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G. ornata

The leaves of Genlisea are of two kinds described by Dr Warming as "spathulate" & "utriculiferous", but on account of the existence of true bladders on the rhizomes of G. filiformis it will be necessary to call his "utricules" by some other name. They are therefore called flasks & the leaves "flask-bearing" instead of "utriculiferous"

Each of These leaves have a proximal portion or petiole which supports the flask. The cavity of the flask communicates with the exterior, by means of the long tube or "neck", through the orifice. Two other tubes the "arms" communicate with the neck close to the orifice— The following are the measurements of a flask:— Neck 5/12 15/36 inch in length 1/100th in width; flask 1/36 inch in long diameter transverse Neck 1/100. In the figure the length of the arms & the direction of the spiral had to be taken from Dr Warmings fig because I couldn't make out either point from the torn off arms I worked at—

Two spiral or ascending vessels run up one in the centre the other in the ventral dorsal side

Spiral vessel

(The spiral vessel in the petiole bifurcates at a kind of ganglion below the flask, The two divisions vessels into which it branches run up, one on the dorsal the other on the ventral side of the neck & X each is continued on into one of the arms)— The arms have the following structure

If a flat ribboned like lamina were wound spirally round a cylinder in such a way that the edges of the consecutive turns of the lamina

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came into exact contact, then if the cylinder were removed we should have a tube, with a line, formed by the junction of the edges of the laminæ, winding spirally round it. If we suppose this spiral line of junction not to be soldered up, but the two edges which come into contact there to be pinched up all the way along. Then we shall have a tube round the outside of which a spiral projecting frill winds spirally just as the thread winds round a screw. The two laminæ This spiral frill being produced by the pinching up of two twines of edges where they come into contact along the spiral line of junction — must be formed of two layers one on the top of the other. These two layers can be easily separated & a needle pushed into the cavity of the arm; in fact in the specimens I examined they were found separated in many places, leaving natural entrances into the arm—

(At the distal extremity of the neck in the angle between the arms, is a transverse slit opening into the cavity of the neck)

The inside wall of the walls of the flask-cavity bears glands & spines

The glands are found at the bottom & also in two patches continuous with the first set & stretching a little way up the ventral & dorsal walls of the flask—. The glands are nearly circular in outline & consist of 4 cells (sometimes 8 Warming)— The spines are found where in the upper parts of the flask & begin to appear where the glands leave off.* According to Warming a few scattered glands are found in the upper part of the flask

The same kind of structure exists in the neck. Its inner surface bears at regular intervals slightly projecting rings formed of rectangular epidermic cells

Each ring bears a row of long thin transparent hairs; so that the whole inner surface of the neck is lined with successive zones of downward projecting hairs

*They are short hairs having rather broad bases & sharp lips formed by each by a separate conical cell. They are arranged side by side in rows, their inside free ends projecting downwards. So that the inside of the flask is covered with a series of rings one above the other each made up spines pointing downwards

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If the neck is split open & laid out flat the inner surface looks like a paper of pins, the hairs being the pins & the little folds of paper through which the pins are thrust representing the belt of epidermic cells which bears the hairs. The hairs swells out into a bulb at their attached ends & end in a sharp point separated from the rest of the hair by a transverse cell-wall. They vary slightly in length, on the average they reach down to the bases of the next row

The glands in the neck are of two kinds. Perhaps it has the [2 words illeg] I have. In its lower part they are spherical & 4 celled, with a side celllike those in the flask. In the upper part they are formed of only two elongated cells: in these the stalk-like cell on which they are supported is not symmetrically placed in the middle of the gland but is close to the its upper end. (The opening into the neck is symmetrically placed between the base of the arms & consists of a transverse slit which is continuous at the sides with the spiral line of junction running round either arm— It is not protected by a valve as far as I could make out & Warming describes nothing of the kind.

The lips of the opening are protected armed with a number of short thick hairs sharply pointed & with somewhat incurved. They also have their points divided off by a cross cell-wall. Similar hairs are found arming the edges of the laminæ which form the spiral frill. The long unsymmetrical glands are found in the arms I could not make out what the ends of the arms are like as I had only one broken specimens to work at

Spiral vessels

Prey

The rim is above the part not is say obscure

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Prey found in Neck

Flasks were full of organic debris & dirt

An annelid with retracted horny jaws; the abdomen of some articulate animal. The glands in the flask & neck are much discoloured as if by absorption. The hairs are flexible which looks as if they were absorbers— (The arms are evidently accessory traps.) I do not know whether the arms are closed at the ends or open. But in either case it is highly probable that the prey enters the arm through the crevices between the edges of the spiral laminæ; otherwise I can see no use in the short incurved spines which arm the margins of the laminæ just as they do the lips of the neck.—

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Genlisea filiformis (Bahia)

 No bottle-bladders could be found — On rhizomes of specimens originally sent from Kew on those I took off the line sheets sent, utric-bladders were found— They are transparent, 1/100th of inch length — The outline is peculiar, the dorsal surface margin being straight while the ventral is curved— The antennæ have bear some large hairs (or branches?) & are separate from each other down to base— The papillæ outside on outer surface are few & there are a few but very large quads having 4 equal cells [sketch] — No prey could be found in the bladders examined—

Genlisea africana. S. Africa

The same general structure as in G. ornata. A minute Acarus was found in one specimen— Where

(Are you sure no disruption of contents of Flask?)

Genlisea aurea* Fragments of neck were found with long hairs & unsymmetrical glands, showing that the general structure is the same—

No Rhizome-utricularia bladders could be found in any sp but Filiformis

*I cut these off from the sheet of dried spec