RECORD: Darwin, C. R. n.d. Milkwort — Polygala Diadelphia. CUL-DAR46.2.C47. Edited by John van Wyhe (Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Transcribed by Christine Chua and edited by John van Wyhe 8.2022. RN1

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volume CUL-DAR46.2 contains Notes for Natural selection.


[C47]

Milkwort — Polygala diadelphia notoctand & therefore not papilionaceous = wings standard & keel united into tube with kind of double shield over anthers.

[annotated sketch] standard, stigma, anthers, pistil, fringed wing or Keel, germen.

Stigma at rt. ∟ to style & separated from the true parcels of anthers by membranous prolongation of style. If nectar lies at foot of standard, bee's proboscis in entering (& partly by alighting on fringed wings or Keel) must scrape over anthers & pass directly in front of stigma, which lies against standard.—

Fumaria— Pistil & two bundle of stamens bend toward standard as in Pap. flowers & Keel (which consists of new cylinder) is open in front & allow organs to protrude, so that Bee would act, as in Pap. flowers. The bee, however, must claw the Keel as seems probable itself & not the wings, wh. have no tendency to open the Keel. (a)

[annotated sketch]

Keel

standard with nect.

wings united into one upright petal, like standard wh. two united & shield the upright keel.

[C47v]

1/2 Peloric Pelargonium both nectary unopen

+++ as the Bee visits either nectaries.— In the other Fumarias, although the arch is slit on both sides, pistil & stamens will readily escape only on the nectary side.—

(a) Smith calls standard upper lip & many united wings lower lip. (wh he says sometimes has nectary with honey like upper.— My Keel is formed of the alternate & similar lips united.— On gloomy day, watched long large beds of small white species saw nothing— perhaps rather late for many seed-pods formed. It is no argument Fumaria having nectary, for so has Heartease for Bees, coming to it— Watched another Purple species & saw no Bees = Shrewsbury saw there Humbles go to Fumaria glauca? a largish species ++— saw one two other on large fleshy red Fumaria.— (++ with mouth quite closed) In the red-fleshy Fumaria there are two nectaries; & it is beautiful to see that the alternate lips which forms the hood, (answering to the Keel of Pap. flowers) will readily slip off in front or back +++


Return to homepage

Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

File last updated 28 August, 2023