[A27]

Plantago laceolota — Hartfield April 28/63

Dichogam & Dimorph

In afternoon plucked off all anthers, next morning many stamens had grown to full length & shed large clouds of incoherent pollen — filaments long dangling — Corolla inconspicuous colourless, — neither attract insects or interfere with wind fertilisation — Pistil long secured with papillæ, adhesive — Poss

& Thyme

Development of flower from base to apex.— Pis

Stigma matures very much earlier that stamen & [illeg] from end of closed flowers — with anther extremely immature — I found stem stigma (in both forms) in this early condition with anther immature already penetrated by pollen-tubes — Hence Plantago is strongly female dichogamy — but a flower all near & filament so long, though beneath the stigmas, must often get own pollen; but by first flowers nearby be crossed by other plant. Moreover

[A27v]

"Female Dichogam. wind fertilised presenting a sub-dioicous dimorphite."

All heads on same plant, apparently belong always to same form, perfect or imperfect.—

[A28]

there in another form with generally longer pistil & with shorter filament, of which the anthers are brighter green when they first appear, & which contain but little pollen & the the grains chiefly of unequal sizes, with some few grains perhaps good, Like Thyme

Ap. 30 the green anthers of 2d form, dehisce; but contain hardly any pollen — nearly a few grains — I doubt about pistil longer —

All pistil on lower half of each head project far with tips bifid long before single stamen protruded — so m all these must be fertilised by distinct head or plant — When pis stamens protrude, the pistil is invariably withered & brown.—

[A28v]

May 1st I have examined a spec. of imperfect Form in which some few anthers contained a moderate quantity of pollen, but grains were of unequal size — other anther on same head contain very little or not no pollen — so amount of pollen variable on same plant.—

I have now compared pistils from 2 heads on "imperfect" & "perfect" flowers & pistil of same length; but I may say sometimes longer in imperfect flowers & stamens filament apparently always shorter in imperfect flowers.—