RECORD: Darwin, C. R. 1873.06.30-.07.01. Drosera. CUL-DAR55.151-153. Edited by John van Wyhe (Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Transcribed by Christine Chua and edited by John van Wyhe 1.2023. RN1

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volumes CUL-DAR 54-61 contain material for Darwin's book Insectivorous plants (1875).


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Drosera

Jun 30 /73/ cut off bit of leaf, which had been in infusion of raw meat for about 2˚. 45'.— There was some distinct segregation in the cells near the glands; but only a few masses of purple protoplasm actually separated; but with small separate spheres of pale purple on sides & free. [sketch]

Lower down the tentacle the purple matter appeared as if a little separated from the wall, as if enclosed in a delicate envelope which had contracted — Perhaps this is a plausible theory; no for perfectly rounded end will send out a prolongation, which will flow yet I can hardly think that all the infinitely many small spheres & masses of protoplasm are bladders, as Huxley think, with purple fluid within— The current of protoplasm wonderfully conspicuous in this specimen; the many particles often flowed in close contact with the so-called bag of protoplasm, gliding along its surface, & not quite rather than close to wall of cell.

The bag was sometimes close to one wall & not close to the other wall or side — the little travelling masses masses of protoplasm

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was decidedly purple, for I saw them pass beneath the pointed end of the bag. So that the bag seems quite distinct.— Sometimes they passed beyond the pointed end & joined it. —

I cannot but think that a molecular change takes place in the homogeneous fluid content of cells, which contain invisibly minute atoms of purple coloured protoplasm & which unite together; but I fully admit, as H. showed me, that masses project from the flowing shell sheet & cross the cells; but the flowing sheet does not seem nearly thick enough to make all the masses after segregation & the flowing sheet is colourless. — It is rather striking that the currents seem to cease cease after very strong segregation from C. of Ammonia. or at least are far less plain: perhaps due to aggregation of all visible patches.

(I also examined at the same time & after similar exposure to C. of Am short tentacles of disc with greenish pedicels — the protoplasm is here greenish & no granules granules of chlorophyll can be distinguished — I saw current currents on wall; in the partially segregated cells. →

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cells, & colourless bag-like masses, of a greenish colour a little separated from the walls, as in the purple cells of longer tentacles — There were also greenish atoms travelling in the current.

The protoplasm being of different colors in 2 sets of tentacles looks as if extraneously coloured.

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July 1. pale leaf left in strong infusion of raw meat for 25˚ & then placed for about 1˚ in water — had become slightly flaccid with tent. well inflected. — Splendid circulation round cells.— great oval masses of purple protoplasmic circulating & apparently leaving leaving a tail tail behind them — Incessant changes of form— Much contraction of the so-called bag, leaving one end of cell almost empty, but filled with palest purple fluid, & here I could see circulation going on, so all protoplasm not drawn away from wall to form the bag— Some of these bags wd appear almost uniform in texture & rather dark purple; & then they these would gradually appear in midst endless spherical & oval oval masses, all changing their form. — This seems quite opposed to doctrine that it is a real bag. — A long prolongation & beaded projected from rounded end of bag & invaded end of cell which had before appeared almost empty.— Spheres appear within spheres. — perhaps these may be vacuities — filled with transparent fluid.

I again saw tails drawn out. I cannot believe bladders with purple fluid. —


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Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

File last updated 14 May, 2023