RECORD: Darwin, C. R. 1866.04. Inheritance…short-styled cowslip-Polyanthus. CUL-DAR108.120. Edited by John van Wyhe (Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Transcribed by Christine Chua and edited by John van Wyhe 12.2022. RN1

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volumes CUL-DAR108-111 contain material for Darwin's book Forms of flowers (1877).


[120]

April 1866 Inheritance

In 1864 I fert. short-styled cowslip-Polyanthus with pollen from red equal-styled Cowslip (so that this is in fact or ought to be a Homomorphic union; as stamens are placed in red cowslip as in short-styled plants) and I raised 7 plants, which judging from stamens consist of 2 long-styled & 5 short-styled, but of latter 2 had longer pistils than they normally ought. —

I also crossed do. long-styled cowslip=—polyanthus with do. pollen & got 29 plants, of which 13 were long-styled, & 15 short-styled & 1 mid-styled — but one of short-sty had pistil above normal length & some of long-styled had it rather too short.—

[in margin:] Feeble transmission when crossed with strong inheritance especially if 2d generation comes true

à I do not think anything here worthy of notice or mention, except feeble transmission.—

[120v]

36 pl. were also raised from a common cowslip of both forms, fertilise crossed by the pollen of the equal-styled; & of these plants one was equal-styled, 20 were short-styled , but in 3 of these the pistil was rather too long , & the [illeg] 15 were long-styled. In this case we see a good illustration of the difference which often occurs, between single inheritance, which may be very strong, & proportion, the difference betw single inheritance & proportion of transmission

a fact which often occur, namely that a character may be finely transmitted by an uncrossed a variety. The

for the equal-styled structure wh self-fertilise, though not strictlyinherited is transmitted [illeg] with much force, but when cross with common cowslip fails almost completely before the its greater force of these of the common cowslip, when the two are crossed.


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Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

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