RECORD: Darwin, C. R. & Francis Darwin. [1879?].09.28-10.15. Phaseolus Sachs Curvature. CUL-DAR209.6.18-23. (John van Wyhe ed., 2002-. The Complete Work of Charles Darwin Online, http://darwin-online.org.uk/)
REVISION HISTORY: Transcribed by Christine Chua and John van Wyhe, edited by John van Wyhe 12.2022. RN3
NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volume CUL-DAR209.6 contains materials for Darwin's book Movement in plants (1880).
[18]
Phaseolus Sachs Curvature
Six fixed damp air on study; chimney piece
Oct 13 12 noon, with hilum downwards [sketch]
They are fixed so that the tip of the root was vertical the basal part making an angle with the vertical θ, which I call inside vertical
Oct 13. 12 noon Oct 14 9.35am Oct 15 8.45am Oct 16 8.30am
[table not transcribed]
Thus the base of the root is at first under the bean as in making the angle θ; this angle afterwards diminishes so that the base becomes vertical or may even go beyond the vertical in the other direction
[19]
Jar D
Phaseolus Sachs Curvature
Oct 13. 12.10 noon
Nine fixed with the bean vertical so that the roots at least the basal part in nearly 7 out of 9 was not vertical but was were from 5° or 6° to 23° outside the vertical. One
[sketch]
23 [-] 5 [=] 18 [÷] 2 [=] 9 [+] 5 [=] 14°.
root had not grown & one is vertical.
Oct 14. 9.20 am after 12 [+] 9. 10 [=] 21.10 21hrs 10m
The angles outside the vertical are, 30° 35, 45 30, 38, 24. Two have not grown & one fell off its pin — The fo
Oct 15 8.34 am 36 + 8 = 44 after 44hrs 34m the angles outside are 70°, 50° 80° 80 50 25. The one which fell off on Oct 14 was put on at 28° outside and this angle is now increased to 45°— One fell off on 15th and one has hardly grown making up the nine beans
23 [-] 5 [=] 18 [÷] 2 [=] 9 [+] 5 [=] 14°.
Must have moved after 48h as much as 45° in opposite direction.)
[20]
Vert beans (of Sep 28) 1.5 mm cut off—
Oct 4. 8. 10 am
main root
1 20° Sachs— (ie 20 with vertical)
2 45 —"—
3 .0° (vertical)
This applies to beans cut off on Sept 28 at 7° 15 P.m
4 40 Sachs
5. 48
6.. 16
From this it appears that Sachs curvature not reduced, & the plumles are now straight.— yet judging by eye I thought that several had become straight — it is certain that radicles & plumules in several now lie in same straight line. —
The tips are now painted with new soot— cap— formed — ie after 6 days from Sept 28th
This applies to Beans cut off on Sept 28 at 7° 15 P.m
From this it appears that Sachs curvature not reduced, & the plumules are now straight.— yet judging by eye I thought that several had become straight — it is certain that radicle & plumule in several now lie in same straight line. —
The tips are now painted with new root- cap-formed — ie after 6 days from Sept 28th—
Conclusion
[21]
Jar D— 10 am Oct 4 Faba
The bean is fixed with its long axis vertical without considering the root which however are nearly vertical
|
min |
|
10 15 Oct 5 |
|
Oct 6. 10 30 am |
1— |
7 |
about V |
not grown |
1 |
not grown |
2. |
15 —, |
— Sachs — |
30 Sachs |
2 |
+32 Sachs |
3 |
11 |
—Sachs |
19° |
—33— |
base +25, tip 2 cm vertical |
4 |
10 |
,—Sachs — |
30° |
4 |
+37° & 20° Rt |
5 |
14 — |
Rt |
,—8°, |
5 |
'+17° |
6 — |
7 |
Left |
to left (far) |
6 |
+30° |
|
rough— |
|
|
7 |
— |
put on study chimney piece
Oct 5 7, 8 put in at 10 15 am length measured at 12 noon
7.— |
26mm |
—+10 Sachs |
— |
7, |
+ 13 |
8 |
15.5 |
—vert |
|
8, |
+ 18 |
9 |
26mm |
put in at 12 — |
+20° Sachs |
9 |
+55°, 40 Rt |
roots now beginning to dip in water
[22]
Jar D Oct. 8 9 am
1 = -10°
2 = +2 °
3 = +18° basal 2 1/2 cm) the apical 4.5cm is about vertical
4 = +25 °
5 + 22 18°
6 +3 6°
7 + 19 15°
8. + 18 15°
9 +5
measured in water
[23]
Jar D Oct. 9 8.35 am
1 = 10° away from Sachs
2 = 29 to Sachs
3 = vertical
4 = 16 Sachs
5 = 16
6 = 29
7 = 18
8 = 1
9 = 54°
Conclusion
Oct 9th Frank concludes from these observations on 9 Radicles not with tips cut off of Beans that Sachs' curvature does not disappear or straighten straighten itself. —
Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)
File last updated 28 December, 2025