RECORD: Darwin, C. R. [1866].10.28-1869. Marjoram. CUL-DAR78.186-189. Edited by John van Wyhe (Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Transcribed by Christine Chua and edited by John van Wyhe 2.2023. RN1

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin.


(28

Marjoram (cultivated Garden var.)

This plant so long cultivated by roots, is very sterile. Plants so incessantly visited by Bees must be crossed by distinct plants, but all in my garden no doubt from same original root-stock. — Plant produced only a few seed— Plants under net excessively sterile; from whole only a dozen or two seeds, of which with many small & bad. — Placed on sand Oct. 28th 5°. 30'

(Oct 30th 10° P.m. a few of Both lots have cracked seed-shells; certainly those under net as forwards or more forwards than those exposed & visited by insects.)

(Nov. 2d the crossed seed on sand have protruded rootlets apparently quicker than the uncrossed & certainly seem more vigorous.)

Pot I. transplanted & ∴ equal Kept in Hothouse Dec 30' moved into greenhouse

Pot 2. do ― ―  ― greenhouse; larger Pot.

(Dec. 1, Pot I no difference in plants hardly more than cotyledons developed.

Pot. 2 crossed rather finest, — In sand self-fert. finest, but these are very few & crossed are crowded.)

(Dec 13 Pot I. Hot H. uncrossed the rather the finest— ditto in Sand. Pot 2 Crossed finest).

(Dec 30' do do do do do)

(Feb 10th Pot 1 self-fert. finest. Pot. 2. crossed finest.)

(March 31. Pot 1 & 2 Crossed & self all nearly equal.)

/over

[28v]

(June 3d. 1867 Some little time, turned out of Pot, & put undisturbed in K. Garden. —

Pot I. self- tallest. Pot 2. crossed tallest.)

(It is probable that the parent plant had all spread from single root)

(28A

Marjoram — Dec. 19 1867

The plants described in last page seedlings possibly from crossed & self-fertilised seed, but almost certainly all derived from same original root-stock. These seedlings were crossed, (ie the whole head) & self-fertilised; but the latter produced more & finer, though extremely few seed, than the crossed heads. —Some of the crossed seedlings were allowed to be spont, crossed by Bees & were produced in much greater number. Seeds of all 3 kinds put on damp sand. Dec 19. 67. —

(Dec. 22d 67 Both sets germinated equally planted Dec 23d. afterwards on opposite sides of Pots. — The supposed spont. crossed marked with bits of zinc. — In Pot III. the crossed poor & germinated late; 2 of the spont. having germ. 18° after the selfs.)

(April 3d. 68 In all 3 Pots on average the crossed have decided advantage & are finer plants.)

(April. 23d very plain difference)

(July 9 (68) Two tallest plants on each side of each Pot. measured — 2 of Pots put out of doors Plant just going to flower. Pot I crossed flowered first

[table not transcribed] (July 15 crossed Plants in all 3 Pots flowered first.)

[table not transcribed] Aug 23d beginning to seed

In Pot I & II. (the former in greenhouse) are been kept in Pots & have very few stems on self-side, compared with crossed side— In Pot III.

[28Av]

the plants were turned out, & so had plenty of good soil to grow in, & they appear equally fine, with equal number of stems & almost equal in height as by measurements. Equal— number of plants originally planted, but now Aug. 26 /68/

Crossed

in Pot I 15 stems

Pot II.           15 —do

[total] 30

—self- side

6 stems

9. do

— 15 stems

Give case of Pot III. which was treated differently as separate case. —

Jun 24 69. I have compared seed, after above plants were left open & not fert.

Pot I crossed plants produced 6 flower heads, & selfs 5 Heads, but the latter produced quite as many or even more seed than the crossed plants. It is, however, scarcely possible to compare seeds accurately

Pot II. The crossed plants produced very many more flower heads & so more seed, but no difference in fertility of the heads in crossed & selfs.

Pot III. Flower heads & seeds apparently equal. —

(It is clear that these plant seedlings of 2d generation produce far more seed, though grown in same garden than the original stock long self-prop. by stolons roots.

28A

[calculations not transcribed]

(28B

Marjoram

May 9' 1869 — Pot I, which from having been long kept in greenhouse did not like being turned out there is wonderful difference in the size of clump, which has increased by stolons.

Crossed

Self

Pot I. plant 10 x 4 1/2 inch

3 1/2 x 2 1/2

tallest young stems   5 1/2 inches

— 4 inches high

Pot II.           18 x 9 inches

tallest stem    8 1/2 inches

12 x 4 1/2

— 6 inches high

Pot III. Both sides equal. —

(Aug 2d. 1869 In Pot I & II the inequality in the size of clumps remains about the same. — The crossed flowers in both flowered before the selfs—)


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Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

File last updated 29 May, 2023