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May 30' 1862. Vanilla from Sion House (per Hooker) (Add at end of Arethusa & give case of Sobralia)

Long tubular green flower: Labellum attached to column close up to its summit; flower thus very tubular: in front of Labellum, some way beneath stigma - there projects largely an extraordinary stiff brush, hinged, like a series of combs with teeth pointing down, apparently to compel insects to back out near to the column, [sketch] as then succession of combs would act like a panel on the abdomen of insect

Another with broad filament which acts like spring & keeps it close on Rostellum.

Rostellum broad highly projecting, hinged, with edges curled in, (like that of Masdevallia); its inferior

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surface coated with viscid matter; when this is lifted up, in withdrawing a small hook, it lifts up the anther & viscid matter being scraped off the hooks removes pollen nicely. The Rostellum rests rectangularly on a [sketch] bilobed membrane, the back of which (ie surface towards anther) is viscid & two little horns (viscid? with one surface stigmatic tissue?? project on each side at the back of these membrane on stigmas - so that each stigma consists of a membrane & a horn; I have just looked at one of the membranes; it is covered with viscid globules & utriculi - The horns stand more in connection with wings of clinandrum, but the concave side seems stigmatic so I am not sure of nature. Certainly these horns project from each side transversely between Rostellum & the 2 stigmas. The large clinandrum, seems

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to allow anther & rostellum to work truly & perpendicularly these wings are furnished with large group of spiral vessels, close to which the horns project, as if produced & bent connective

Pollen extraordinary, single grains; no threads, but anterior point semi-waxy & abounds with pale brown viscid matter, even the posterior & more friable pollen has a little of this. Thus Vanilla is allied to Cypripedium & leaves no doubt that elastic threads are this matter modified. (I suspect Cephalanthera is one of Arethusa. Can the horns be inflexed "auricles" & wings do, extra developed?? Certainy arise near spiral vessels of wings of clinandrum.

Insect having pollen sticking to head, when retreating would lift up rostellum & daub the 2 stigma -

The Stigmatic surfaces are enclosed in a Box - closed on sides. Rostellum is the lid & stigma itself in front.