RECORD: Darwin, C. R. 1878.09.14-16. Phalaris. CUL-DAR209.7.80-81. Edited by John van Wyhe (Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Transcribed by Christine Chua and edited by John van Wyhe 8.2022. RN1

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volumes CUL-DAR209.7-8 contain notes on heliotropism (phototropism) for Darwin's book Movement in plants (1880).


(1

Sept 14 1878.

Phalaris

Heliotropism with very obscure light

in Study: pot pot 42 inches from N.E. window:— with ordinary linen linen & two muslin blinds whole trial,— (Day after 9° 15 become cloudy, but clouds high & bright) from 10° 15' to 1° 32' a towel was also put over pot. — Sides of Box black; at back open to room. See 2 Tracings — Fig I made one great circumnutation, with longer axis in this case & in all others in direction of light & apparently thus caused, though so influenced by slight light; & then zig-zagged to light till 6° 36', by which time evening very darkish.

Fig 2. nearly like Fig (1)) in most respects

These ca 2 figures certainly indicate— circumnutation, modified & in latter part of day overcome by light & as evening advanced, line line becomes zigzag, or returned, showing renewed circumnutation. —

Sept. 15th— dark cloudy day, exposed pot behind window linen & muslin blinds & towel with roof to cover — sides black open to room behind, with roof above none of plants bent at all to light, yet pencil cast slight shadow. Early morning brighter, & they began by bending to light a little, then returned & circumnutated.

In the other compartment— plants exposed below window & muslin blind & 2 muslin blinds, these all become bent slightly to light, but the 2 Seedlings to which filaments hadbeen attached did not bend!!! yet circumnutated; Here we have another case of apex affected, preventing movement of base; I used however rather a large mass of shell-lac.

Pots 42 inches from N.E Study window

(2

Sept 16 1878 Phalaris Heliotropism obscure light

Study window: pot 42 inches from window — side of box blackened roof above, covering covering Horizontal glass on which tracing made was made: (also also a vertical glass in front.)— Window linen & muslin blind down all day — Also 2 muslin blinds over case. — At 4° 28' P.m removed window muslin blind, & at 5° 55' 1 of muslin blinds over case as evening grew darker so that till 4° 28' light came through linen & 3 muslin blinds— pencil cast faint shadow in morning.— Day in early morning sky bright & then become moderately cloudy with gleam of sunshine. [sketch]

Right-Hand seedings 1.9 high. — never moved off horizontal glass see Tracing A. At first whilst morning bright moved in slightly zig-zag line to light. From about 11° to 6° clearly attempted to circumnutate; afterwards did not advance more more to light, & returned during night; too old to return far. — Engrave only to 5° P.m or 5° 58' or both 6° 30'

Left Hand Seedling 1.6 high at first until 11° moved in very slightly zig-zag line with great great rapidity so as to cross whole Horizontal pane; & at 11° first dot made on vertical glass: afterwards the descending line shows bending bending to light in zig-zag line. — rose a little at night & late in evening, but had not power power to ascend ascend much during night — Not so good a tracing as the last — Do not give.

(over)

[81v]

This left Hand seedling had bit of short glass tube slipped over end to which filament was fixed & this weight probably prevented the seedling from rising at night


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Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

File last updated 25 September, 2022