(C.

1881 [sketch] Clarkia pulchella elegans

Dark red anthers with red filaments exterior face sepals & more than twice as long, but not broader than the interior anthers (of pale pink with purplish filaments) which face petals & form the interior whorl. — In both sets of anthers plenty of pollen, which when water added seem equally good & grains swell in the same manner The pollen in the long stamen not red, exterior anthers are slightly more pink, & perhaps la perhaps a trifle larger (?) The pale interior anthers dehisce before the exterior red ones. The flower is strongly protandrous. Before the stigma opens the pale anthers have dehisced. At this stage pistil bent down rectangularly.

July 26 1 crossed by pale pollen of interior anther — (These abort in C. pulchella distinct plane White Thread

― 27' 1 crossed by purple pollen of exterior & longest stamen red anther do— Black Thread

― 1 White Thread.— 28 1 Black 1 White 29th 1 Black 1. White 30 2 white 1 Black

31st 1 Black 1 White 1 Black Aug 1. 1 Black 1 Double white 1 White 1 White 1 Black 1. Black

Aug 2d 3 Double White 2 Black 1 Double White 2 Black (over)

 

[82v]

July 26th I see there are 4 sepals & 4 petals & dark red face sepals & are exteriors

To day at 2˚ 30' P.m one stigma fully open. — several flowers fully expanded early yesterday morning

Pistil straighten itself & puts itself into gangway when stigma expands

The stigma sometimes touches red anther of exterior stamen & thus get self-fertilised.

Double white mean that red anther were previously castrated or as seem through leaves stigma quite clean, so there cd be no self-fertilisation. —

Clarkia pulchella only 4 fertile stamens & these face sepals; the 4 which which face petals are very minute quite rudimentary. In one anther alone I found a very few grains of pollen which did not swell in water — It is therefore clear that the (smaller) pale-pink anthers in C. elegans are tending to become rudimentary.─

[83]

Aug 7th I have been looking again at pollen of C. elegans & the dry grains of the dark red anthers are generally a little smaller than those of the pale red anthers, which are tending to be rudimentary

Aug 10' Plants in separate Pots 2 or 3 in each Pot

[data not transcribed]

[83v]

In the genus Enchiridium, which is close to Clarkia there are 4 stamens, so 4 quite aborted — In several other allied genera there are 8 stamens.

Clarkia plants in pots in Greenhouse