RECORD: Darwin, C. R. 1860.07.11-17. Columbine has circle of nectaries & 5 pistils with points slightly reflexed. CUL-DAR76.A7-A8. Edited by John van Wyhe (Darwin Online, http://darwin-online.org.uk/)

REVISION HISTORY: Transcribed by Christine Chua and edited by John van Wyhe 12.2022. RN1

NOTE: See record in the Darwin Online manuscript catalogue, enter its Identifier here. Reproduced with permission of the Syndics of Cambridge University Library and William Huxley Darwin. The volumes CUL-DAR 76-79 contain material for Darwin's book Cross and self fertilisation (1876).


[7]

July 11. 1860. Hartfield— Columbine has circle of nectaries & 5 pistils with points slightly reflexed from axis of flower. In Delphinium grandiflorum — the anthers at first all turned down: & covered & separated from mouth of nectary by 2 little petals, which a Bee separates in sucking — first about half anthers turn up & afterwards the otherhalf turn up & shed pollen.— Subsequently the 3 pistils turn up at right angles & the stigma then open, after nearly all own pollen-shed. At first the pistil are turned downwards so that whole direction is changed.—

(Delphinium)

I see lots of Flies routing in small flowers of Spiræa — Saw several times several flies on Stellaria graminea (onothersp. of Spiræa, Hive Bee

Hartfield July 1860

Euphorbia— pistil curls at right angles; Is not Euphorbia in very list of [3 words illeg] & curls at right angles; & come right into gangway— stigma is rough.— anthers coherent, furnished with spines projecting into gangway, & when touched by insect's back shake strike out large sized little coherent pollen-grains.— Each anther cell has a spine; lower cell of lower anther have much longest spines.=

July 17th saw many small Flies of 2 kinds sucking with long proboscis (over)

[7v]

these flowers; saw many repeatedly sucking —

Specimen preserved.— I watched through lens them crawling almost quite into flower — in doing this the 2 long spines interfered much & I saw then rub against them & shake lots of pollen on their backs.—

[8]

July 11th 1860. Hartfield. Ch. 3

Saw same little Fly in vast numbers on the Tormentilla officinalis — & legs-sides, eyes & antennæ were dusted with pollen— This rather insignificant flower must be largely crossed. — Flies suck base of pistils. Saw at least one small Bee collecting pollen.— Saw in Eliz garden 3 species of Saxifraga all visited by various small flies sucking base of pistils — several of these were quite coloured & dusted with pollen. —

One species, (Saxifraga sarmentosa from China & Japan) with 2 lower petals large & white — 3 upper smaller & prettily spotted: has 2 pistils in upright in young flowers, but bend slightly towards upper side when fully open: at base of pistil large orange rough secreting surface, which alone secrete & is visited by Flies & here pistil bends slightly upwards in proper direction

/over

[8v]

Hooker says in some Saxifrages & in Parnassii stamen more to pistil in pairs. In Kalmia they spring up, but being shorter than pistil they do not reach it yet jerkwould often throw some on —


Return to homepage

Citation: John van Wyhe, ed. 2002-. The Complete Work of Charles Darwin Online. (http://darwin-online.org.uk/)

File last updated 1 June, 2023