RECORD: Darwin, C. R. & Francis Darwin. [1879?].09.28-10.15. Phaseolus Sachs Curvature. CUL-DAR209.6.18-23. (John van Wyhe ed., 2002-. The Complete Work of Charles Darwin Online, http://darwin-online.org.uk/)
REVISION HISTORY: Transcribed by Christine Chua and John van Wyhe, edited by John van Wyhe 12.2022. RN3
NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volume CUL-DAR209.6 contains materials for Darwin's book Movement in plants (1880).
[18]
Phaseolus Sachs Curvature
Six fixed damp air on study; chimney piece
Oct 13 12 noon, with hilum downwards [sketch]
They are fixed so that the tip of the root was vertical the basal part making an angle with the vertical θ, which I call inside vertical
Oct 13. 12 noon Oct 14 9.35am Oct 15 8.45am Oct 16 8.30am
[table not transcribed]
Thus the base of the root is at first under the bean as in making the angle θ; this angle afterwards diminishes so that the base becomes vertical or may even go beyond the vertical in the other direction
[19]
Jar D
Phaseolus Sachs Curvature
Oct 13. 12.10 noon
Nine fixed with the bean vertical so that the roots at least the basal part in nearly 7 out of 9 was not vertical but was were from 5º or 6º to 23º outside the vertical. One
[sketch]
23 [-] 5 [=] 18 [÷] 2 [=] 9 [+] 5 [=] 14º.
root had not grown & one is vertical.
Oct 14. 9.20 am after 12 [+] 9. 10 [=] 21.10 21hrs 10m
The angles outside the vertical are, 30º 35, 45 30, 38, 24. Two have not grown & one fell off its pin — The fo
Oct 15 8.34 am 36 + 8 = 44 after 44hrs 34m the angles outside are 70º, 50º 80º 80 50 25. The one which fell off on Oct 14 was put on at 28º outside and this angle is now increased to 45º— One fell off on 15th and one has hardly grown making up the nine beans
23 [-] 5 [=] 18 [÷] 2 [=] 9 [+] 5 [=] 14º.
Must have moved after 48h as much as 45º in opposite direction.)
[20]
Vert beans (of Sep 28) 1.5 mm cut off—
Oct 4. 8. 10 am
main root
1 20º Sachs— (ie 20 with vertical)
2 45 —"—
3 .0º (vertical)
This applies to beans cut off on Sept 28 at 7º 15 P.m
4 40 Sachs
5. 48
6.. 16
From this it appears that Sachs curvature not reduced, & the plumles are now straight.— yet judging by eye I thought that several had become straight — it is certain that radicles & plumules in several now lie in same straight line. —
The tips are now painted with new soot— cap— formed — ie after 6 days from Sept 28th
This applies to Beans cut off on Sept 28 at 7º 15 P.m
From this it appears that Sachs curvature not reduced, & the plumules are now straight.— yet judging by eye I thought that several had become straight — it is certain that radicle & plumule in several now lie in same straight line. —
The tips are now painted with new root- cap-formed — ie after 6 days from Sept 28th—
Conclusion
[21]
Jar D— 10 am Oct 4 Faba
The bean is fixed with its long axis vertical without considering the root which however are nearly vertical
|
min |
|
10 15 Oct 5 |
|
Oct 6. 10 30 am |
1— |
7 |
about V |
not grown |
1 |
not grown |
2. |
15 —, |
— Sachs — |
30 Sachs |
2 |
+32 Sachs |
3 |
11 |
—Sachs |
19º |
—33— |
base +25, tip 2 cm vertical |
4 |
10 |
,—Sachs — |
30º |
4 |
+37º & 20º Rt |
5 |
14 — |
Rt |
,—8º, |
5 |
'+17º |
6 — |
7 |
Left |
to left (far) |
6 |
+30º |
|
rough— |
|
|
7 |
— |
put on study chimney piece
Oct 5 7, 8 put in at 10 15 am length measured at 12 noon
7.— |
26mm |
—+10 Sachs |
— |
7, |
+ 13 |
8 |
15.5 |
—vert |
|
8, |
+ 18 |
9 |
26mm |
put in at 12 — |
+20º Sachs |
9 |
+55º, 40 Rt |
roots now beginning to dip in water
[22]
Jar D Oct. 8 9 am
1 = -10º
2 = +2 º
3 = +18º basal 2 1/2 cm) the apical 4.5cm is about vertical
4 = +25 º
5 + 22 18º
6 +3 6º
7 + 19 15º
8. + 18 15º
9 +5
measured in water
[23]
Jar D Oct. 9 8.35 am
1 = 10º away from Sachs
2 = 29 to Sachs
3 = vertical
4 = 16 Sachs
5 = 16
6 = 29
7 = 18
8 = 1
9 = 54º
Conclusion
Oct 9th Frank concludes from these observations on 9 Radicles not with tips cut off of Beans that Sachs' curvature does not disappear or straighten straighten itself. —
Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)
File last updated 28 December, 2025