(3

Short-styled — white thread on 3 plants— Polyanthus pollen; heteromorph

Heteromorphic — cowslip-polyanthus seedings

All this page. Heteromorphic seedling — cowslip — Polyanthus mainly as standard for Homomorphic union.—

41 45 12? very poor 33 very fine seed 0

Long-styled

14 very fine seed

45 poor unripe seed

0 thread lost

8 flowers crossed

Longstyled by Pollen of Cowslip. white silk

43 immense seed

36 fine seed

39 fine, but not ripe

0 — lost shanked off

(4 fertilised)

Heteromorphic seedling Heteromorphically by other heteromorphic seedling of same lot White wool Short-styled

17

51

0

0 not ripe enough to be counted 4 fertilised

Long-styled

38

43 good seeds 45 fairish 23 very moderate 54 pretty good.} 5 fertilised

(average 38.7)

Short-styled by Heteromorphically by Homomorphic pollen no 6.

27 39 26 29 very fine 0

[calculations not transcribed]

Homomorphic (30.2 average)

average of all 15 other Heteromorphic cross crosses (excluding one with only 12) gives 37.8 seed per capsule

/over

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Aug. 16th 1863. I have compared all seeds on other sides. Those fertilised by cowslip are finest — the rest are nearly the same; perhaps those fertilised by Polyanthus not quite so fine. —

Those fertilised by Homomorphic pollen are as good as any others!

I do not understand how it comes that the naturally fertilised pods, which I believe were certainly fertilised before plants put under net are so poor — they were earliest flowers & plants fertilised flowered early in season.

Altogether I suspect that plants, perhaps from pots, not very fertile: compare with wild Cowslips. —

No great difference in seeds from cut-off short-styled (for crossing) short-styled & long-styled flowers. —

The pods crossed & Homomorphic cross pollen only a little rather less fertile

A plant which I believe had never been under the net produced only 2 pods with very poor seed. — I do not think that this plant was exposed to the tobacco fumes.— Are these plants sterile from being havng 1.2 Polyanthus? Does potting male them sterile? Whole case very odd.

(5

Final conclusion.—

I am utterly puzzled— Plants capriciously & oddly sterile: I am inclined to think owing to conditions. — But all evidence, as far as it goes, points to the homomorphic unions & even heteromorphic unions of being homomorphic p seedlings being excessively sterile

Aug 19th I have now compared all seed with wild cowslip seed. Of the Heteromorphic seed those fertilised by cowslip & by Polyanthus & one Heteromorphic with another are good average. N. B seed of wild cowslip varies greatly in size; when few they are often of very large size.

Of the Homomorphic seeds those fertilised by cowslip & polyanthus are fair average —

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Homomorphic Cowslip-Polyanthuses

Results of Experiments of 1863 & 64