RECORD: Darwin, C. R. 1862.05.28-30. Viola canina. CUL-DAR111.A3-A5. Edited by John van Wyhe (Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Transcribed by Christine Chua and edited by John van Wyhe 1.2023. RN1

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volumes CUL-DAR108-111 contain material for Darwin's book Forms of flowers (1877).


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May 28th 1862 ─ Viola canina

Marked 2 plants with red left behind Azalea bed with no large pods, only small green flowers ─ some adjoining plants have such large pods, that I think must have come from early flowers.

The calyx of these large pods identical in size & structure with small flowers, which have only scales instead of petals, which became brown at tip & enclose stigma ─ One larger scale apparently represent lower lip.

Stamen foliaceous distinct. Each with 2 minute sacks which apparently dehisce at upper end ─ in younger state each sack contain extremely few pollen-grains─

There are sub-triangular with angle very

[1v]

Mark flowers next spring with thread & (measure pollen; examine stigma) came up one plant─ I observe many wild plants with extremely few seed;

Bees do not much visit these flowers

Mem. Ch. Fernand on Pansy.─

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much ended; in 3 spec. 12/7000 of inch in largest diameter ─ water produced no effect (perhaps not ripe); certain spherical brownish grains of unequal size, & ∴ do not look good.─

Ovarium with only few ovules.─

Stigma a hollow cup at end.─ bent at right angles.─ Calyx closed ─ no nectary.

May 30. V. canina.─ I found to day long pollen tubes protruding from small anther cells; the latter now empty & tips of tube broken; & in another specimen I traced these tubes at one end into anther cells of one stamen, & at other end into Stigmatic tissue.

Which causes the pollen-grains in middle of anther to protrude tube─ does stigma seat fluid which penetrate, I doubt /over

[2v]

May 28th V. odorata

(The white violet in K Garden which brown into large riped pods, has also very few small flowers like those of dog, but germen very hairy ─ 5 stamens 4 or 5 scales for petals.

Anther-ells similar: I saw add thread from each anther cell. (In double purple scented violet, what appear like imperfect flowers became very numerous scales are in fact double.─ Curious case of correlation

Ex can how then aborted flower may differ in different species from correlated clumps in old flowers.─)

do white violet K. Garden

May 30 ─ I find 5 scales alternating with sepals & with spherical vessels & 5 separate anthers; each with largish bundle of pollen tubes proceeding from open end of anther ─ as scale on top of anther extended some way beyond the 2 cells, the tubes have th run some way from opening opening of cell before reaching the stigma; for the flower was old & they could not have been drawn out after having penetrated stigma [sketch]

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May 30 Double purple violet odorata

I have examined with great care minute flowers of in thyme ─ I judge from time of year & single sweet white having three flowers & all others long out of flower & position from their position low down, like single little flowers of sweet white, that they are really like "small flowers" & former not opening. These flowers show section like a like section of red cabbage with central column, no trace of stamens or stigma (or ovarium?)) [sketch]

31' Viola canina in K. garden ─ end of stigma bent down on itself, much more than rectangularly bent bent ─ 4 petals excessively minute scales with long papillæ on edging; but the lower lip facing the bent stigma is 4 or 5 times bigger & has smooth edges.─ Two stamens alone produce pollen, & lie curled & with terminal scales crossing each other, with stigma almost within in the anther, filament footstalk narrower, & with scale-like hood; whole rather larger─: the 3 other stamens show no trace of anther-cells, separate, terminal scales flattened & not hood-like

[Continued at pp. 3, 4, 5, 6 of CUL-DAR111.A6-A11]


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Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

File last updated 4 May, 2023