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Conclusions. 1865. Long-styled Homomorphic purple Primrose from seed sent by Mr Scott from a Homomorphic union. —

I raised 23 plants, of these 18 were purple of different shades & 5 were yellow— Some One of the yellow was quite like wild plant; in the other the eye of corolla was brighter orange. Two or three of the purple were a little streaked & freckled with yellow; — an attempt at reversion. The corollas were generally larger than the wild & the plants were profuse flowerers— Two of the plants were had semi-monstrous flowers. — All were long-styled, with stamens seated low down, as in common long-styled. — The pollen resembled that of common wild long-styled, except that it contained many more shrivelled small & bad grains — The pistil in some was rather shorter than in wild; in many it stood up much higher above mouth of corolla; but this varied on the same plants.— The stigma itself varied in form & elongation.)—

(Ten plants, 8 purple & 2 yellow, were subjected to experiment, & there was no marked difference in fertility in these differently coloured seedlings. No less than 72 fl. on these several plants were homomorphically fertilised by pollen of same plant, & these yielded only eleven pods, (so proportionally very much fewer than with long-styled wild primrose homomorphically fertilised); & these pods contained on average only 11.5 seed each. instead of 52.2.

No pods were spontaneously produced by any of these plants another proof of very great sterility.—

[Repeat of 89b]

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Conclusion long-styled purple Primroses 1865

(To show that these 10 plants were sterile in some degree on both female & male side, 72 flowers on the several plants were homomorphically fert. by pollen of long-styled common wild Primrose, & these set 39 pods with average of 31.4 seed (instead of 52.2) & even if we omit 10 ten of the poorer pods, with less than 15 seed each, we can force up the average to 40.6.

(26 flowers were also fert. heterotopically by pollen of short-styled common primrose, & of these only 18 set pods giving 36.4 seed (instead of 52.2); but omitting 2 pods with less than 15 seed we get the average only up 41.2.

It is strange that thus pollen of wild short-styled is hardly more more effective in very slight degree than that of wild long-styled.— —i.e a heteromorphic than homomorphic union.

(To show that these ten long-styled purple & yellow plants were all sterile in their male organs, I fert. homomorphically 20 fl. of long-styled wild primrose with their pollen & got 14 pods, which yielded average of 15.4 seed per pod; & if 3 poor pods be omitted the average is 18.9. I also fert. heteromorphically 6 10 fl. of short-styled wild Primrose & got 6 7 5 pods which yielded average of 30.5 seed; or omitting one poor pod average of 35.4.

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Conclusions. long-styled purple Hom. Primroses

Hence we see that these Homomorphic seedlings are sterile to a considerable degree whether we test them by the action of their own pollen on their own stigmas, either by the number of pods which set or by the which gives average of seed as which is as 11.5 to 52.2.

Or by their spontaneous ave setting & average, which is as 0 to 12.7 seed.

Or by the action of pollen of pure wild long-styled & short-styled pollen on their stigmas, when we get proportion of 31.4 to 52.2 & of 36.4 to 66.9.

Or by the action of the pollen of their po seedlings on stigmas of wild long-styled & short-styled primroses, when we get proportion of 15.4 to 52.2 & of 30.5 to at least 65.0

Nevertheless, had we no other facts, it might have been argued that this demonstrated sterility in male & female organs of these purple as well as yellow seedlings was not due to their homomorphic nature, but to parent plant being accidentally struck with sterility & having transmitted this failure of power; something may be due to this, but having other facts, we may safely allow much to their homomorphic nature.

I was led to make these experiments in such detail from Mr Scotts statement about parent plant. He admit experiment not sufficient— anyhow in the 10 offspring of his plants, facts directly opposed to his conclusions.—