RECORD: Darwin, C. R. [1849-1853]. Drafts of A monograph on the sub-class Cirripedia, etc. CUL-DAR210.7.[1]. Edited by John van Wyhe (Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Transcribed by Christine Chua and edited by John van Wyhe 8.2022. RN1

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin.

These draft fragments are in CUL-DAR210.7. They were later used as rough paper by the Darwin children. Eight cut-up pages have here been restored to form four draft pages of Living Cirripedia.

See also Darwin, C. R. [1854-1855]. Catalogue of the appendages and other parts of Cirripedes, mounted as microscopical slides. UMZC-Histories3.454.


[1]

(2

Pedunculated Sessile (1) Division, Pedunculated or sack-formed

(2) Lepadidæ

Capitulum usually

(2) Peduncle without calcareous scales, with attached by two

cement-glands & two undivided cement-tubes, opening out near centre of surface of attachment

(1) Capitulum with as generally flattened with calc. pieces sometimes absent or rudimentary, formed of with consisting of a carina sometimes rudimentary with two tergal valves & with Heptalasmis tt brachial valves often either absent or rudimentary

Calcareous pieces not overlapping each other, with the parts

Upper portion of calcareous pieces not projecting free.

valves only capable of being short & opened

[illeg] Sack, not furnishedwith voluntary muscles, excepting the

adductor of tergal valves

[illeg] lodged obliquely within capitulum – Stomach generally furnished with cœa

Branchiæ  attached to post-abdomen & abdomen.

no sack-branchiæ an ovum-fold in rostral portion?)

palpi flattened (but towards each other not touching

[line excised]

[2]

(3

[folio numbered:] 575

Genus Tetraclita

[illeg] [vesiculæ]. On along the occludent edge, [illeg] margin the inflected end extremity of each alternate growth-ridge is generally much thickened,─ a set of teeth being thus formed, by which the two valves are locked together, but In the T. porosa, this character is variable, for instead of sometimes every alternate ridge, & sometimes only every two or three ridges, distend separated from each other by several growth-ridges, are thus developed into teeth. The articular ridge is either moderately prominent or extremely prominent as in T. vesiculæ cœrulescens or [illeg];but the lower edge in no case depends as a freely free, hinge-like style as sometimes in Balanus. The adductor ridge is generally well developed & [illeg] distinct from the articular ridge: in T. purpurascens it is very blunt: in T. serata it joins is united to the articular ridge half way up it, thus forming a deep tubular cavity above the crests from the lateral depressor muscle running up to the apex of the valve: in T. vesiculæ cœrulescens, the adductor ridge is very short & is united to or almost continuous with the bottom lower end of the articular ridge, a small sub-cylindrical tubular cavity being thus formed. Small crests exist for the attachment of the rostral & lateral depressor muscles in all most of the species, excepting T. purpurascens & costata, in which, however, more especially in the former there are, instead of crests, [sketch] sma minute

[Living Cirripedia (1854): 322-3: "Along the occludent margin, the inflected extremity of each alternate growth-ridge is generally much thickened,—a set of teeth being thus formed, by which the two valves are locked together. In T. porosa, this character is variable, for sometimes every alternate ridge, and sometimes only two or three ridges, separated from each other by several growth-ridges, are thus developed into teeth. The articular ridge is either moderately prominent, or is extremely prominent, as in T. cœrulescens; but the lower edge in no case depends as a free, hinge-like style, as sometimes in Balanus. The adductor ridge is generally well developed and distinct from the articular ridge: in T. purpurascens it is very blunt: in T. serrata it is united to the articular ridge half way up it, thus forming a deep tubular cavity running up to the apex of the valve: in T. cœrulescens, the adductor ridge is very short, and is united to, or almost continuous with, the lower end of the articular ridge, a small sub-cylindrical tubular cavity being thus formed. Small crests exist for the attachment of the rostral and lateral depressor muscles, in most of the species, excepting T. purpurascens and costata, in which, however, more especially in the former, there are, instead of crests, minute pits for the attachment. These crests vary much in prominence in the same species."]

[3]

(4

(4) Clisiadæ

Shell unsymmatrical circular formed of six pieces, 4 opercular valves &  a Carina & rostrum the rt opercular valves capable of being  opened; the left attached & forming part of circle, with the carina & rostrum, which [illeg] on right side, do not overlap each other, but lock together  by a serrated suture but on left side do not approach each other. Base membranous.

Animal placed in a plane parallel to base.

Branchiæ none. Mouth upper lip not bullate, nor notched nor toothed. Mandibles with single row of teeth Inner maxillæ, cutting face irregular.

Cirri with three posterior  pair on same type.

[4]

or notched [excised]

formed. Nearly as large as [2 words illeg] inner maxillæ

without prominent [nostants].

Cirri 6 pair; with branches having two pair segments first pair differently constructed form the five posterior. 2nd pair with ant. arms also slight differently constructed

Tail absent or rudimentary & double, not articulated

Hermaphrodite or bisexual Penis single, seated beneath on abdominal face of anus.

Ovaria, unite within peduncle & sometimes round the sack extending under the opercular valves.

Larvæ in first stage uniorcular without prehensile antennæ

attached generally to floating or moving dead or alive objects (alepas?) as [illeg] Echnoidæ

[5]

(1 Section – Pedunculated or sack-formed

(II Arthrobalanidæ

Peduncle & Capitulum for conformed into a sack-formed bag. Valves represented by a toothed calcareous rim with two calceus bars running downwards towards the rostral

margin of sack.)

animal. Branchiæ none: abdomen post- abdomen narrow elongated & transversely articulated. Mouth with upper lip produced into a long flabelliform moveable organ. Palpi

cylindrical pointing upwards mandibles & inner maxillæ furnished with apodemes

Cirri with only 3 posterior pairs present; branches not articulated. Penis double

seated on dorsal surface anteriorly to anus.

[line excised] or represented by at single style beneath

[6]

Cirripedia

Sub-families Genera

Arthrobalanus

(1)

[remainder of page not transcribed]

[7]

(1) Chthamalidæ

Catophragmus 1

Octomeris 2

Chthamalus 4

Priolepas 5

Chamailasma 3

Thuralepas 6

Artrolepas 7

(2) Coronulinæ (?)

Coronula

Platylepas

Tubicinella

(3) Balanidæ

A flat or irregular

Balanus

[Diklutus]

Coria

Elminius

B. Base cup-shaped parasitic in or on corals & sponges

Acasta

Conopea

Adna

Prygoma

Creusia

Nobia

Desaria

35 genera

[8]

Sub-class

Orders Families

Order I.

Tripoda or Tribrachiata no has [illeg] feet. ie 3 pairs Arthrobalanidae

Order II

Saxpoda or Sax brachiate

Pedunculata

movements involuntary except adductor

Hexpoda no has

Tripoda 12 feet

Apoda

A Inequalia no movement [illeg] add Pomarisa from P. [illeg] a li

[9]

the anus.

Larvæ in first stage b[excised]

prehensile antennæ.

Living buried in shells.


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Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

File last updated 21 November, 2022