RECORD: Darwin, C. R. 1866-1871. Cabbage. CUL-DAR78.139-156. Edited by John van Wyhe (Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Transcribed by Christine Chua and edited by John van Wyhe 2.2023. RN1

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin.


(20

Cabbage, Cattell's Early Barnes.

1866

One plant under net, 10 flowers were fert. by other plant of same var. growing about 12 yards off in same soil & yielded 8 pods, (of which one contained only 3 seed & is omitted); & the 7 pods contained (max. 20 min. 10) on average 16.3 seed.

Ten flowers fert. by pollen of same flower & these yielded only 3 pods, containing, 2, 1, & 5 seed! As this plant was protected there was much pollen & L. [Lettington] placed on a quantity; whereas in above cross with other & uncovered individual, there was but little pollen, & yet great fertility — looks as if excess of pollen was injurious, as with Ipomæa.

14 self-fertilised pods were gathered from under net & these contained (max 10. minim 1) average of only 4.1 seed; & even if we take the 6 finest

(21

(Cabbage)

pods, average is only 7.1. These seeds self & spont. fert were fine & 58 in number & weighed 3.88 gr, whilst 58 seed fertilised by above cross with another individual weighed a little less viz 3.76 gr. —

(Seed Planted out 13th 3° 30

Crossed Plants with distinct Plant

Spont fert.

Crossed with own flower's pollen few seed

Oct. 15 2° P. m. 1 (Planted)

Oct 15 2° P. m 2 (Planted)

 

 

These lots germinated at about same rate; if any advantage crossed seed had it. I picked out many equal pairs equal & planted then in Pot I in greenhouse. I also picked up Oct 18 all the crossed plants which were most advanced & left the remainder in equal state to grow as well as they could in pure sand. —

Oct 26. The crossed seedlings, as yet only with cotyledons in glass of sand & pot, a little taller than the uncrossed. —

Nov 2d only a shade taller.─

[21v]

Cabbage 1866

[table not transcribed]

(21 A

Small Pot (no?) Cabbage

Oct 30' Transplanted later out of sand-glass, 4 plants of each kind into pot,— only cotyledons, nearly equally developed; but an uncrossed one, marked with zinc, decidedly smaller than fellow. —

(Nov. 10 Pot I. — Tallest crossed to base of Cots. 1.1. inch tallest uncrossed. 7 of inch — Leaves of former larger.

(Dec. 2' Pot I. crossed plants altogether look decidedly the finest, though rather most crowded). In pot 2 no difference)

(Dec. 30' Pot I both lots equal. Pot. 2. do, self-fert if any difference are finest)

(March 31' Pot I Crossed a shade finest plants)

(June 3d Pot. 2. was bisected about a month ago & the two halves separately planted in K. Garden; the crossed plant now look decidedly finest & have largest Leaves— In Pot I lately transplanted, without being at all disturbed ball turned out of Pot in K. G, the crossed look most healthy & finest. —)

(June 14th Pot 3 the crossed & self are each of nearly equal height; the tallest crossed is 8 inches high— tallest Self- 6 inches high. — The crossed look healthy

Pot II Tallest selfs equal tallest crossed; but crossed look finer plants. —

(see p. 21. B)

(21 B

Cabbages

I left two lots to grow & struggle in pure sand in tumbler in Hothouse. Now Nov 2d are much drawn up & though so having only cotyledons are 2 inches high to tips of leaf—; the crossed are on average a little taller than the uncrossed. (Dec. 1 Both lots very unhealthy, have no marked differences)

(Feb 10th Both lots in sand equal.)

(March 31 do do do & very poor state. dying.)

June 30' 1867 In Pot I. equal great difference as on 14th & there is now great difference in size in Pot II

1868 April. 26 In both cases in the crossed & self sets there is great difference in size of Plants. The crossed in both cases flowered sometime before the selfs. —

(May 8th 1868. The cabbages have now thrown up flower stems & having plenty of soil, there is no great difference, the tallest crossed 6 inches taller than tallest self). (One self plant put under net & crossed plants again crossed.)

(21 D

21 C in Paper

 

Purple Magenta & green lac. Cabbage, 7 flowers were crossed with pollen of foregoing white & green lac. cabbage, & produced 6 pods with average of 11.5 seed, with max. of 22 & min. of 5.

12 pods spont. self-fert. contained on average 15.7 seed with max of 26 & min 6.

Oct 19. 1868 measured to tips of leaves.

Magenta & green crossed by white & green.

[table not transcribed]

(May 15 1869— Of the five Pots with plants of the 2 crossed lots 7 of the crossed have produced flowering stems & are in full flower & only 3 of the self- Plants, out of whole lot—)

(May 23. 1869 I have now looked more carefully & find that with the Purple lot (crossed by gre white & green) 5 crossed plants ↘

[21Dv]

have flowered & only 2 selfs have flowered; & although the crossed plants were taller when young in Pot. I (in which alone there are flower stems on both sides) the flower stems of crossed is quarter of an inch shorter. I think the pure & hardy self-side— plants will ultimately beat the crossed plants crossed with the delicate var. (In Pot III. in which Cineraria had grown the self- plants yet have the advantage.)

May. 23. 1869. ―

In the lot from green & white crossed by purple the crossed side, as might be expected has an immense advantage over the self- side; in this lot 2 crossed & one miserable self-plant have flowered. —

21 E

Lancinated Cabbages. (same plants as in last page)

White & green crossed by magenta & green Aug 5' 1869 — measured to tips of leaves

Pot I crossed 30 inches pure green umbel var.

do 6 purplish

Pot I Self 10 & 8

Pot II. Crossed 28 marked with purple

do 9

Pot. II. self. 6 inches. (the other dead)

Magenta & green crossed by white & green seedlings

[table not transcribed]

N. B the appearance of the pure green var, has spoiled the experiment.)

The crossed plants were smothered by enormous pla curled leaves pure green var. somewhat

These tall self- plants are pure green var., with somewhat curled leaves

(12 E (21

Barnes Cabbages 1868

The self-fertilised of last or first generation was allowed under net to spont. self-fertilise themselves, & produced splendid pods some one containing 21 seed, & average about a dozen.

The crossed plants of 1st gen. were again crossed, by L. but produced from some unknown cause very poor pods, containing only 4.5 seed per pod, with max of 10.

The following plants will form 2d generation

Seed germinated on damp sand. —

(Oct. 20' 1868) Young Plants measured to tips of leaves.

June 6th 1869 do do

[table not transcribed]

(12 E (21

[Repeated]

21E

[calculations not transcribed]

(21F

Cabbage same plants as on last page.

August 5 1869 — Measured to tips of leaves or buds

[table not transcribed]

The plants were turned unbroken out of pot & injured by this &.: has not grown

N. B. the plant with great Heads had grown so crooked, that I cd not measure it anyhow

The 3 self smothered by crossed plant with great, immense Head. —

21E 21F

[calculations not transcribed]

(21G

Cattells' Barne's Cabbage — 1870

Same plants as on p. 21 F, after being turned out into open ground & are now just done flowering

(Heights of flowering stems to tip measured)

[Table, excised, not transcribed]

Hence the self plants seem as tall or taller than the crossed; but this chiefly owing to few stems running up. — In general vigour & size wonderful difference.

The eight crossed plants from Pots I, III & IV. cut off at ground weighed altogether 13lb. 11oz — whereas the 8 self- plants weighed only 5lb. 2oz. —

Table 28

21G

[calculations not transcribed]

(21H

Barnes' Cabbage 1870. —

The self-plants of last Page were allowed to set spontaneously

18 fl. on crossed self-plants crossed by distinct self plant (a) of same lot, white thread

18 flowers on self-plants crossed by pollen of plants of same var. fr grown in Cattells' nursery white wool. —

August 1870

But owing apparently to very hot & dry season, all the early flowers which were crossed set miserably in both cases. The spontaneously & later s self-fert. flowers were also rather sterile; but many pods were produced producing & 10 of these yielded only 3.7 seed per pod. —

Only 4/18 pods were produced by crossing the self- plants & these contained altogether only 7 seeds!

Only 4/18 pods were produced by crossing the self-plants & these contained altogether only 7 seeds!

Only 4/18 pods were produced by those crossed by Cattells' Plant & these contained 24 seed, ie average of 7.7 seed; but too few to judge of increased fertility. —

All 3 lots of Seeds germinated on sand & planted

The result on following page

[21Hv]

(a)

(The self-plants were intercrossed because I was afraid to leave crossed plant. (owing to Meligethes &c) under the same net with the selfs)

21I.

Barnes Cabbage— Jan 16— 1871

Young plants kept in greenhouse, raised from seed described in last page measured to tip of leaves

[table not transcribed]

21I

[calculations not transcribed]

21K

[text excised] Cabbage — same plants as on last page— The larger one has formed fine Heads— & I thought weighing would show difference best

[Table partly excised, not transcribed]

[21Kv]

[blank]

21K

[calculations not transcribed]


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Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

File last updated 29 May, 2023