contains materials for Darwin's book Movement in plants (1880).

Notes for Movement in plants. Text F1325.


[1]

July 12' 1879. "Cause of Sachs' curvature".— (see what Sachs says)

Phaseolus Scarlet run [sketch] Hilum A B

A Position of radicle of A. B sown in damp─ Peat in upright position with hilum on one side Sachs' curvature curvature causes radicles to grow close down to & covers the hilum. So it grows at first in damp air, but afterwards grows out like B.— now no doubt primordially the cotyledons were lifted out of the earth, & the part marked red red, between petioles & summit of radicle, wd from universal rule have grown arched for sake of breaking out of ground; & a retention retention of this movement causes Sachs curvature. It is due to epinasty, & has been carried on to epicotyl, when where growth of same upper surface causes it to keep arched for some time, subsequently from greater growth hyponasty on now concave surface straight the epicotyl, & similar growth on summit part between petiole & summit of radicle, (marked red) caused the bow bowing out in damp air figured at B. This cannot take place, or only slightly in the Earth.

(2

Phaseolus Scarlet Runner.

When seed germinates in this position with hilum downwards— Sachs' curvature is much more conspicuous at first, but left in damp air radicle subsequently diverges as at B. Everything goes on same principle, as in the last case.— [sketch] B A

Peas with hilum downwards ha has Sachs curvature & bending of Epicotyl on same principle.

Common Beans— everything exactly same principle— (see old Franks drawings

(After 2 or 3 days, whilst suspended over water, the radicles of the Beans straightened themselves, as at B in F. old pencil sketch but did not bow outwards like the radicles of Phaseolus.—)

There must be M.S of F. about Phaseolus


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Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

File last updated 16 January, 2023