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CUL-DAR76.B86-B87
Note:
[Undated]
Dichogamy / Isotoma (a Lobeliad) white flow[ered] has curious long
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The Complete Work of Charles Darwin Online [86] Dichogamy Isotoma (a Lobeliad) white flow. Nov. 12. 1862. has curious long pointed filament from summit of united anthers, projecting rectangularly into gangway to nectary; when struck as must be by insect, pollen is shaken on to insect. After most pollen spilt, stigma protrudes open, as in Lobelia. — 5 plants in pots in Greenhouse, were not watched, but were not seen to be visited by bees produced 12 24 good pods 13 very poor pods (= 37 pods
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CUL-DAR76.B100
Note:
[Undated]
Tropaeolum tricolorum — Greenhouse / 12 fl[owers] crossed & produced 6
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The Complete Work of Charles Darwin Online (104 Tropæolum tricolorum — Greenhouse 12 fl crossed produced 6 fruit, ie. 1/2 set containing 9 seed. ie. 1.50 at average with max of 2. 18 fl self-fertilised produced 11 fruit ie 11/18 set, containing 19 seed, ie average 1.73 seed with max. of 3. Hence self-fertile in all ways more fertile. — [calculations not transcribed
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CUL-DAR66.59-77
Note:
[Undated]
[Bloom continued] [notes on plants cleaned with damp sponge; application
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Coppery Oxalis with yellow flowers. out of doors corniculata. (var coppery) June 1'. 9˚ 30'. F. fastened open 6 leaves cleaned 1 leaflet on each towards end where sticks are fastened. June 6. Only 3 of the old ones still attached fastened 7 more at 10. 15 AM — these are known by either being fastened directly to the bog main board or by a pin driven into the little laths. ( VI is pencilled opposite each new one — but this might wash out) The leaflet most away from greenhouse wall cleaned on
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CUL-DAR66.59-77
Note:
[Undated]
[Bloom continued] [notes on plants cleaned with damp sponge; application
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July 7th of the 4 leaves, 2 seem decidedly injured in comparison with others of same age. — not so 2 others — Bloom not renewed—, The bloom but I rubbed them again.) (Aug 19th— The 4 pairs of leaves cleaned on May 24 are all without exception much more decayed much more transparent than opposite ones which had not been touched.) Less chlorophyll in cleaned leaves for alcohol now deeply stained green in those which had not been cleaned Dec. 12 On a plant which stood in greenhouse had drops on
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CUL-DAR66.59-77
Note:
[Undated]
[Bloom continued] [notes on plants cleaned with damp sponge; application
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of Common Laurel have greasy appearance water does not well adhere to it; whatever it is, it is not easily rubbed off by dry, Handkerchief or S. Ether, but is perfectly removed by slight — rubbing with finest precipitated moistened chalk end of finger. The surface of Cassia common greenhouse is in exactly the same state as under surface of Laurel. — Youngish Holly-leaves polished considerably wetted. — Young leaves of Mahonia (aquifolia) splendidly polished — they after dipping are not
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self-fertile Cytisus scoparius. few comparatively to uncovered plants or none, or only a moderate number Ononis minutissima ─ pod perfect flower produce Plenty of capsules (8/12.) Bartonia aurea— Produced no capsules, in pots in greenhouse but then not healthy. — Passiflora gracilis highly fertile Apium petroselinum, apparently as fertile as uncovered plants Lactuca sativa — produced some seeds, season unfavourable orchis (Gesneria Adonis Hibiscus Pelargonium Phalaris Tropaeolum Clarkia Scabiosa
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CUL-DAR68.112-113
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[ny].07.13--[ny].07.23
Tropaeolum minus — dwarf var / 1h am / Rubbing with sponge at 90° does
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July 16. S. Ether removes silver, but surface still repellent waving nearly full-sized leaf for 1' in water at 92° no effect— same leaf in w at 101° for 1' silver largely removed on upper surface less removed on lower surface. — (July 20th at 9° 30' P.m To my great surprise the plants in 2 Pots in middle greenhouse were not asleep — none of leaves; whereas the plants in further House — the coldest in cool hot-house were asleep— Perhaps the plants in middle house were too dry; I have wetted one
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CUL-DAR68.70
Note:
[ny].07.17--[ny].07.20
Elymus condensatus / 8h 30 sponged space of leaf with [water at]
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The Complete Work of Charles Darwin Online [70] Elymus condensatus July 17th — 8° 30' sponged space of leaf with water 80-90 in greenhouse, removed bloom — placed on several drops of salt-water. (I see 4 sp in U. States are sea-shore plants) (19th trace of yellowish) July 20' 12° dead involuted. — Anyhow E. arenarius of Britain inhabits sandy sea-shores Hooker
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CUL-DAR68.140-151
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[ny].08.00--[ny].09.00
[experiments on effects of attempted removal of bloom]
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Carnation from greenhouse Young shoot. Aug 10 11° — one leaf cleaned on both sides, marked with blue on top, hung up to dry. — (Aug 29' no difference in the 2 sets) do— marked done in same manner to stand in water. Sept 17th no clear difference in the two sets of leaves [143
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CUL-DAR68.140-151
Note:
[ny].08.00--[ny].09.00
[experiments on effects of attempted removal of bloom]
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Greenhouse Sedum. Aug. 10' 11° Right side of several leaves cleaned both surfaces — old youngish— (The 2 leaves with black worsted where had drops on them not cleaned—) The cleaned leaves marked on tips with prussian blue. Hung up dry Aug 31. right-side of 1 old leaf rather more discoloured Aug 16' 9° 30 cleaned some leaves of branch blue dot put with cut-off end to stand in sol. of corrosive cultimate. [145
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CUL-DAR68.82-84
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[ny].07.31--[ny].08.23
Triticum repens in Greenhouse [application of fresh water, salt water]
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The Complete Work of Charles Darwin Online [82] Triticum repens in Greenhouse Aug 14th. 8° 35' put bunch of leaves in Bottle of fresh water. Aug 21st tips of blades yellow ― 23d nearly all the blades yellower. [83
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Pea Aug. 1'. I yesterday at night cut off tendril secured internode close to base of peduncle of leaf: I watched this peduncle from 9°. to 7° P.m., there was no true movement: there was depression towards light, as on other leaves; for plant had been all early morning in greenhouse. So I may conclude peduncle no movement, slight bending observed yesterday caused probably by weight of the 2 leave tendril.— [in margin:] Petiole does not move I watched all day from about 9° till 9° recorded on
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CUL-DAR68.82-84
Note:
[ny].07.31--[ny].08.23
Triticum repens in Greenhouse [application of fresh water, salt water]
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Triticum in Greenhouse Aug 8th 9° placed one bundle of blades, slightly sponged another bundle in bottle in salt-water, to see if they decay quicker in light. — Cleaned blades black wool: — white uncleaned. — Aug 12th 10° 30' — 3 leaves white wool with tips badly injured yellow. — Black wool 2 leaves with tips yellowish. (warmth light seem to make great difference in decay. Aug 13th leaves considerably injured, especially tips where bent — no difference between cleaned uncleaned. — [84
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CUL-DAR68.115
Note:
[ny].07.16--[ny].08.29
Carnation — Greenhouse / 11h 30 — Removed bloom with sponge & water at
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The Complete Work of Charles Darwin Online [115] Carnation — Greenhouse July 16th 11° 30' — Removed bloom with sponge water at 90° from whole of upper surface of 4 leaves. The uppermost has axis (which is bloom-covered) also cleaned water shall be lodged in the axis angle between leaf axis. — (19th water often renewed but frequently rolls off) (July 28th left off putting on water: no effect, except leaves a little spotted with dusky tint, as are very many other leaves which had not had their
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CUL-DAR157.2.65-67
Note:
[ny].07.01--[ny].08.16
Virginian creeper [Ampelopsis] / 3h 30 pm set stick to base of upper fork
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immersion a little warmed of some hours duration, except such as are deeply embedded these are only mechanically held. (I tried all this repeatedly) Essential oils dissolve caoutchouc. — The difference was marked compared with anything else tried. — Aug 10th On Brick in Greenhouse after 38° touching, end of tendril showed plain signs of swelling. [4v
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CUL-DAR68.67-69
Note:
[ny].07.12--[ny].08.01
Marine Grass — Salt-water / 9h 1 leaf cleaned with salt water
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Triticum repens. var.? Marine grass — Greenhouse July 13th 8° 50' after cleaning blade with sponge water at 85-90°. — put on salt-water on 2 with black wool 1 with white wool distilled water— on 15th in at 4° P.m 12° not shut evening forgot put on salt-water afterwards continued salt-water The leaf was then quite green (15th none of these leaves shut) (July 16th — 8° A.m The 2 leaves with black wool, now look yellowish-green. —) (All the killed leaves become closely involuted.) (July 18th 8
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A793.3
Beagle Library:
Kirby, William and Spence, William. 1815-26. An introduction to entomology. 4 vols. London: Longman, Rees, Orme, Brown, and Green. vol. 3.
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becomes a covering for her eggs, which it incloses on every side. To make this intelligible to you, further explanation is necessary. You must have noticed those singular immovable tortoise-shaped insects, which are such pests to myrtles and other greenhouse plants. These are the young of a species of Coccus (C. Hesperidum L.), and their history is that of the whole race. Part of them never become much bigger than the size of which you ordinarily see them, and when full-grown disclose minute two
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Sunday, 23 January 1887 Monday, 24 January 1887 Alice Hutchinson came Tuesday, 25 January 1887 went to Greenhouse !! 48º Wednesday, 26 January 1887 !! Thursday, 27 January 1887 Friday, 28 January 1887 Saturday, 29 January 1887 Alice H. went January – February 1887 Sunday, 30 January 1887 out twice Monday, 31 January 1887 r dead Tuesday, 1 February 1887 Wednesday, 2 February 1887 frost brisk Thursday, 3 February 1887 52º v. unwell all day Friday, 4 February 1887 53º out in chair Saturday, 5 February
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A585
Book:
Lindley, John. 1840. The theory of horticulture; or, an attempt to explain the principal operations of gardening upon physiological principles. London: Longman.
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with a substance whose moisture is continually diminishing; or in a greenhouse, where the pots are syringed, the heat of the earth in contact with the roots is lowered by a copious evaporation from the sides of the pot, just when, in nature, the bottom heat should be the greatest. The evil consequences of this are well known to gardeners, who however seldom take any sufficient precautions to prevent it. Greenhouse plants exposed to the open air in summer always suffer severely from the irregular
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A585
Book:
Lindley, John. 1840. The theory of horticulture; or, an attempt to explain the principal operations of gardening upon physiological principles. London: Longman.
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depends upon their peculiar habits, and the temperature of the climate of which they are native. So far as general rules can be given upon such a subject, it may be stated that the temperature of the earth most favourable for germination is 50° to 55° for the seeds of cold countries, 60° to 65° for those of greenhouse plants, and 70° to 80° for those of the torrid zone. No [page] OF SEED-SOWING. 16
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A585
Book:
Lindley, John. 1840. The theory of horticulture; or, an attempt to explain the principal operations of gardening upon physiological principles. London: Longman.
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first and greatest end attained by potting is, the power of moving plants about from place to place without injury; greenhouse plants from the open air to the house, and vice versa; hardy species, difficult to transplant, to their final stations in the open ground without disturbing their roots; annuals raised in heat to the open borders; and so on: and, when this power of moving plants is wanted, pots afford the only means of doing so. It also cramps the roots, diminishes the tendency to form
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A585
Book:
Lindley, John. 1840. The theory of horticulture; or, an attempt to explain the principal operations of gardening upon physiological principles. London: Longman.
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must have all such roots cut off again; for there are no means of bringing them within the limits of a pot. For these and similar reasons, no good gardener will expose his greenhouse plants to the open air in summer, if he can help it; unless they are duplicates, or unless there is some object to be attained very different from the strange notion that they are hardened by this process. The effect that is really produced upon them is, to give them a sort of artificial winter in summer, that is
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A585
Book:
Lindley, John. 1840. The theory of horticulture; or, an attempt to explain the principal operations of gardening upon physiological principles. London: Longman.
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Excretory functions of roots, 284. Exogens, 29. 30. their pith central and distinct, 25. Exposure of greenhouse plants, 288. Eyes, propagation by, 191. F. Fall of the leaf, 54. Fertilisation, 68. of the Mistletoe, 332. Fibrous roots, production of, by root-pruning, 308. Fibre, woody, originates in the leaves, 24. Fibres of vegetables, 138. Field crops, germination of, in wet seasons, 182. Fig-trees, mode of pruning, 253. Filament, 56. Floral envelopes, 56. Flowers, their action, 55. their
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A585
Book:
Lindley, John. 1840. The theory of horticulture; or, an attempt to explain the principal operations of gardening upon physiological principles. London: Longman.
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. deceptions practised with regard to it, 233. on roots, 235. herbaceous plants, 247. pines, 248. Grafting clay, its use in preventing evaporation and affording aqueous food to the scion, 246. Grafting plasters, their unfitness to supply aqueous food to scions, 246. Grass, its fibrous texture favourable to the emission of heat, 139. Granulation, 297. Granulations of cellular tissue, 245. formed by the living tissue, 308. Grapes, a cause of their shrivelling, 110. Greenhouse plants, their exposure, 288
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to Caerdeon. In the morning we all drove up to the poor old Mount had the misfortune to find the then tenant, an underbred clergyman at home. He never left us alone for one moment I admired my Father's gentleness forbearance in the way he answered all his prosy impertinent questions as to the changes made since the old time. Afterwards he said as we were driving away with a pathetic look of regret, If I could have been left alone in that greenhouse for five minutes I know I should have been
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A222
Periodical contribution:
Groom, Henry. 1844. Pelargoniums. Gardeners' Chronicle and Agricultural Gazette no. 36 (7 September): 605.
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feathered race except that bold, impudent, sooty, but withal comical fellow the house-sparrow. My small garden is some half mile off in the suburbs, and I do all in my power to induce the birds to frequent it, through, as an incessant warfare is kept up against them by every idle apprentice who can lift a gun, they are far from numerous. I formed this garden in the winter of 1836, and immediately a robin took up his quarters with me in the greenhouse, going out during the day, and returning at night
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CUL-DAR28.1.A1-A55
Draft:
[1848--1882.04.00]
The action of carbonate of ammonia on the roots of certain plants
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(With the following plants the solution produced some slight effect. with The roots of a fern, Nephrodium molle, were immersed for 20 hrs in a solution of 4 to 1000; this caused the deposition of some brown granular matter in the cells near their tips of the roots; and more or less confluent globules could be seen in the underlying parenchyma-cells. So it was with an unnamed greenhouse species of fern; and in this case the almost loose cells of the root-cap contained brown granules. The roots
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CUL-DAR28.1.A1-A55
Draft:
[1848--1882.04.00]
The action of carbonate of ammonia on the roots of certain plants
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exterior layer likewise contained a extremely fine granular matter. Still higher up the same root there were considerable spaces in which none of the cells contained granules. But again higher up these granules reappeared. The root-hairs were numerous; but I did not one was seen which arose from a cell containing granules.) (Roots of Oxalis sepium, corniculata, and of a greenhouse sp. with small yellow flowers, were immersed in a solution of 7 to 1000, and granular 3
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The Complete Work of Charles Darwin Online [221] Imatophyllum (A) the pot see fig stood in cool greenhouse 16 1/2 inch from BC which represents the glass partition between the green house grape house. A straight bit of glass 6 cm (2.4 inches) was fastened onto leaf observed by foreshortening, dots made on the vertical glass. Observed for 3 days Oct 8, 9, 10; everyday it f sank during the morning rose during afternoon morning evening. The angle O was about 50 O only altered when the leaf at its
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CUL-DAR116.1-43
Abstract:
[1854--1855]
Gaertner C.F von `Versuche und Beobachtungen über die Bastarderzeugung im Pflanzenreich' [Stuttgart 1849]
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or no (K) that hardly this shakes me.— It is very evident From G Koelreuter, Herbert, Sageret c having succeeded where he Gærtner failed, that perhaps he was not very good gardener, I suspect had no greenhouse. It is evident, as he admits, that numberless experiments must be tried to know the real fertility of any cross: yet it is strange that he shd have got 17/33 20/45 (K) when crossing vars. or pure species, only one or two or (
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Nov. 1867. I put a goose's foot on ground both by day night some very many small Helices of at least 3 species 2 sp. of slugs crawled on it. The goose-foot suspended in greenhouse another on tree by day, the Helices slugs all dropped off in about 5 hours. When goose or duck flies foot closes. Try this during night whilst shells slugs are very young— [remaining 3 pages are blank] [inside back cover] p 30 31 Sweet Peas crossed. , 39 p 1. Hollyhocks [printed label:] FROM HALLIFAX CO. Account Book
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smaller— a shade yellower— certainly a far greater number of minute aborted ovules.— Two flowers in do. state from greenhouse.— vast difference in size of seeds their colour.— Here too male had far more aborted ovules.— I mixed pollen of male female together when viewed under 2/3 object glass I certainly think male pollen darker than can be accounted for by greater diameter, which is often as 5:8.— Under very high power this difference in brownness opacity not very decided.— Compared seeds of male
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The dark purple grows nearest walk: pale crimson further off. 9 1856 June 13th. The Lychnis (red) dioica which has had so much salt Guano, in the female stems are certainly less red less hairy than to other plants which only occasionally have had Guano alone. June 16th Lathyrus Nissolia— seeds raised in Hotbed Greenhouse planted out some— now in flower— clump about 18 inches high. Flowers always 2 on stalk — Lake Red . — Has been treated all spring with salt Guano water.— Specimen dried.— Nos
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same flower. The parents marked C. were fert in Greenhouse were all long-styled children of long-styled Hom. fertilised— So that plants to be marked C.C. which will perhaps flower in 1867 some did flower will be great-grandchildren of first Hom. Union, all intermediate unions having been illegitimate or Homomorphic. 80-8
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government botanic gardens in Perth, Western Australia.] (53 1861 May 8th 1860 I observe about 1/2 cowslips in Greenhouse Experimental beds have long pistils very short anthers with several pollen at base of tube other 1/2 plants (all same on same plant) have very short pistil with long anthers at mouth of tube with much pollen– Case of Compensation.— The long short pistil is rather large rudimentary (in male flowers) is flattened at pole smoother— pale coloured.— The long pistil, which I presume is
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F1583
Book:
Stauffer, R. C. ed. 1975. Charles Darwin's Natural Selection; being the second part of his big species book written from 1856 to 1858. Cambridge: Cambridge University Press.
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effects2 of the pollen not being always applied at exactly the right time or not often enough at successive periods, /10/and of the plant being cultivated in a pot placed not in a greenhouse but in a chamber, lastly of the early castration of the anthers. He admits3 that in order to get the proper maximum, many flowers in successive years should be experimentised on. Hence it is much to be regretted that he did not take for his standards of comparison the same species artificially fertilised
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CUL-DAR157.1.34
Note:
[Undated]
Kidney bean / Young plant raised in Greenhouse — stake supporting huge
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The Complete Work of Charles Darwin Online (1 Kidney Bean.— Young plant raised in Greenhouse— slates supporting huge cotyledon 4 5/8 inch — first internode 7 1/2 inch long— Neither of these move. The upper internode, together with very young one an inch long, were 6 1/2 inches in length, but shortened by tip being curved. This upper internode made a first most regular circle in 2° — A second circle in 1°. 55' — A third in 1°. 55'.— Moves reverse of Watch.— I see a trace of twisting in this
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CUL-DAR157.1.27
Note:
[ny].03.28--[ny].03.29
Stauntonia latifolia — Hothouse properly Greenhouse plant / against sun
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The Complete Work of Charles Darwin Online [27] Stauntonia latifolia — Hothouse properly Greenhouse plant. against sun. Mar. 28 3°. 30' 29 3°. 45 [Climbing plants, p. 15
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CUL-DAR157.1.32
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[ny].04.04--[ny].04.07
Sollya drummondii in Greenhouse moves against sun
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The Complete Work of Charles Darwin Online [32] Sollya Drummondii in Greenhouse moves against sun. — Ap 4— 4°. 25' Ap 5— cold day — 8°. 0' ― 6— 6°. 25' 7— 7°. 5' A plant with long stems like string [Climbing plants, p. 16
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CUL-DAR157.1.63
Note:
[ny].04.04--[ny].04.05
Clematis montata in Greenhouse move against sun
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The Complete Work of Charles Darwin Online [63] Clematis Montana, in Greenhouse [Move] against sun. Ap 4 — 4° 20' ― 5 — 5° 10' Cold day shoot leans well over. After taking a turn in one direction; then kept straight reversed its spiral. (I can now see by looking at the several Clematis; Trop. tricolorum Rhodichiton, Bign. unguis (in contrast with B. buxifolia) that all these plants by aid of leaves make a much more open spiral; becoming in part straight then reversing spire, they ascend to a
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The Complete Work of Charles Darwin Online [33] Glycina in Greenhouse: moves against sun. — 1864. May 13th 3°. 5' ― 3°. 20'} shoot young ― 16th 2°. 5' ― 24 3°— 21' ― 25 2°— 37' ― 25 2°— 35
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CUL-DAR49.50
Note:
1858.05.16
I suspect good generalisation, that wherever there is nectary on one
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In the greenhouse Balsam, there is spur-nectary on lower side of flower, anther and pistil bend downward into gangway. Tropæolum canariense, when pollen shed stigma probably ready, the whole pistil is always slightly bent up to gangway
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The Complete Work of Charles Darwin Online [49] Scyphanthus elegans Greenhouse (with sun) June 13th 1°.— 45' ― 1°. 17'. quickest ever made ― 14.' 1°. 36' ― 1°. 59' (sometimes takes 2 turns) in one direction Yes) ― 2°. 3' [Climbing Plants, p. 18.] Loasa aurantiaca. Greenhouse.......? Greenhouse against sun June 20th. 7° 20 2°. 37' ― 2°. 13' ― 4°. 0' June 21' 2°. 35' ― 22d 3°. 26' ― 23d 3° 5' July 11' another plant mov followed sun 1° 51 1° 46' 1° 41' 1° 48} very hot day {one plant first
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CUL-DAR49.60
Note:
1861.04.08--1861.04.10
Vinca major in Pot — greenhouse [with illustrations]
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The Complete Work of Charles Darwin Online [60] Vinca major in Pot— greenhouse Apr 8th 1861 [annotated sketch] [illeg] orange-coloured curtain More accurate [annotated sketch] upper part of anther forming a cap without pollen. anthers: pollen shed in 2 packets in recesses amongst the woolly white hairs of depending from top of pistil above the circular humid disc - without flower [illeg] no pollen cd get on circular stigma. corolla) pistil Insect must pass proboscis between two of anthers on
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F1715
Periodical contribution:
Darwin, C. R. 1861. Cause of the variation of flowers. Journal of Horticulture and Cottage Gardener 1 (18 June): 211.
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present in my greenhouse, but I know not its name), the central flower in each truss every year comes regular, loses the two dark patches of colour on the two upper petals, and, what is very curious, loses the nectary, which may be seen in all the other flowers cohering to the flower-stalk. In the common Carrot the central floret in the umbel is dark purple and very different from the others; and I find that this central little flower is extremely variable. Are there not other cases of species which
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F1716
Periodical contribution:
Darwin, C. R. 1861. Vincas. Gardeners' Chronicle and Agricultural Gazette no. 37 (14 September): 831-832.
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difference in the results obtained on the one hand by F. A. P. , and on the other by the writer from Kew and Mr. Horwood? Will F. A. P. have the kindness to state, if he sees this notice, whether his plants were in a greenhouse with the windows left open, so that the moths could get access at night?3 Charles Darwin, Down, Bromley, Kent. 1 F.A.P. 1861 wrote: I am surprised at C. W. C.'s assertion in your number for July 27, that Tropical Vincas never produce seed under cultivation if left to themselves
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Nov. 27 /62 Nolana prostrata 9 Plants uncovered, seed weighed 79 gr. ∴ 12 plants would have produced 105 gr. ✔ 12 Plants under net in Greenhouse white thread 3 seeds Black thread 4 all the other pods had opened got mixed with remaining seeds; so that 14 flowers were artificially fertilised in this lot. — Every single plant of the 12 produced some seed, so that none can be strictly dimorphic. — These 12 Plants produced only 64 gr, yet 14 flowers were artificially fertilised; so that Bees aid
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The Complete Work of Charles Darwin Online (66 Nolana prostrata (old notes in Dichogam or Dimorph. [illeg] Germination of seed Portf Portfolio) I believe 30 flowers crossed, 27 pods produced each with 5 seed (In greenhouse) (1867) 32 flowers were crossed with own-pollen (26 having been crossed from same flowers 6 from flower on same plant, but the latter plant not the least advantage) These 32 flowers produced only 6 pods each with 5 seed.— so very sterile with own pollen. In 3 cases 2 seeds
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CUL-DAR108.70
Note:
1862.spring
Wild Cowslips in Pots experimented on in Greenhouse / Short-styled
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The Complete Work of Charles Darwin Online [70] Wild Cowslips in Pots, experimented on in greenhouse in Spring of 1862. Short-styled — (stick) [data not transcribed] (Used in Lythrum paper) (These were the products of the umbels which did not perish from the heat.) 50 crossed first homomorphically with cowslip in 24° heterotrophically with polyanthus 48 59. do do do do do Hybrids Primrose Cowslips Cowslip by pollen of Primrose [data not transcribed] p. 76 Experiment Book gives record of what
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CUL-DAR108.63-64
Note:
[1862].03.07
I have examined Mr Frys "mid-styled" flowers stigma in all stands in very
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As long-styled seed most freely homomorphic seedlings are inherited their own form, may not preponderance of long-styled in our Greenhouse be thus accounted for.—? Ap. 24 62. Mr Crocker has looked says certainly 3 forms in Chinese Primrose at Chichester
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CUL-DAR76.B40
Note:
1862.05.24
Cabbages / Prepotency / 10h 45 put plenty of pollen on 3 lately opened
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not to be easily crossed even late application of pollen produce effect— [Right margin:] Prepotency of pollen of other Var. Quoted do. put plenty of own pollen on 3 open flowers of Cattells early Barnes' Cabbage after 23 hours put pollen of long stamens of Ragged Jack. — marked with black thread. Same remarks applicable in this case as in last. — (Plants killed) June 4th castrated 4 flowers of Ragged Jack Stems nearest to Greenhouse put under net: 6' fertilised 2 with pollen of short stamens of
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CUL-DAR108.135
Note:
1862.05.26
a few Pods — Chinese Primroses brought me by Mr Horwood [numbers of
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The Complete Work of Charles Darwin Online (5 May 26 — 1862— a few Pods — Chinese Primroses; brought me by Mr Horwood (N. B. I have counted all the seeds, except the very small manifestly bad.) The empty pods not brought there were many Horwoods seedlings: thumb-eyed or mid-styled (picked pods by Mr Horwood) These no doubt were so far picked as the upper Ducks were, so that none with no seed were brought [Data and calculations not transcribed] Horwoods seedlings 4 plants kept in my greenhouse
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CUL-DAR157.1.103
Note:
[1862].04.14--[1862].06.15
Solanum jasminoides Greenhouse moves against sun
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The Complete Work of Charles Darwin Online [103] Solanum jasminoides Greenhouse an upright bush. with thin twigs Moves against sun. — sometime stood still or moved but little Ap. 4. 3° 35' — 3° 10' Ap 6 3° 45' — 3° 15' 3°. 26' [calculations not transcribed] June 15th Plant put into Hothouse a good-sized leaf well caught vertical stick in 7°. In greenhouse movements extremely slow, footstalk not acted on during several days by 15 inches of red string — It is remarkable that full-sized leaves
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CUL-DAR51.B10-11
Note:
1862.06.06
Pelargonium / In bedding out scarlet I find 5 sepal-facing anthers longer
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-stamens new fact in Dichogamy probably in all regular flowers has same meaning. But in some of the scarlets there is one small anther on opposite side of nectary, which can be of little use, is sometime absent; in the large fancy Pelargon in Greenhouse is always absent - This probably parent of Beaton's dwarfs. [B10v
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CUL-DAR109.A11
Note:
1862.07.29
Satureia hortensis Summer Savoy / I raised 11 Plants one alone had
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The Complete Work of Charles Darwin Online [A11] Satureia hortensis. Summer Savoy. July 29.62 I raised 11 Plants, one alone had perfect anthers purple ie was Hermaphrodite. The 10 others were female. The only difference in pistils was that in female they were a shade longer. Largely visited by Humble Hive Bees ─ In pot in greenhouse─ Culture under heat perhaps must have increased the Females.─ See about seeding ─ Case exactly like Thyme. ─ Anthers of female a more colourless transparent double
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F1717
Periodical contribution:
Darwin, C. R. 1862. On the two forms, or dimorphic condition, in the species of Primula, and on their remarkable sexual relations. [Read 21 November 1861] Journal of the Proceedings of the Linnean Society of London (Botany) 6: 77-96.
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. Weight of seed in grains. The two homomorphic unions..... 100 45 31 11 100 24 100 35 The two heteromorphic unions..... 100 77 71 39 100 50 100 54 With P. auricula I was unfortunate; my few seedlings, except one poor plant, all came up short-styled; and of these plants several died or became sick, owing to the hot weather and the difficulty of excluding insects and ventilating the corner of my greenhouse enclosed with net. I finally got only two pods from one union, and three from the other. The
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F1717
Periodical contribution:
Darwin, C. R. 1862. On the two forms, or dimorphic condition, in the species of Primula, and on their remarkable sexual relations. [Read 21 November 1861] Journal of the Proceedings of the Linnean Society of London (Botany) 6: 77-96.
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of insects, the six short-styled plants ought to have produced 92 grains' weight of seed instead of only 1.3; and the eighteen long-styled plants, which produced not one seed, ought to have produced above 200 grains' weight. The production of the 1.3 grain of seed in the smaller lot was probably due to the action of Thrips or some minute insect. This evidence is sufficient, but I may add that ten pots of Polyanthuses and Cowslips of both forms, protected from insects in my greenhouse, did not set
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CUL-DAR205.8.6
Note:
1863.04.24--1863.07.21
Clarkia elegans / Plant in Greenhouse / 1 fl[ower] with pollen of little
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The Complete Work of Charles Darwin Online [6] Clarkia elegans Ap. 24th Plant in greenhouse 1 fl with pollen of little anther Black Thread 1 fl with pollen of large anther White Thread For future part more pollen of small anthers. Pollen of short anther mature long before long anther – Both mature long before stigma but pollen having about 8 threads to old anther gets often on the stigma without insect ovarium swells. May 20th I find 6 flowers in which there was no pollen on stigma were
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CUL-DAR109.A12
Note:
1863.07.13
The Summer Savoy raised from the seed from plants in Greenhouse last
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The Complete Work of Charles Darwin Online [A12] July 13' 63 The summer Savoy raised from the seed from plant in greenhouse last summer are quite true all uniform not in least crossed with winter, which shows that the one (?) hermaphrodite sufficed to impregnate all ─ very curious case
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CUL-DAR157.1.89-90
Note:
1863.08.09--1863.08.12
Tropaeolum canariense / Young plant 2 lowest internodes do not move
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Aug 10th I found after night that footstalk of young leaf had wound round stick in direction of reverse of watch — Took away stick in 6°, peduncle which was greatly hooked, quite straightened itself. Accordingly I tied internode above below a young leaf; it moved greatly toward light; but cd. detect no other movement. — Aug. 11' In greenhouse I can see 2 leaves on same stem wound round stick in opposite direction — in one case peduncle completely hooked— In study a young leaf which was
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CUL-DAR157.1.89-90
Note:
1863.08.09--1863.08.12
Tropaeolum canariense / Young plant 2 lowest internodes do not move
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Does not climb Tropæolum Canariense (Peregrinum) Aug 11th I have now ascertained that quite young leaves even leaves 1 1/4 inch in diameter have whole peduncle sensitive if touched on any side in from 1/4 to 1° curl up to that side, thus clasp stick if stationary): Leaf Leaf does not move.— As yet not seen leaf move move that to form a hook, ie. upper basal ends parallel. — I doubt if this plant winds?? Movement to bring peduncle into contact — In greenhouse a leaf which had clasped stick has
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CUL-DAR79.93-97
Note:
1863.08.22--1868.10.22
Delphinium consolida? / Tall branching Larkspur [illegible] white var
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232 self seed weighed 4.42 ∴ less than half the weight this shows how universally small. The crossed seed in weight are to equal numbers of self-seed, as 100:45. (July 8 /68) [Table partly excised, not transcribed] Oct. 22 68. All the crossed plants together, uncovered in greenhouse produced 282 mature immature pods; all the self plants produced only 159 do pods.─) [95
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Cobæa Scandens Oct 29 1863 On flat surface in greenhouse hooks, with rare exceptions, become neatly attached. Whole tendril corresponds to terminal leaflet with foot-stalk. Upper surface of t- angular green. lower surface rounded purple. The Green stripes very narrow on branchlets run to upper surface of terminal hooks; difference of colour probably determines the movement. No movement till tendril comes into contact. It is mere case of leaf turning to light. Hence slow adaptation of branches
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CUL-DAR17.1.B1a-B62
Draft:
1864
'On the movements & habits of climbing plants by Ch Darwin F.R.S. F.L.S'
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Sollya Drummondii (Pittosporaceæ) moves against the sun; kept in greenhouse. April 4' — 5 — 6 7 1s circle 4° 25' 2d do 8° — very cold day 3d do 6° 25' 4th do 7°. 5' Polygonum dumetorum (Polygonaceæ) this case is taken from Dutrochet (p. 299), as I observed no allied plant: moves follows the sun; star three shoots cut off placed in water, made circles in 3° 10', 5° 20' and 7° 15'.— Wistaria chinensis (Leguminosæ), in greenhouse— moves against the sun. May 13th — — 16 1' circle 3° 5' 2d [circle
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CUL-DAR17.1.B1a-B62
Draft:
1864
'On the movements & habits of climbing plants by Ch Darwin F.R.S. F.L.S'
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[First 5 lines already transcribed in the previous page] they ought to have revolved in between 2° and 2° 30'. Shoots of the Kidney-bean were similarly but retarded but in a less degree. I have repeatedly observed that carrying a plant from the greenhouse to my house, or even (4
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CUL-DAR17.1.B1a-B62
Draft:
1864
'On the movements & habits of climbing plants by Ch Darwin F.R.S. F.L.S'
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Tamus communis (Dioscoreaceæ)— a young shoot from a potted tuber in a pot in the greenhouse; follows the sun.— July 7th do — 8' 1st circle in 3° 10' 2d do 2° 38' 3d do 3° 5' July 8' do do 4'' circle 2° 56' 5'' circle 2° 30' 6'' circle 2° 30' Lapagerea rosea (Philesiaceæ), in greenhouse, follows the sun. March 9th March 9th — 10th — 11 — 12 — 13 — 16 1st circle 1' circle do shoot young 1' circle 26° 15' shoot young semicircle 8° 15 2d circle 11° 3d — 15°. 30' 4'' 14° 15' 5 8° 40' placed in hot
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CUL-DAR108.137
Note:
[1864orafter]
Horwoods 4 Plants kept in my Greenhouse — midstyled — self-fertilised
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The Complete Work of Charles Darwin Online [137] Horwoods 4 Plants kept in my greenhouse— midstyled — self-fertilise — produced 180 pods, then I examined 30, of which only 3 produced no seed: Hence. average [calculations not transcribed] 35.5 But if all beneath 14 be omitted (I chose this by chance from results of 1863) we have this average [calculations not transcribed] 42.5 averag
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CUL-DAR17.2.A63-A93
Draft:
[1864]
'On the movements & habits of climbing plants' [continued]
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(Leaf climbers) It is considered a greenhouse plant; but when kept there, the petioles took several days to clasp a stick: in the hothouse a stick was clasped in 7°. In the greenhouse a petiole was not affected by a loop of string, suspended during several days weighing 2.5 2 1/2 grains; (Diagram 3] (a) In this specie plant leaf-climber plant, in no other species leaf-climber seen by me, a leaf grown to its full-sized leaf was capable was capable of clasping; but the movement was so
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CUL-DAR17.1.B1a-B62
Draft:
1864
'On the movements & habits of climbing plants by Ch Darwin F.R.S. F.L.S'
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from one part to another of the greenhouse to another part, always stopped the revolving movement for some a time; hence I conclude that during windy weather twining plants plants natural growing would not during very windy stormy weather make their revolutions. A decrease in temperature was always caused a considerable retardation in the rate of revolution; but Dutrochet (Tom. 17 p. 994, 996) has given such good precise observations on this head with respect to the tendril-bearing pea, that I
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CUL-DAR27.2.A1-A54
Draft:
[1864]
On the sexual relations of the three forms of Lythrum salicaria
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(22bis * Footnote to p. 17 In the spring of 1862 I crossed a number of forty cowslip flowers (P. veris) heteromorphically homomorphically. The plants were accidentally exposed in the greenhouse to too hot a sun, a number of umbels suddenly perished. Some, however, remained in fairly moderately good health, on these there were twelve flowers which had been fertilized heteromorphically eleven which had been fertilized homomorphically. The twelve heteromorphic unions yielded seven fine capsules
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CUL-DAR17.1.B1a-B62
Draft:
1864
'On the movements & habits of climbing plants by Ch Darwin F.R.S. F.L.S'
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the angle between two stems leaf-stalk. I may here state that I ascertained the weights of the string thread used by in all cases by carefully weighing 50 inches of string of thread in a chemical balance, then by cutting I cutting off measured portions lengths.*a The main foot-stalk petiole carries three leaflets; each having a short, but separate petiole; there were but their short petioles are of the leaflets are not sensitive. A young inclined shoot (the plant being in the greenhouse
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F1731
Periodical contribution:
Darwin, C. R. 1864. On the sexual relations of the three forms of Lythrum salicaria. [Read 16 June] Journal of the Linnean Society of London (Botany) 8: 169-196, 1 text figure.
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contained, on an average, 107.3 seed. * In the spring of 1862 I crossed forty Cowslip flowers (P. veris) heteromorphically and homomorphically. The plants were accidentally exposed in the greenhouse to too hot a sun, and a number of umbels perished. Some, however, remained in moderately good health, and on these there were twelve flowers which had been fertilized heteromorphically and eleven which had been fertilized homomorphically. The twelve heteromorphic unions yielded seven fine capsules
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CUL-DAR157.1.82
Note:
1864.01.31
Tropaeolum tricolorum / Tendrils very sensitive on upper & lower surface
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Horace observed that plant climbed in Hot-house in one-way then in other way. I see same fact in Greenhouse — Bears on reversed movement in Pea echinocystis A common twining plant could not do so
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CUL-DAR157.1.84
Note:
[1864].02.06
Tropaeolum tricolorum grandiflorum — in Greenhouse — Tip of t[endrl]
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The Complete Work of Charles Darwin Online [84] Feb 6. Tropæolum tric. grandiflorum— in Greenhouse— Tip of t. pointed, a little flattened, slightly furrowed on upper surface — sometimes a trace enlarged — formed of concave cells thin petiole— le evidently repeating leaf— but no trace of divisions, or [illeg] nature. The tip in movement rudiment of separated laminæ of true leaf— In Greenhouse move spontaneously to stick twine beautifully. — so omit all allusion to Hothous
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The Complete Work of Charles Darwin Online [22] Lapagerea Rosea. Mar. 64— Moves with Sun; in greenhouse first circle Mar 7. not complete in 26°— Mar 9. completed 26°. 15' Mar 10. Semi circle 8°. 15' = 16°. 30' Mar 11 circle in 11° Mar 12. Do — 15°. 30' Mar 13 Do 14°. 15' At Commencement shoot was very young. In Hothouse Mar 16 in 8°. 40'— then shoot remained stationary the whole of the next day.— [Climbing plants, p. 15
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CUL-DAR157.1.60
Note:
1864.03.10
Tecoma radicans / I observe in hothouse & greenhouse the plant not having
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The Complete Work of Charles Darwin Online [60] Mar 10 1864 Tecoma Radicans I observe in hothouse greenhouse the plant not having been moved for several days that the shoots move from side to side slightly, caused I presume by the changes in the sun, for a plant brought into the room, merely moved down towards the window during the day, quickly upwards after dark. There were irregularities in some cases caused perhaps by changes in the light. The young shoot 3 or 4 inches long bearing the
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The Complete Work of Charles Darwin Online [24] April 9th 1864 Hop— Plant in Greenhouse (put in House. 1 ft. high — moved with sun. two circles each in 4° 16' ie on average °2°. 8'.— Average of 7 revolutions = 2°. 8' warm weather during April August [calculations not transcribed] [24v
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CUL-DAR157.1.64
Note:
1864.04.25--1864.04.27
Clematis sieboldii / Leaves with long & sensitive peduncle
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Rhodichiton volubili leaf-stalks either take full turn or half turn then bend up in odd manner. — Cl. Sieboldii in greenhouse — against sun. Ap 25th. — 2°. 55.' ― 2°. 45' ― 26th 3°. 55' ― 27th 4°. 30' (shoot getting older) [calculations not transcribed] average of 3° 11
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CUL-DAR157.1.31
Note:
1864.05.18--1864.05.26
Hibbertia dentata placed in Hothouse (Greenhouse plant)
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The Complete Work of Charles Darwin Online [31] Hibbertia dentata, placed in Hot-house (greenhouse plant) 1864 Moved with sun May 18th — 7°. 20' May 19th against sun 7°. 0' ― 20th. moved against sun about 1/3 of a circle then, stood still. (26th went 2/3 of a circle with the sun then returned in (11°, 46' I placed a stick, incessantly it curled round it uncurled again — this was repeated an endless number of times— The curling uncurling being often 1/2 2/3 of a circle— Internodes very long
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CUL-DAR157.1.100
Note:
1864.06.03--1864.06.12
Lophospermum scandens purpureum / Young plants with long internodes
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June 12th. Rubbed 4 very young growing stems in Hothouse 1 in Greenhouse in 2° they were all 5 well bent after a dozen rubs,— but some older stems did not move. — Tail of worsted caused movement in leaf-stalk average 3° 15' [calculations not transcribed
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F1733
Periodical contribution:
Darwin, C. R. 1865. On the movements and habits of climbing plants. [Read 2 February] Journal of the Linnean Society of London (Botany) 9: 1-118, 13 text figures.
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regularly at an average rate of 3 h. 26 m. The shoots, however, sometimes stand still. It is considered a greenhouse plant; but when kept there, the petioles took several days to clasp a stick: in the hothouse a stick was clasped in 7 h. In the greenhouse a petiole was not affected by a loop of string, suspended during several days and weighing 2 1/2 grains; in the hothouse one was made to curve by a loop weighing 1.64 (and, on the removal of the string, became straight again), but was not at
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F834a
Book:
Darwin, C. R. 1865. On the movements and habits of climbing plants. London: Longman, Green, Longman, Roberts & Green and Williams & Norgate.
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regularly at an average rate of 3 h. 26 m. The shoots, however, sometimes stand still. It is considered a greenhouse plant; but when kept there, the petioles took several days to clasp a stick: in the hothouse a stick was clasped in 7 h. In the greenhouse a petiole was not affected by a loop of string, suspended during several days and weighing 2 1/2 grains; in the hothouse one was made to curve by a loop weighing 1.64 (and, on the removal of the string, became straight again), but was not at
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F1733
Periodical contribution:
Darwin, C. R. 1865. On the movements and habits of climbing plants. [Read 2 February] Journal of the Linnean Society of London (Botany) 9: 1-118, 13 text figures.
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; follows the sun. Three shoots cut off and placed in water made circles in 3 h. 10 m., 5 h. 20 m., and 7 h. 15 m. Wistaria Chinensis (Leguminosæ), in greenhouse, moves against the sun. h. m. May 13, 1st circle . . . . . . 3 5 „ 13, 2nd „ . . . . . . 3 20 „ 16, 3rd „ . . . . . . 2 5 h. m. May 24, 4th circle . . . . . . 3 21 „ 25, 5th „ . . . . . . 2 37 „ 25, 6th „ . . . . . . 2 35 Phaseolus vulgaris (Leguminosæ), in greenhouse, moves against the sun. h. m. May, 1st circle . . . . . . 2 0 „ 2nd
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F834a
Book:
Darwin, C. R. 1865. On the movements and habits of climbing plants. London: Longman, Green, Longman, Roberts & Green and Williams & Norgate.
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; follows the sun. Three shoots cut off and placed in water made circles in 3 h. 10 m., 5 h. 20 m., and 7 h. 15 m. Wistaria Chinensis (Leguminosæ), in greenhouse, moves against the sun. h. m. May 13, 1st circle . . . . . . 3 5 „ 13, 2nd „ . . . . . . 3 20 „ 16, 3rd „ . . . . . . 2 5 h. m. May 24, 4th circle . . . . . . 3 21 „ 25, 5th „ . . . . . . 2 37 „ 25, 6th „ . . . . . . 2 35 Phaseolus vulgaris (Leguminosæ), in greenhouse, moves against the sun. h. m. May, 1st circle . . . . . . 2 0 „ 2nd
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F1733
Periodical contribution:
Darwin, C. R. 1865. On the movements and habits of climbing plants. [Read 2 February] Journal of the Linnean Society of London (Botany) 9: 1-118, 13 text figures.
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(MONOCOTYLEDONS, continued.) Asparagus (unnamed species from Kew) (Liliaceæ) moves against the sun, placed in hothouse. h. m. Dec. 26, 1st circle . . . . . . 5 0 „ 27, 2nd „ . . . . . . 5 40 Tamus communis (Dioscoreaceæ). A young shoot from a potted tuber placed in the greenhouse; follows the sun. h. m. July 7, 1st circle . . . . . . 3 10 „ 7, 2nd „ . . . . . . 2 38 „ 8, 3rd „ . . . . . . 3 5 h. m. July 8, 4th circle . . . . . . 2 56 „ 8, 5th „ . . . . . . 2 30 „ 8, 6th „ . . . . . . 2 30
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F834a
Book:
Darwin, C. R. 1865. On the movements and habits of climbing plants. London: Longman, Green, Longman, Roberts & Green and Williams & Norgate.
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(MONOCOTYLEDONS, continued.) Asparagus (unnamed species from Kew) (Liliaceæ) moves against the sun, placed in hothouse. h. m. Dec. 26, 1st circle . . . . . . 5 0 „ 27, 2nd „ . . . . . . 5 40 Tamus communis (Dioscoreaceæ). A young shoot from a potted tuber placed in the greenhouse; follows the sun. h. m. July 7, 1st circle . . . . . . 3 10 „ 7, 2nd „ . . . . . . 2 38 „ 8, 3rd „ . . . . . . 3 5 h. m. July 8, 4th circle . . . . . . 2 56 „ 8, 5th „ . . . . . . 2 30 „ 8, 6th „ . . . . . . 2 30
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F1733
Periodical contribution:
Darwin, C. R. 1865. On the movements and habits of climbing plants. [Read 2 February] Journal of the Linnean Society of London (Botany) 9: 1-118, 13 text figures.
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-bean were similarly retarded, but in a less degree. I have repeatedly observed that carrying a plant from the greenhouse to my house, or from one to another part of the greenhouse, always stopped the movement for a time; hence I conclude that naturally exposed plants would not make their revolutions during stormy weather. A decrease in temperature always caused a considerable retardation in the rate of revolution; but Dutrochet (tom. xvii. pp. 994, 996) has given such precise observations on this
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11% |
F834a
Book:
Darwin, C. R. 1865. On the movements and habits of climbing plants. London: Longman, Green, Longman, Roberts & Green and Williams & Norgate.
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-bean were similarly retarded, but in a less degree. I have repeatedly observed that carrying a plant from the greenhouse to my house, or from one to another part of the greenhouse, always stopped the movement for a time; hence I conclude that naturally exposed plants would not make their revolutions during stormy weather. A decrease in temperature always caused a considerable retardation in the rate of revolution; but Dutrochet (tom. xvii. pp. 994, 996) has given such precise observations on this
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F1733
Periodical contribution:
Darwin, C. R. 1865. On the movements and habits of climbing plants. [Read 2 February] Journal of the Linnean Society of London (Botany) 9: 1-118, 13 text figures.
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growth the finest fibres and secrete an adhesive cement. Eccremocarpus scaber (Bignoniaceæ).—Plants in the greenhouse, though growing pretty well, showed no spontaneous movements in their shoots or tendrils; but, removed to the hot-house, the young internodes revolved at rates varying from 3 h. 15 m. to 1 h. 13 m.: at this latter unusually quick rate one large circle was swept; but generally the circles or ellipses were small, and sometimes the course pursued was extremely irregular. An internode
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F834a
Book:
Darwin, C. R. 1865. On the movements and habits of climbing plants. London: Longman, Green, Longman, Roberts & Green and Williams & Norgate.
Text
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PDF
growth the finest fibres and secrete an adhesive cement. Eccremocarpus scaber (Bignoniaceæ).—Plants in the greenhouse, though growing pretty well, showed no spontaneous movements in their shoots or tendrils; but, removed to the hot-house, the young internodes revolved at rates varying from 3 h. 15 m. to 1 h. 13 m.: at this latter unusually quick rate one large circle was swept; but generally the circles or ellipses were small, and sometimes the course pursued was extremely irregular. An internode
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F1733
Periodical contribution:
Darwin, C. R. 1865. On the movements and habits of climbing plants. [Read 2 February] Journal of the Linnean Society of London (Botany) 9: 1-118, 13 text figures.
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measured lengths*. The main petiole carries three leaflets; but the short petioles of these leaflets are not sensitive. A young inclined shoot (the plant being in the greenhouse) made a large circle opposed to the course of the sun in 4 h. 20 m., but the next day, being very cold, the time was 5 h. 10 m. A stick placed near the revolving stem was soon struck by the petioles which stand out at right angles, and the revolving movement was arrested. The petiole then began, being excited by the contact
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7% |
F834a
Book:
Darwin, C. R. 1865. On the movements and habits of climbing plants. London: Longman, Green, Longman, Roberts & Green and Williams & Norgate.
Text
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PDF
measured lengths*. The main petiole carries three leaflets; but the short petioles of these leaflets are not sensitive. A young inclined shoot (the plant being in the greenhouse) made a large circle opposed to the course of the sun in 4 h. 20 m., but the next day, being very cold, the time was 5 h. 10 m. A stick placed near the revolving stem was soon struck by the petioles which stand out at right angles, and the revolving movement was arrested. The petiole then began, being excited by the contact
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CUL-DAR110.A56
Note:
1865.04.19
Crossed 12 fl[owers] (white thread) of long-styled Pulmonaria
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all fl. which had opened off one plant, at corner nearest greenhouse, see if these seed. N.B. whiles other sp was in flower, the bees sucked some on margins, whilst under net, so observe whether any in this position set seed.— No not more than elsewhere
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CUL-DAR78.17-40,42-45
Note:
1866--1872
Mimulus [comparison of crossed and self-fertilised plants in height,
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taller from greenhouse (5
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CUL-DAR78.139-156
Note:
1866--1871
Cabbage [comparison of crossed and self-fertile plants in number of seed,
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Barnes Cabbage— Jan 16— 1871 Young plants kept in greenhouse, raised from seed described in last page measured to tip of leaves [table not transcribed] 21
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CUL-DAR108.119
Note:
1866
Long-styled Cowslip — Pot / Marked C in Greenhouse — Plants from a
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The Complete Work of Charles Darwin Online [119] Long-styled Cowslip—Pot. — 1866 Marked C. in Greenhouse — Plants from a Hom. plant— so that 2 generations of Hom. union: or grandchildren of first. Hom: union — Six plants preserved 25 flowers fertilised by pollen of own flower. (see note about state of pollen, where I enumerate how many came long-styled.) [Data and calculations not transcribed] 16.3 seed per pod 60/1000 pods set— Some of All the plants were more sterile than others produced no
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CUL-DAR78.122-131
Note:
[1866]--1871
Candytuft crimson — Iberis umbellata var Kermesiana [crossed vs
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N. B. Under net in Greenhouse Lettington saw small fly, flying from flower to flower sucking. Hence it is just possible plants last year were not well protected the self were crossed, but not at all probable. — (June 26— 68 some seeds, germinated were sown on both sides of Pot with Calceolaria— crossed flowered first; but tallest crossed only 4 1/2 inch high tallest self 7 inches high — so selfs great advantage— (p 16
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CUL-DAR78.164-182
Note:
[1866]--1871
Petunia [comparison of crossed and self-fertile plants in number of seed,
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The Complete Work of Charles Darwin Online (25 Petunia seed of common garden var. [dingy] purple. viz Violacea?) 6 4 Plants in separate pots full of same size in greenhouse under net. — 6 flowers crossed by distinct plant. produced 6 pods:— 6 fert by pollen of own flower produced 3 pods not one pod was produced by self-fertilisation: though 4 were marked Seeds from self-fert. by eye less than with from crossed. The 6 seed from the 6 weighed 4.44 grains; therefore from three ( all produced
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CUL-DAR78.122-131
Note:
[1866]--1871
Candytuft crimson — Iberis umbellata var Kermesiana [crossed vs
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Crimson Candy-tuft 1869 June 5th 1869 measured the tips of Leaves Aug 2d. 1869 measured the end of Pods [Table, partly excised, not transcribed] Sept 9. 1869 The above Plants, left uncovered in greenhouse Pots rather small, so plants unproductive many capsules on all plants abundant Pot I Crossed Plants number of seed self-fert Plants number of seed I (3 plants) 74 79 III (4 Plants) 132 75 206 154 (I do not think there any evidence of the self-plant having become sterile.—) 16
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CUL-DAR78.192-196
Note:
[1866]--1867
Lupinus luteus [comparison of crossed and self-fertile plants in number
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Image
some germinated seed was planted on each side of pot of Calceolaria Crossed 21 1/2 — 18 in. Self 20 — 9 1/2. So that of all 8 eight matched plants the crossed in every case the tallest. (The plant were left to fertilise themselves uncovered in greenhouse, where few Bees entered they set very badly produced only bad empty Pods. All the crossed Plants produced only 9 good pods containing 31 seeds, average 3.4. All the self Plants produced only 7. good pods containing 5 21 seed, on average 3. 0 3
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CUL-DAR78.186-189
Note:
[1866]--1869
Marjoram [comparison of crossed and self-fertile plants in speed of
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Image
seed-shells; certainly those under net as forwards or more forwards than those exposed visited by insects.) (Nov. 2d the crossed seed on sand have protruded rootlets apparently quicker than the uncrossed certainly seem more vigorous.) Pot I. transplanted ∴ equal Kept in Hothouse Dec 30' moved into greenhouse Pot 2. do ― ― ― greenhouse; larger Pot. (Dec. 1, Pot I no difference in plants hardly more than cotyledons developed. Pot. 2 crossed rather finest, — In sand self-fert. finest, but these
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CUL-DAR78.200
Note:
[1866--1867]
Parsley Plants growing close together one covered up — several left
Text
Image
Hothouse.) almost all killed by H. H moved into greenhouse. — (Pot I. (Dec. 1') the self-fertilised tallest! Pot 2. 3 Transplanted seeds no difference in Hothouse ― 3 If any difference self-fert. have advantage.) (Dec. 16th Parsley in sand self-fert. perhaps tallest) — (Pot 2 perhaps do, this pot has been moved into greenhouse, for wd not stand Hot-house. Pot 3 do —) (Dec. 30 Parsley in sand crossed plant finest) (Feb. 10th in sand, crossed rather the advantage: Pot 2. crossed rather finest plants
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CUL-DAR78.122-131
Note:
[1866]--1871
Candytuft crimson — Iberis umbellata var Kermesiana [crossed vs
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Iberis umbellata — Plants from last Page. Plants measured to tip of convex heads of flowers on July 1871. In Pots in greenhouse. — All the crossed plants, except those marked P. showed by their pale pink flowers, that they had been crossed— The self-plants rich purple. (—This good case of colour not blending, 6 of crossed plants exactly as purple as pure selfs — all the rest pale pink like ♂. —) (The refuse seed, more or less germinated was planted in 2 separate rows in open ground — The self
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CUL-DAR78.133-137
Note:
[1866]--1868
Sweet pea [comparison of crossed and self-fertile plants in height,
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Image
July 8 68/ Some crossed self-fert. sweet Peas were sown in Box with Brugmansia, where the convolvulus wd not grow, but these grew well. — The tallest plants measured, — (These Peas belong to lots measured on p. 18 C. In Greenhouse July 8' Aug 23' Crossed ft in 6 — 5 1/2 5 — 8 1/2 inches 9 0 1/2 Selfs ft inches 6 — 5 1/2 7. 11. Produced 7 Pods Produced 9 Pods The self flowered first, have always until last measurement on Aug. 25 beaten crossed Some of same 2 lots were sown out of doors in poor
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CUL-DAR78.139-156
Note:
1866--1871
Cabbage [comparison of crossed and self-fertile plants in number of seed,
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planted then in Pot I in greenhouse. I also picked up Oct 18 all the crossed plants which were most advanced left the remainder in equal state to grow as well as they could in pure sand. — Oct 26. The crossed seedlings, as yet only with cotyledons in glass of sand pot, a little taller than the uncrossed. — Nov 2d only a shade taller.─ [21v
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CUL-DAR78.164-182
Note:
[1866]--1871
Petunia [comparison of crossed and self-fertile plants in number of seed,
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Image
Petunia, Augt 30 /71/ Same plants as on last page p. 26 H now growth nearly completed — drawn up, tied to stick — Green House — measured to tips of stems (— measured to base of calyx. —) [Table excised] These plants in greenhouse all seem sterile. — The self-fertilised plants are almost quite uniform in colour— flesh coloured or pale purple— The intercrossed more variable from nearly white to pale pink purple. The plants crossed by Cattell pollen all much more purple, showing really crossed
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CUL-DAR78.186-189
Note:
[1866]--1869
Marjoram [comparison of crossed and self-fertile plants in speed of
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Marjoram May 9' 1869 — Pot I, which from having been long kept in greenhouse did not like being turned out there is wonderful difference in the size of clump, which has increased by stolons. Crossed Self Pot I. plant 10 x 4 1/2 inch 3 1/2 x 2 1/2 tallest young stems 5 1/2 inches — 4 inches high Pot II. 18 x 9 inches tallest stem 8 1/2 inches 12 x 4 1/2 — 6 inches high Pot III. Both sides equal. — (Aug 2d. 1869 In Pot I II the inequality in the size of clumps remains about the
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CUL-DAR78.192-196
Note:
[1866]--1867
Lupinus luteus [comparison of crossed and self-fertile plants in number
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(Dec. 30th Plant no' 4 is now a shade taller than no. 10.) (Feb. 10th Plant no' 9 the finest much most healthy of all 3 plants ― — — 10' do do do. do. —) (March 21. Crossed plants no 9 10 are now on point of flowering, but only 9 inches high healthy — It is clear planting in autumn in greenhouse most injurious: One of self-fert plants, which is very unhealthy, yet is going soon to flower is 8 inches high: all the others are wretched will mostly die. —) (June 3d Only 1 self-fert plant set pod
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CUL-DAR78.192-196
Note:
[1866]--1867
Lupinus luteus [comparison of crossed and self-fertile plants in number
Text
Image
Pots, plants 4 inches to tip of leaves crossed rather finest tallest. — great difference in plants in red Calceolaria Pot) (Jun 20 In pot I, II III The self have flowered first!) (see next page for measurement) (31 C Lupinus luteus. The plants on p. 31. B in greenhouse have now arrived at full height. — [31Cv
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CUL-DAR78.72-111
Note:
1866--1872
Ipomoea purpurea / convolvulus [comparison of crossed and self-fertile
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Ipomoea— Plants on last page. In Pot IV. Plants became unhealthy from some unknown cause especially on crossed side, experiment, not fair. At first (see last page plants crossed by Cattell plant were taller than those the intercrossed Red cross means flowered first two crosses means same time. I took by chance Pot V. collected all pods, spontaneously fert. in greenhouse, uncovered large small, (some not some very small hardly worth counting on both sides: the 3 crossed Plants produced 104 Pods
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CUL-DAR78.122-131
Note:
[1866]--1871
Candytuft crimson — Iberis umbellata var Kermesiana [crossed vs
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Candy-tuft crimson in greenhouse 1867 The plants supposed to be crossed were artificially crossed ie whole capitulum smeared with pollen from distinct plant. 2 or 3 times 3 Heads thus done, produced 108 seed, which weighed 3.57 gr. Several Heads on plants which had been self-fert. were smeared with own pollen spont. self-fertilised under net these heads were quite as fine as the crossed heads. The seed was apparently finer, but hard to judge for it differed much in size from different heads
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CUL-DAR78.122-131
Note:
[1866]--1871
Candytuft crimson — Iberis umbellata var Kermesiana [crossed vs
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Candy-tuft. The plants on last page had individual flowers, 24 in number, carefully crossed by self, 21 set seeds, all these except 7 pairs produced 2 seeds. — The Heads under net were carefully protected this year in Greenhouse, set seeds spontaneously very badly; so that 12 Heads produced only 39 seeds; 8 of these heads yielded only 3 seeds; few flowers produced 2 seed; unfortunately I did not observe whether the crossed self-fert. plants were equally spontaneously sterile. — All the heads
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CUL-DAR78.133-137
Note:
[1866]--1868
Sweet pea [comparison of crossed and self-fertile plants in height,
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Sweet Pea. 1867 in greenhouse. Two plants, descended from Painted Lady Purple crossed, had some 6 flowers crossed, produced 6 pods, with average of 6.0 (Max 7. Min 5) peas per pod— which 36 peas weighed 51.12 gr. Therefore 46 Peas would have weighed 65.32 gr. Ten pods spont. self-fert. under net on these crossed plants were saved they contained on average 4.6 peas per pod (with max. of 6 min of 3) these 46 peas weighed 62.77 gr. So crossed peas more numerous rather heavier. (A painted Lady
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CUL-DAR78.17-40,42-45
Note:
1866--1872
Mimulus [comparison of crossed and self-fertilised plants in height,
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The Complete Work of Charles Darwin Online (5 Mimulus, seeds bought as mixed vars were raised all in same pan in greenhouse covered with net — vars. nearly similar, except in tint of colour. Flowers yellow spotted with orange-brown. They are fertile when not artificially fert. Six pods fert, with pollen from same flower Six pods ― ─ … from distinct plant All 12 flowers produced pods. Seeds too numerous to count, but placed in equal watch-glasses the crossed seeds perhaps appeared rather most
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CUL-DAR78.183-185
Note:
[1866]--1868
Lobelia ramosa [comparison of crossed and self-fertile plants in number
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advantage rather more of them being cracked. (Nov. 4th certainly the plumu radicles of crossed seed are now more fully protruded than in any of uncrossed.) Pot. I seeds were sowed on both sides, (but did not germinate well) Pot 2. seedlings equal transplanted — Hothouse Pot 3. do do. do Greenhouse (Dec. 1 Pot. 2. Hot House Crossed uncrossed equal, except one crossed tallest Pot 1. all equal. — March 31. crossed rather finest In sand, the crossed perhaps the healthiest. (Dec 16 Pot. 2. Crossed a
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CUL-DAR78.186-189
Note:
[1866]--1869
Marjoram [comparison of crossed and self-fertile plants in speed of
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. Pot I crossed flowered first [table not transcribed] (July 15 crossed Plants in all 3 Pots flowered first.) [table not transcribed] Aug 23d beginning to seed In Pot I II. (the former in greenhouse) are been kept in Pots have very few stems on self-side, compared with crossed side— In Pot III. [28Av
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CUL-DAR78.72-111
Note:
1866--1872
Ipomoea purpurea / convolvulus [comparison of crossed and self-fertile
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in feet (A) 7: 2 1/2 7:4 Self 4: 2 feet 7: 3 (A) 5: 2 Pot III Crossed 7: 3 1/2 feet 4: 6 1/2 4: 0 1/2 Self 5: 4 1/2 feet 3} 3: 2 1/2} aphides stopped growth add viz these 3 Pots Pot. IV. This pot was placed in Greenhouse near outside weather was cold, all 3 self were killed, 3 crossed have survived are twining though injured. Pot V. This pot. similarly placed, with 6 plants on each side. (over) [10Nv
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CUL-DAR78.72-111
Note:
1866--1872
Ipomoea purpurea / convolvulus [comparison of crossed and self-fertile
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Ipomoea, Few seed of self (not tall Plant A), crossed again crossed in usual manner from 9th generation; so these plants will be 10th generation — only 6 on each side [Table excised] seeds germ on sand planted in beginning of July 1871. — The crossed plants in this pair sick with crumpled leaves, the self- plant extra vigorous Nov. 21— 71 The 6 crossed plants in Greenhouse spontaneously fertilised produced 101 ripe unripe, large small pods. weight of 100 (by calculation 454 gr The 6 self
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CUL-DAR79.2-8
Note:
1866--1872
Carnations (Dianthus caryophyllus) A large bed of plants raised from
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Greenhouse.— [38Bv
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CUL-DAR78.72-111
Note:
1866--1872
Ipomoea purpurea / convolvulus [comparison of crossed and self-fertile
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(In Pot II One self- plant flowered one day before crossed, X which I never saw before. Sept 6th 1867.) In the same Pot I, placed under net, 30 fl. we on the crossed side were fert. by crossed plants yielded 26 pods with 123 seed, ie 4.73 seed per pod. 30 flowers were self- art. fertilised during autumn in greenhouse on the self-plants produced 23 pods produced 102 seed, ie 4.43 seed per pod; very small difference in actual fertility. The 123 crossed seed weighed 53.22 gr ie 100 weighed 43.27
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CUL-DAR70.58-60
Note:
1866.01.10--1866.03.19
From J Traherne Moggridge from Mentone received Dec 30 1865 [Ophrys
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(3) March 19th 1866. No 4. (a) like D} 2 sticks Greenhouse But 2 spikes in same pot differed in colour of sepals petals (Jan 25th do. Both same 1867) One marked 1 stick has green petals edged with brown green sepals like B. Nr 4 (b) like C. but Labellum with no point N 4 (e) like B. (see Back) but with little point to Labellum Feb 4th 1867- like D in colour do do. do Nr 4 (f) like D. with Lab. approaching in colour to G. Jan. 25 /67/ the same but nothng peculiar in Labellum No 5. like C. but
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17% |
CUL-DAR78.4-10
Note:
1866.06.18.--1869.04.23
Lythrum / Antirrhinum (peloric) / Myosotis alpestris / Cowslip-Polyanthus
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Long-styled Cowslip-Polyanthus These are P. Sinensis experiment June 1868 — Four of the 20 long-styled gr-gr-grandchildren (mentioned at p. 423 of my paper see Exp. Book p. 79 for planting — Illegit. Progeny) Cowslip-polyanthus were kept in greenhouse, 30 fl. were self-fertilised yielded 17 pods, containing on average 32.0 seed with max. of 50 min. of 20 — seed mostly fine good — This is surprising fertility for illegitimate union, now many times repeated — I think, (compared with my paper p
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CUL-DAR78.4-10
Note:
1866.06.18.--1869.04.23
Lythrum / Antirrhinum (peloric) / Myosotis alpestris / Cowslip-Polyanthus
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hours 2 more had done so These plants germinated more freely or in greater numbers than those below. — [in margin:] The parents of the seed grown in greenhouse. For comparison sown in same glass, seeds of Cowslip-Poly, which had been spontaneously fert by both forms growing close together (see Exp. Book p. 79); but those plants were children of long- short styled Hom. plants heteromorphically fertilised. More than twice as many of these seeds sown. — ✔ Aug 2d. 10° P.m. interval 2 days 14 hr. one
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The Complete Work of Charles Darwin Online (6 Calceolaria, plants raised from seed, called common mixed a plant under net, spontaneously set its own pods— Six flowers fert. by pollen of own flower six by pollen of a distinct plant of same general appearance in greenhouse.— owing to minuteness of seed, adhering to capsule I could not compare quantity carefully in the two lots; certainly not much difference Aug 9th 7° 30' P.m — planted in sand in same glass— Aug 14th 2° 30' P.m one of the
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(Calceolaria) Aug 22d 3° P.m. a great number of seeds have now germ. in both lots formed cotyledons; by holding up to light both lots in as nearly equal a state as possible; if any advantage the uncrossed have it so I picked up one or two tallest: now planted on each side of same pot placed in greenhouse. — Sept. 21st By holding pot horizontally I can now see that clump of tallest crossed plants are taller than tallest clump of uncrossed. Sept. 28 —The largest leaf of crossed here shown some
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CUL-DAR108.121
Note:
1867
Cowslip-Polyanthus Long-styled (C) which seeded during 1866 in Greenhouse
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The Complete Work of Charles Darwin Online [121] C 1867 7. do 15 poor seeds 9 16 poor some few better 11 poor mouldy Cowslip-Polyanthus Long-styled (C.) which seeded during 1866 in greenhouse was very fertile; some plants were planted in Orchard, but I neglected to record how many 20 flowers were fertilised with own pollen, probably 10, produced only 5 pods, with seed as above wretched. — But then the weather was atrociously bad
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CUL-DAR76.B99
Note:
1867
Cuphea purpurea — At first many flowers were fertilised owing to my not
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The Complete Work of Charles Darwin Online (64 Cuphea purpurea. —1867 At first many flowers were fertilised, owing to my not knowing when stigma was mature, so no account kept of numbers crossed self-fertilised.— None set spontaneously.— Average of 6 crossed. 6.8. ― of 6 self-fert. 8.8. So of few pods self- gives largest average.— Crossed─ 8 3 12 6 6 6 Self-fert (In greenhouse) 14 10 16 3 8 2 6/41 6/53 (Keep) [calculations not transcribed] 112.8 say 113 Introduce often Rosaceæ Foot-not
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CUL-DAR79.65-71
Note:
1867--1870
Viola tricolor / Three plants, very similar, seedlings, large-flowered,
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Viola tricolor 1869 April 27 — 1869 — measured to tip corner of little leaves at end of latters stem June 2 1869 ─ most Plants nearly done flowering [table not transcribed] April 27 The plants have flowered yesterday in Pot I, II III. *The self plants in Pot III (Jan 2d 69) are yellower less [illeg] than the crossed June 2d 69 Owing to plant kept in greenhouse, with few Bees confined in growth, only few pods produced; these few confined to Pot III IV on crossed plants alone.─ /over 59
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CUL-DAR78.161-163
Note:
1867--1869
Verbascum lychnitis / These plants were raised from self-fertilised
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capsules of placed in watch glasses, seemed not much different in number. — But the 4 by own flower pollen weighed 0.61 gr 4 by distinct plant of same parentage weighed 0.64 4 by wild plant, seemed clearly finer seed weighed 0 88. Planted in large flower pot on Oct. placed in greenhouse (2
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CUL-DAR78.197-199
Note:
1867
Lupinus pilosus [comparison of crossed and self-fertile plants in height
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left in pot.) (June 21. 1868 the crossed flowered first, 27 inches to tip of flower-stem the self-plant, 21 1/2 inches high) (Aug 24. Same plant after regrowth. Crossed. 33 inches — self 26 1/2 inches) (The crossed plant spont. fert, uncovered in greenhouse produced 8 pods with 22 seeds average 2.77. seed. The self Plant produced only 2 pods with 5 seed, average 2.50 — seed. — [calculations not transcribed
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CUL-DAR79.101-104
Note:
1867--1869
Papaver vagum from Dr E Bornet of Antibes / 10 flowers crossed with
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Papaver vagum Pot I— Both Two plants on both sides in pale flowers Both crossed 23 inches high, self 25 1/2 high so taller!! July 6 /68/ Plants now in full flower.— Pot Crossed— Self-fertilised I 23 inches 22 25 1/2 25 III 20 1/2 21 IV Plants from 10-11 inches high not in flower self-plants rather the highest 1868 I crossed some myself some flowers on the plants in Greenhouse with care acted on young flowers, to prevent their having been first self-fertilised. — These plants from being grown
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CUL-DAR79.117-136
Note:
1867--1871
Tobacco in greenhouse / Nicotiana tabacum / 12 flowers crossed gave 10
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The Complete Work of Charles Darwin Online (94 Tobacco in greenhouse 1867 Nicotiana tabacum 12 flowers crossed gave 10 pods 5 pods contained by weight only 14.40 gr. of seed. 12 fl. self-fert. gave 11 pods 5 pods contained 25.78 gr. of seed; so very much fewer, more as far as 5 pods serves. I was so much surprised at this result that I cleaned 5 more pods of each sort. now 10 crossed pods weighed 31.70 gr of seed 10 self-pods [do] 47.67 gr. So self much more fertile!!! (May 20' 68/ In Pot 2 3
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CUL-DAR108.122
Note:
1867.04.00
Some plants of C which flowered in Greenhouse last year have been placed
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The Complete Work of Charles Darwin Online [122] Some plants (see Back) of C. which flowered in Greenhouse last year, have been placed out of Pots in Orchard covered with net and flowers have been fert with pollen of own flower [122v
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CUL-DAR108.122
Note:
1867.04.00
Some plants of C which flowered in Greenhouse last year have been placed
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April 1867 Cowslip-Polyanthus — in Greenhouse Marked C. C. (ie children of C. great-grand-children of first Hom. union, see p. 79 Exp. Book.) Pollen just examined appears good. Also examined pollen of plant of C. growing in orchard this seems good. These are From plants of C.C. ; are as two three have flowered, both all long-styled two b 10 flowers have been fert, with pollen of same flower.— Experiment book, p. 79
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CUL-DAR108.123
Note:
1867.07.00
Cowslip-Polyanthus Long-styled C.C. seedlings of C artificially self-fert
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The Complete Work of Charles Darwin Online [123] 1867. July. Cowslip-Polyanthus Long-styled. C. C. seedlings of C. self artificially self-fert in Greenhouse in the spring.— Average of 5 pods 21.0 seed. 10 flowers were fertilised. [calculations not transcribed
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CUL-DAR185.18
Correspondence:
Darwin Emma née Wedgwood to Darwin William Erasmus
[1867?][.04.00?]
Darwin Emma née Wedgwood to Darwin William Erasmus
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he thinks the frequent doses of Chalk Mag. [Magnesium] do him real good the Podophillin once in 4 or 5 days makes him feel wonderfully well for a day. The pipes are ordered for the new greenhouse I think it will be begun soon. Luckily Mack has taken a turn, as most strangely Fanny has not taken alarm from the Telegrams letters they have been to Naples since expect [2v
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Nemophila insignis. 1870 Some plant on last page, in which the crossed selfs differed so greatly in development were allowed to seed excluded from insects. So self-plants, are self-fert for 2 generations. (Trained up sticks; greenhouse) [Table partly excised, not transcribed] All the plants grew very poorly in Pot 3 4 could hardly live died while young.— I cannot account for advantage of self-, as on both sides there was self-fertilisation — The case is a lesson of prudence — I suppose crossed
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CUL-DAR79.30-41
Note:
1868--1871
Escholtzia californica / Crossed 12 fl Self fert 18 flowers
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produced no pods spontaneously; but as plants were not healthy, nothing can be judged those in Greenhouse produced very few.─ 50
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CUL-DAR79.30-41
Note:
1868--1871
Escholtzia californica / Crossed 12 fl Self fert 18 flowers
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Escholtzia. July 5. 68 the growth of the tallest flower stems measured [Table excised] Plants in Pots in greenhouse 1868. Some Plants in the Pots were experimented on, so sterile 6 fl were crossed gave 3 pods, with moderate lot of seeds— 12 flower were self-fertilised by Lettington only 2 flow pods were spontaneously self-fertilised produced these contained very few seed — the best of 2 with only 14 seed. No pods were produced spontaneously.— The above plants on same pods were kept, after
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CUL-DAR79.30-41
Note:
1868--1871
Escholtzia californica / Crossed 12 fl Self fert 18 flowers
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The 19 plants in the 7 Pots crossed by garden plants produced (including 12 in paper of which seeds have to be counted) up to Aug 28, when all small great were gathered 240 Pods— The greenhouse not favourable to fertility.— The 18 intercrossed Plants produced 130 pods (so that the 19 wd have produced about 137 137.22 pods calculate this accurately) The 19 self plants produced 152 pods — so again the intercrossed show disadvantage. (The refuse seed, more or less germinated, was sown in 3
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F1742
Periodical contribution:
Darwin, C. R. 1868. On the character and hybrid-like nature of the offspring from the illegitimate unions of dimorphic and trimorphic plants. [Read 20 February] Journal of the Linnean Society of London (Botany) 10: 393-437.
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. Nevertheless these plants were not very sterile; for 25 flowers, fertilized with their own pollen, produced 15 capsules, containing an average of 16.3 seeds. As already stated, the probable average with legitimate plants for a union of this nature is rather above 20 seeds. But it should be observed that these plants were remarkably healthy and vigorous, being placed under highly favourable conditions, and grown in pots in the greenhouse; and we shall hereafter have occasion to show that such treatment
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.86 ∴ 127 would weigh 4.46 The 127 self- seeds weigh as 100 : 105 to self-fert, seeds. (May 17th in Pot. II. all the self plants, after se germinating seeds had been planted died. The seeds have all germinated badly.) (June 7th Pot III. is the only one with both sides alive the self flowered first) (July 13th Pot III. Plants have now done flowering crossed Pl. 28 1/2 inches 2 ft : 8 1/2 inches high (In Greenhouse) self Pl. 1 ft : 9 1/4 inch inch high 4
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CUL-DAR79.75-76
Note:
1868--1871
Broom / Pot I The three crossed plants very much healthiest and finest
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The Complete Work of Charles Darwin Online (63 A Broom 8 Plants on one side. (April 3. 68/ Pot I. The three crossed plants very much healthiest finest; tallest 3 1/4 high— tallest self- 3 inches. — in Pot II — The one crossed finer than the one self.— Four more young plants placed on each side of the same pot. ) so 5 on each side (June 14 68. the Pots were bedded out, unbroken, for plants began to look unhealthy in greenhouse; but thy, especially the self, suffered painfully from change. the
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F1744
Periodical contribution:
Darwin, C. R. 1868. On the specific difference between Primula veris, Brit. Fl. (var. officinalis, of Linn.), P. vulgaris, Brit. Fl. (var. acaulis, Linn.) and P. elatior, Jacq.; and on the hybrid nature of the common Oxlip. With supplementary remarks on naturally-produced Hybrids in the genus Verbascum. [Read 19 March] Journal of the Linnean Society of London (Botany) 10: 437-454.
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following facts may be worth giving:—Some Cowslips which had been transplanted from the fields into a shrubbery were again transplanted into highly manured land. In the following year they were protected from insects, artificially fertilized, and the seed thus procured was sown in a hotbed. The young plants were afterwards planted out, some in very rich soil, some in stiff poor clay, some in old peat, and some in pots in the greenhouse; so that these plants, 765 in number, as well as their parents
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F879.1
Book:
Darwin, C. R. [1868]. The variation of animals and plants under domestication. With a preface by Asa Gray. New York: Orange Judd and Co. vol. 1.
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without looking any more drawn than if it had stood in a common greenhouse; and Blanche Fleur seems as if made on purpose for growing in winter, like many bulbs, and to rest all summer. These odd constitutional peculiarities would fit a plant when growing in a state of nature for widely different circumstances and climates. Flowers possess little interest under our present point of view, because they have been almost exclusively attended to and selected for their beautiful colours, size
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F1742
Periodical contribution:
Darwin, C. R. 1868. On the character and hybrid-like nature of the offspring from the illegitimate unions of dimorphic and trimorphic plants. [Read 20 February] Journal of the Linnean Society of London (Botany) 10: 393-437.
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cultivated in the greenhouse the high average, from seven capsules, of 58.7 seeds, with a maximum in one capsule of 87 seeds; but from plants grown out of doors I never obtained a higher average than 41 seeds. Now two of the equal-styled plants, grown out of doors and spontaneously self-fertilized, gave averages of 44 and 45 seeds; but this high fertility may perhaps be attributed to the stigma receiving pollen from the surrounding anthers at exactly the right period. Two of these plants, fertilized
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F877.1
Book:
Darwin, C. R. 1868. The variation of animals and plants under domestication. London: John Murray. 1st ed., first issue. vol. 1.
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reproduced their proper leaves and fruit; but the seedlings were not quite equal in merit to their parents. The red-fleshed orange, on the other hand, fails to reproduce itself. Gallesio also observed that the seeds of several other singular varieties all reproduced trees having a peculiar physiognomy, but partly resembling their parent-forms. I can adduce another case: the myrtle-leaved orange is ranked by all authors as a variety, but is very distinct in general aspect: in my father's greenhouse
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F877.1
Book:
Darwin, C. R. 1868. The variation of animals and plants under domestication. London: John Murray. 1st ed., first issue. vol. 1.
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even pine-apple top and bottom heat, without looking any more drawn than if it had stood in a common greenhouse; and Blanche Fleur seems as if made on purpose for growing in winter, like many bulbs, and to rest all summer. These odd constitutional peculiarities would fit a plant when growing in a state of nature for widely different circumstances and climates. ——————————————— 166 Loudon's 'Arboretum et Fruticetum,' vol. ii. p. 834. 167 Loudon's 'Gardener's Mag.,' vol. ix. 1833, p. 123. 168 Ibid
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F877.2
Book:
Darwin, C. R. 1868. The variation of animals and plants under domestication. London: John Murray. 1st ed., first issue. vol. 2.
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fertile, though sterile when left to themselves, for bees are unable to crawl into the narrow tubular flower. The peloric flowers of Corydalis solida, according to Godron,91 are barren; whilst those of Gloxinia are well known to yield plenty of seed. In our greenhouse Pelargoniums, the central flower of the truss is often peloric, and Mr. Masters informs me that he tried in vain during several years to get seed from these flowers. I likewise made many vain attempts, but sometimes succeeded in
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F877.2
Book:
Darwin, C. R. 1868. The variation of animals and plants under domestication. London: John Murray. 1st ed., first issue. vol. 2.
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either pliability of organisation or some constitutional difference. The same individual cherry-tree, when forced, has been observed during successive years gradually to change its period of vegetation.75 Few pelargoniums can resist the heat of a stove, but Alba multiflora will, as a most skilful gardener asserts, stand pine-apple top and bottom heat the whole winter, without looking any more drawn than if it had stood in a common greenhouse; and Blanche Fleur seems as if it had been made on
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F878.1
Book:
Darwin, C. R. 1868. The variation of animals and plants under domestication. London: John Murray. 1st ed., second issue. vol. 1.
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reproduced their proper leaves and fruit; but the seedlings were not quite equal in merit to their parents. The red-fleshed orange, on the other hand, fails to reproduce itself. Gallesio also observed that the seeds of several other singular varieties all reproduced trees having a peculiar physiognomy, but partly resembling their parent-forms. I can adduce another case: the myrtle-leaved orange is ranked by all authors as a variety, but is very distinct in general aspect: in my father's greenhouse
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F878.1
Book:
Darwin, C. R. 1868. The variation of animals and plants under domestication. London: John Murray. 1st ed., second issue. vol. 1.
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even pine-apple top and bottom heat, without looking any more drawn than if it had stood in a common greenhouse; and Blanche Fleur seems as if made on purpose for growing in winter, like many bulbs, and to rest all summer. These odd constitutional peculiarities would fit a plant when growing in a state of nature for widely different circumstances and climates. 166 Loudon's 'Arboretum et Fruticetum,' vol. ii. p. 834. 167 Loudon's 'Gardener's Mag.,' vol. ix. 1833, p. 123. 168 Ibid, vol. xi. 1835, p
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F878.2
Book:
Darwin, C. R. 1868. The variation of animals and plants under domestication. London: John Murray. 1st ed., second issue. vol. 2.
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fertile, though sterile when left to themselves, for bees are unable to crawl into the narrow tubular flower. The peloric flowers of Corydalis solida, according to Godron,91 are barren; whilst those of Gloxinia are well known to yield plenty of seed. In our greenhouse Pelargoniums, the central flower of the truss is often peloric, and Mr. Masters informs me that he tried in vain during several years to get seed from these flowers. I likewise made many vain attempts, but sometimes succeeded in
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F878.2
Book:
Darwin, C. R. 1868. The variation of animals and plants under domestication. London: John Murray. 1st ed., second issue. vol. 2.
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either pliability of organisation or some constitutional difference. The same individual cherry-tree, when forced, has been observed during successive years gradually to change its period of vegetation.75 Few pelargoniums can resist the heat of a stove, but Alba multiflora will, as a most skilful gardener asserts, stand pine-apple top and bottom heat the whole winter, without looking any more drawn than if it had stood in a common greenhouse; and Blanche Fleur seems as if it had been made on
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F879.1
Book:
Darwin, C. R. [1868]. The variation of animals and plants under domestication. With a preface by Asa Gray. New York: Orange Judd and Co. vol. 1.
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another case; the myrtle-leaved orange is ranked by all authors as a variety, but is very distinct in general aspect: in my fathers greenhouse, during many years, it rarely yielded any seed, but at last produced one; and a tree thus raised was identical with the parent-form. Another and more serious difficulty in determining the rank of the several forms is that, according to Gallesio,16 they largely intercross without artificial aid; thus he positively states that seeds taken from lemon-trees (C
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CUL-DAR79.20-22
Note:
1868.05.24--1869.06.05
Bartonia aurea / 12 flowers crossed — 68 flowers self fert
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Bartonia aurea A fresh set of crosses 1868 6 flowers twice crossed by self-produced 4 pods these contained by eye rather more seed than the best of 7 pods spontaneously of the self-fertilised. — young flowers were crossed. self-fertilised 12 flowers were self-fertile by self produced only 7 pods as above. These plants in pots in greenhouse produced no pods spontaneously. (May 16— 1869 young plants about 1 inch high no difference) (May 28 1869 In Pot II III. a self-plant has died, but no other
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CUL-DAR79.158-161
Note:
1869
Phalaris (seed averages calculated) / Canary grass — treated just like
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Canary grass. — treated just like maize — ie. Art. crossed roughly spont. self- - fert— seeds (May 28 1869 — two rows of seeds were sown out of doors, equally thick, but wonderfully more of crossed have come up — so that crossed row looks quite thick self row now quite thin.) (June 6. 1869.— Greenhouse measured to extreme tips.) (Aug. 24th 1869 measured to tips of tallest culm in each pot.) [table not transcribed] [Pot 1-3 note:] All the crossed plants (in rather small pots in greenhouse
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CUL-DAR79.150-157
Note:
1869
Maize / Plants in greenhouse crossed artificially and singly — others
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The Complete Work of Charles Darwin Online (111 Maize Plants in greenhouse crossed artificially roughly— others spont. self-fert seed Put to germinate on sand. May 3d— 1869 June 6— 1869 Plants in Hothouse in Pots measured to extreme tips. 11
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CUL-DAR79.162-164
Note:
1869
Beet — seed from plant growing in group — & seed from solitary plant -
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The Complete Work of Charles Darwin Online (113 1869 Beet —seed from plant growing in group — seed from solitary plant— no additional aid. — (Jun 5 the row out of doors of self-plants has come up very badly L. [Lettington] turned up earth found that very many seed had germinated then died.) The seeds were sown on to 2 sides of 2 great Pots. Kept in Greenhouse in the 2 rows in open ground: the latter were protected during winter by a little straw, but many leaves were killed by the frost.— None
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CUL-DAR245.293
Correspondence:
Litchfield Henrietta Emma née Darwin to Darwin George Howard
[1869].04.19
Litchfield Henrietta Emma née Darwin to Darwin George Howard
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wd. be a pleasant companion dead then living. Horace will be here when u come home. Father hasn't been getting on or at least I don't kno' that he has until today when he certainly is more comfortable. He was driven up to the greenhouse on the truck was none the worse. The thing is the immense bruise wh. is partly inflamed - he thinks some nerve is crushed – so I am glad to say Paget is coming down. tho' Father is low at the thoughts thinks Paget will think him an [2v
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The Complete Work of Charles Darwin Online (50H Escholtzia. 1871. The From the plants on pots in Greenhouse (on last page) 12 pods were gathered from each lot; but from the self-fert. lot 7 out of the 12 pods contained all or almost all bad seeds. In the other 2 lots several of the pods were bad. So that there were not enough pods to give fair average of seed. The averages, such as they were, were very low; being for the self-fert. plants only 28.8 seeds. It is clear that greenhouse is very
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from the crossed-garden-plants, 4 were bad. Of the 35 from the intercrossed, 5 were bad. From the self plants 9 were bad. I carefully counted the seeds in 21 pods taken by chance from each of the 3 lots of 45 each. Of the crossed-garden-plants the average was — 67.00 Of the intercrossed average — 56.00 O Do Self-fert — 48.52 Hence we clearly see that the cross from new pollen in comparison with the intercrossed has increased, (not only the number of pods, judging from the greenhouse plants
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F2104
Periodical contribution:
Darwin, C. R. 1882. [Letters to Riley, 1871, 1881 and recollections of Darwin]. In Charles Valentine Riley, Darwin's work in entomology. Proceedings of the Biological Society of Washington DC 1: 70-80, pp. 77-80.
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In the brief hours I then spent at Down the proverbial modesty and singular simplicity and sweetness of his character were apparent, while the delight he manifested in stating facts of interest was excelled only by the eagerness with which he sought them from others, whether while strolling through the greenhouse or sitting round the generously spread table. Going to him as a young entomologist with no claim on his favor, he seemed to take delight in manifesting appreciation. I had occasion in
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F3505
Periodical contribution:
Darwin, C. R. 1884. [Words attributed to Darwin, 1871]. In J. D., A Reminiscence of Mr. Darwin. Harper's New Monthly Magazine 69, issue 413 (October): 759-63.
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invitation of Francis Darwin to go with him for a walk about the grounds, in the course of which we followed some of his father's favorite rambles along shaded paths in a neighboring field, coming back finally to the greenhouse, where some interesting experiments on the revolving movement of plants were at that time in progress. The work of the forenoon was the careful observation of a number of tender shoots that were growing in pots, each under a separate bell glass, and all ranged on a table exposed
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A2118
Periodical contribution:
F. W. B. 1871. Fertilisation of Leschenaultia formosa. Gardeners' Chronicle (26 August): 1103.
Text
The Complete Work of Charles Darwin Online [page] 1103 It was Darwin who first pointed out that self-fertilisation was injurious in the long run, and this important fact we continually see confirmed by weighty evidence, not in the vegetable kingdom alone, but also in the animal world. I happened the other day to examine rather closely certain flowers of that somewhat uncommon greenhouse plant, Leschenaultia formosa, and was agreeably surprised to find it furnished with everything needed in
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F3505
Periodical contribution:
Darwin, C. R. 1884. [Words attributed to Darwin, 1871]. In J. D., A Reminiscence of Mr. Darwin. Harper's New Monthly Magazine 69, issue 413 (October): 759-63.
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little further on were welcomed immediately by Mr. and Mrs. Darwin to a spacious and cheerful parlor or family room, whose broad windows and outer door opened upon a wide and partly sheltered piazza at the rear of the house, evidently a favorite sitting-place, judging from the comfortable look of easy-chairs assembled there, beyond which was a pleasing vista of fresh green lawn, bright flower beds, and blossoming shrubbery, gravel-paths, and a glass greenhouse, or perhaps botanical laboratory
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CUL-DAR55.69
Note:
1872.09.10
After rather cold night, plants in greenhouse at 8h 20 I routed several
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The Complete Work of Charles Darwin Online (p. 2A Sept 10th — After rather cold night, plant in greenhouse, at 8° 20' I routed several tent, 2 moved a little: I put on 3 bits of blue glass; 2 corks 2 cinders; all but the latter was carried inwards. I then put plant under Bell-glass with hot water raised temp to 75° 80˚; but when I routed several tentacles, repeatedly, not one moved 187
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(24) dipped in water at 56 slightly spotted, shaken for 1m. in water at 78° not very many near perfectly wetted. [slip of paper pasted on] (8) Acacia, I think certainly somewhat like my greenhouse one ─ upper surface much wetted by cold water; other surface less─ whole leaf edges absolutely vertical. [slip of paper pasted on] (2 (5) — ? both sides wetted cold water edges of leaves very vertical. (6) — — upper side well wetted lower side considerably— cold water. edges of leaf vertical — (
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CUL-DAR209.12.85
Note:
[Undated]
Cassia bicapsularis / Cassia schinifolia / Cassia barclayana
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The Complete Work of Charles Darwin Online [85] Cassia bicapsularis (Kew) Ind. Or. occ. Madera like greenhouse var.— when shaken leaves become inclined a little— sleep in normal fashion.— 27 [-] 8 [=] 19° 67 [-] 33 [=] 34° Cassia Schinifolia = C. Barclayana. new. Holland no bloom on narrow the rigid leaves. At noon the many leaflets on opposite side of petiole lies in nearly a horizontal plane middle ones formed angle together of 104°. [sketch] At night they rise a little towards each other
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The Complete Work of Charles Darwin Online [24] Cassia floribunda old Greenhouse var. (I have got name somewhere A cut down bush had thrown up vigorous shoots — a young leaf selected — bearing 4 pairs of leaflets, main petioles 3 3/4 long — filament affixed to one of terminal leaflets, petiole secured to stick close behind joint apex 5 1/2 from vertical glass.— This leaflet 1 10/12 5/6 long— [sketch] Observed under skylight traced on vertical glass, from midday of 13th to morning of 17th— see
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CUL-DAR209.3.79
Note:
[Undated]
Red cabbage / Proof sheet of Forms of flowers, published pp. 182-3.
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in the stem leaf how much in the stem. ) probably in chief part to the latter.) Lastly a full-grown cabbage had been kept for some time in a pot in the greenhouse, so that the head had become a little elongated with the leaves not close together: a long glass filament was fixed to the mid-rib of an almost central a leaf was 5 1/2 inches in length near the centre which was inclined at an angle of 44° above horizon, The movement obs was traced on a vertical glass. The plant was kept in complete
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cabbage which had formed Head same plant was left in the greenhouse until its head had become somewhat elongated (Nov. 10th) with the leaves not so closing pressed together; a filament was so affixed to a leaf near to centre, 5 1/2 inches in length, which stood ou t projected at 44° above the horizon Its movement was traced on a vertical glass, The plant being was kept in complete darkness, except for about a minute or two when being observed with every hour by aid of a tape. The movements were
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The Complete Work of Charles Darwin Online (1 Caladium. esculentum (a common Greenhouse Plant) (sleeps) May be said to sleep Caladium esculentum vel Colocasia antiquorum Schott— this is first Proper name A young leaf only 4 inches in length from apex to forked base, with petiole only 3 1/4 high was observed under skylight glass-filament attached to forked brim traced on [sketch] vertical glass. Apex of forked base 11 inches from glass. When tracing rises the apex of leaf falls.— (Temp on 22d
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CUL-DAR55.33-68
Note:
1873.06.00--1873.07.00
[Drosera continued] [application of olive oil, atropine, valerianate of
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July 25 8˚ 30' — Cheese in nearly same state: of the 3 cube of albumen, 1 still square white though reduced; the 2d sub-rounded, but still with much white; the 3d with more trace of opake white in the centre. — The albumen was placed on moss on the 17th weather very hot,— kept in greenhouse.) ((Casein flocculant white rounded at base of cup.) Control (4
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CUL-DAR111.A43
Note:
1873.06.20
Cleistogene Flowers / Drosera rotundifolia kept in cool — Hot-House -
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June 25. D. Anglica produces in greenhouse rather more perfect fl. showing more petals, aperture just open tempering, but hardly enough from insect to visit it — Pollen in shed, but remain round anthers in a cross penetrate the stigma
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35 Introduction were exposed to slightly different conditions as the seasons varied, and as they were raised at different periods. But in other respects all were treated alike, being grown in pots, in the same artificially prepared soil, watched being watched at the same time, and kept in the same greenhouse or hothouse. They were therefore not exposed during successive years to such great vicissitudes of climate as are plants growing out of doors. (open) On some apparent and real causes of
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my study, which was much cooler than greenhouse, all soon closed, shows that it is temperature in accordance with Pfeiffer. It is, however, odd that flower before with water at 98° closed— was it due to evaporation? Sunshine is not necessary for opening of flowers, for they opened on floor in morning when sun did not reach them. — I cd introduce this as * to show caution necessary about water not being the active agent
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dropped off. The leaflet of above leaf, of which tip had been immersed, are now still sensitive though only to slight degree.— (12° in greenhouse I can now see that leaves are dry that they are killed.) (over) [46v
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CUL-DAR209.12.89-119
Draft:
1873.10.07--1873.12.12
Desmodium gyrans / Draft of Expression (fragment).
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in the least affected by the cold water. The greenhouse Cassia from cool hothouse put left in the cold water for 3h 10', had its leaflets a little beveled, which is the first movement in going to sleep. Cytisus [few words illeg]— Trifolium repens in greenhouse— Passiflora gracilis (which depresses leaves when asleep) Tobacco, (growing at out of doors, which slightly raises leaves) were none of them in the least affected.) Sensitivity See Back [29v
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wonder whether Barbary stamens sleep?) — but 2 flowers which remained open at night had their stamens sensitive p. 20 in warm greenhouse, he perceived frenissment spontané dans les etamines — (I might allude to this Cereus under circumnutation of Flowers.) p. 20 Stamen when touched or even blown move rapidly for pistils pretty quickly back from it— p. 23 excitement transmitted from one stamen to other
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CUL-DAR66.26-27
Note:
1873.10.24--1873.10.26
Oxalis acetosella & sensitiva / Potted kept in cool Greenhouse I observed
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The Complete Work of Charles Darwin Online [26] Oct 24th /73/ Oxalis acetosella— sensitiva Potted kept in cool greenhouse. This morning I observed yesterday during very heavy rain that plant out of doors had leaves depressed, but as I have proved that cold thus acts I did not know whether it was beating of water or cold that acted; so this morning I syringed strongly the above plants with water at 72°; in a few minutes almost every leaf young old had 3 leaflets depressed on an average about 45
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CUL-DAR66.26-27
Note:
1873.10.24--1873.10.26
Oxalis acetosella & sensitiva / Potted kept in cool Greenhouse I observed
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moved. I then syringed with water at 62° this produced far more effect, on the young leaves the leaflets shut pretty well. — Water certainly seems to produce more effect than shaking or blowing. — All the leaves young old were well spotted with water after syringing O. acetosella. (Oct 25' 12°) plant had stood in greenhouse, temp. 55° I blew on it violently with bellows for 1 1/2m, on half the plants, a little but a little effect was produced — I then syringed gently the same half of pot for short
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CUL-DAR66.26-27
Note:
1873.10.24--1873.10.26
Oxalis acetosella & sensitiva / Potted kept in cool Greenhouse I observed
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Oxalis acetosella Oct 26 /73/7° 30' Am Plants in greenhouse at temp of 50° put 3 leaves in water of same temp. after 20' only trace of depression of leaflets— Put at same time other 3 leaves in water at 71°, after 20; only trace of depression. — I brushed several times lightly with my open hand the leaves of all the plants, after 3' perhaps a trace of effect after 20' a slight depression of som most of them I then syringed strongly the plants on one side of large pot with water at 62° the
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CUL-DAR77.44
Note:
1873.10.25
Amongst the Convolvulus plants experimented on in last year (72) there
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average only 3.20 seeds per pod (two pods having 6 seeds) this I think much below average. There was a second plant, raised from cross by pollen from another flower on the same plant, which was somewhat less sterile a few seeds were sown in same pot in greenhouse as above, the 24 pods as above yielded average of 3.96 seeds per pod (with 3 containing 6). So that (put in note) transmitted their degrees of sterility to their offspring— (More pods were produced than the 24 gathered off each lot
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CUL-DAR209.12.45-49
Note:
1873.11.00--1873.12.00
Cassia [application of water, sulphuric ether]
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. 2d. at 12° 1° brought plant from Greenhouse shook it intentionally during journey for 2m.; leaflets were depressed at the end of journey.— Before starting I measured angle of main petiole of younger leaf (red thread) with the perpendicular it was 73° so was after the Journey.— A second leaf white wool older (third leaf on axis on stem) had angle before journey of 62 1/2° after journey of 71°, so it had fallen become depressed 8° 1/2; which is contrary movement to sleep movement of main petiole
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CUL-DAR209.12.141-147
Note:
1873.11.07--1873.12.05
Eucalyptus globulus / Eucalyptus amygdalina [application of water, ether]
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Eucalyptus globulus (young plant with horizontal leaves) Dec' 5- 9° — Washed a leaf rubbed with soft camel with water at 90° so got off bloom put on large drop of water.— (Washed leaf to left at tip did not leave on water.) Removed water on 7th 12° 10'— Greenhouse Cool— Hot House little warmer than Greenhouse— Evaporated water, I think more matter than can be accounted for by dust during 52°.— (Weather dull during whole 52°) time cloudy). (Dec. 9. 3° P.m. no effect produced) Dec. 9th 8° A.m
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CUL-DAR209.12.141-147
Note:
1873.11.07--1873.12.05
Eucalyptus globulus / Eucalyptus amygdalina [application of water, ether]
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water.— Removed water on Dec. 7th 12° 10'; evaporated it, I think more matter than can be accounted for by dust during the 52°.— (Weather dark all the time) (Plant in cool Hothouse, not much warmer than greenhouse.) Dec. 9th no effect, but then weather whole time cloudy
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The Complete Work of Charles Darwin Online [80] Common Cabbage (Movement of leaf of full-grown plant in darkness) Nov. 10th A large plant in pot in greenhouse of which I measured distance of opposite leaves some little time ago, had now run run up a little so that head not so no longer compact. Chose One one of young leaves on the summit, (within which several smaller leaves,) 5 1/2 inches in length, inclined 44° above horizon. I fixed long glass filament to mid-rib with little mark with black
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shown.— Plant was moved 2 days previously previously from greenhouse to hot-House to accelerate growth.— (Dyer) Cereus speciosissimus. (garden var) (sometimes Phyllocactus multiflora
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The Complete Work of Charles Darwin Online [15] (12 Cassia Dec. 24'. Brought plant from warm greenhouse into study yesterday at midday today (Temp of Study 54°-56°, perhaps too cold for spont., movements) observed movements of tips of leaf on glass-plate. Began at 8° A.m, when just light. Leaf in nearly the same position to light as on previous days — Day rather clear. —Two leaves observed. First terminal leaflet rather torpid leaf pointed almost directly from light, yet slightly there was
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The Complete Work of Charles Darwin Online [162] Robinia pseudo-acacia 1874 April 4th. Leaf expanded in greenhouse not very healthy — Syringed violently, terminal leaflets pretty well protected almost dry after shaking them, but some which looked unhealthy very wet.— I think younger leaflets certainly are depressed from the syringing the whole main petiole, I certainly think depressed. (Look at Elizabeth tree during rain.) — 9°. 45' P.m. H. F. brought me a leaf beautifully asleep, with all
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The Complete Work of Charles Darwin Online [76] ap. 30 /74/ 10° 50' Mimulus Luteus Greenhouse Plants Syringed hard on stigma of 7 flowers none shut when afterwards touched with needle blotting Paper did not shut! can cold water of Temp. of room paralyze them? Temp. ─ 56° One did shut another partially so I tried 9 flowers. (At 11° 25' (ie after only 35m) 6 of these flowers shut when touch with fine needle for human hair is not stimulus enough. (12° most of them open again) (2° 30' touched the
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The Complete Work of Charles Darwin Online [50] Cassia corymbosa— Greenhouse May 1. 74. (12°) syringed with Hot-house water for 1m leaves became reflexed inverted, almost as if asleep, but did not touch as when (as I believe) asleep. The white (blooms?) surface is turned to stream of water. I rather suspect peduncle became depressed. did not soon recover open leaves. (May 2d I carried plant to Hothouse shook it slightly all the way, leaflets become a little depressed but nothing like what
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CUL-DAR66.34
Note:
1874.05.01--1874.05.02
Oxalis (Greenhouse/grown out of doors) with large leaves & concentric
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The Complete Work of Charles Darwin Online [34] June May 1. 74 Oxalis (Greenhouse/ grown out of doors) With large leaves concentric purple mark on leaves— flower pale purple.— syringed leaflets closed closed to certain extent, like O. acetosella. — May 2d Oxalis with Coppery foliage yellow flowers— I syringed 2 lots of plants in Hot House one lot in pot showed slight signs of depression of leaflets. The other showed no signs.— I then put pot outside of H. House — rather cold day leaves after 3
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CUL-DAR57.18
Note:
1874.05.07--1874.05.15
8 a.m / Six sticks with Black wisp / atoms of old albumen moistened with
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tried were acid May 13' 8° 10. dusted with blue glass, particles of, first ascertained though without much care not acid. 8 or 9 leaves, in 10' much inflection. (Given below as caution) I have no doubt the explanation of this apparent anomaly, that the 8 cinders though not small did not act at all when placed on disc the 8 bits (rather large) of glass did acted very slightly only on few leaves, is due to cold weather the plates having been taken out very warm greenhouse — give this as caution
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The Complete Work of Charles Darwin Online [51] May 25' 1874— I placed Greenhouse Cassia little sp. which has been photographed out in rather heavy warm rain no wind— the leaves were hardly at all depressed not at all inverted.— I then shook plant a little there was some inversion, so that it seems to be movement not drops of water which causes movement
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CUL-DAR59.1.157
Note:
[1874].07.18--[1874].07.22
Utricularia / Fed w[ith] cartilage 6.30 p.m
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The Complete Work of Charles Darwin Online [157] Utricularia July 18. Fed w cartilage 6.30 PM July 20 9.30 AM one bit has [illeg] slipped right in— In another of the pieces I could see that the glands were not inflected so as to touch it— 3 bit of Cartilage Raw Meat July 18— Put into a dish with 2 or 3 drops of raw meat infusion— (high) filled up with tank water— July 20— 9.30 AM— Globular glands opaque yellow much shrunk— — July 20 Examined specimens straight from GreenHouse found shrinking
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F836
Book:
Darwin, C. R. 1875. The movements and habits of climbing plants. 2d ed. London: John Murray.
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, cut off a plant, and placed in water, made circles in 3 hrs. 10 m., 5 hrs. 20 m., and 7 hrs. 15 m. Wistaria Chinensis (Leguminosæ), in greenhouse, moves against the sun. H. M. May 13, 1st circle was made in . . 3 5 ,, 13, 2nd ,, ,, ,, . . 3 20 ,, 16, 3rd ,, ,, ,, . . 2 5 ,, 24, 4th ,, ,, ,, . . 3 21 ,, 25, 5th ,, ,, ,, . . 2 37 ,, 25, 6th ,, ,, ,, . . 2 35 Phaseolus vulgaris (Leguminosæ), in greenhouse, moves against the sun. H. M. May, 1st circle was made in . . 2 0 ,, 2nd ,, ,, ,, . 1 55
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CUL-DAR110.A12r
Draft:
[Undated]
Draft of `Cross and self fertilisation'?: 246 [top righthand quarter only]
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The Complete Work of Charles Darwin Online (246 [Left side and bottom of page excised] (246 (Primula) was first fertilised by pollen taken from ong-styled forms; the long-styled union again fertilised with The grandchildren thus raised, were carefully sel cultivated st in the greenhouse were very but when turned out of doors were became considerably sterile. The of fertilisation was continued from generations; the gr. gr. grandchildren to detract will be [illeg] given as long as they were
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F836
Book:
Darwin, C. R. 1875. The movements and habits of climbing plants. 2d ed. London: John Murray.
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(MONOCOTYLEDONS, continued.) Asparagus (unnamed species from Kew) (Liliaceæ) moves against the sun, placed in hothouse. H. M. Dec. 26, 1st circle was made in . . 5 0 ,, 27, 2nd ,, ,, ,, . . 5 40 Tamus communis (Dioscoreaceæ). A young shoot from a tuber in a pot placed in the greenhouse: follows the sun. H. M. July, 7, 1st circle was made in . . 3 10 ,, 7, 2nd ,, ,, ,, . . 2 38 ,, 8, 3rd ,, ,, ,, . . 3 5 ,, 8, 4th ,, ,, ,, . . 2 56 ,, 8, 5th ,, ,, ,, . . 2 30 ,, 8, 6th ,, ,, ,, . . 2 30
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CUL-DAR91.95r-98r
Draft:
[Undated]
Drafts of Cross and self fertilisation, sheets 191-194, published pp. 101-103
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Brassica but several grew extremely crooked, from having been drawn up to the light whilst in the greenhouse. As it was scarcely possible to measure their heights, the finest plant on each side of each pot was cut down close to the ground and weighted. In the following Table we have the result. Table 30. Cabbages Weight of whole plants after they had formed heads. No. of Pot Crossed plants from pollen of fresh stock Self-fertd: plants of the 3rd generation I 130 oz 18 2/4 oz II 74 34 3/4 III
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F836
Book:
Darwin, C. R. 1875. The movements and habits of climbing plants. 2d ed. London: John Murray.
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weighing only .82 of a grain (53.14 mg.) We have seen that the petioles of some other leaf-climbing plants are affected by one-thirteenth of this latter weight. In this species, and in no other leaf-climber seen by me, a full-grown leaf is capable of clasping a stick; but in the greenhouse the movement was so extraordinarily slow that the act required several weeks; on each succeeding week it was clear that the petiole had become more and more curved, until at last it firmly clasped the stick
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F836
Book:
Darwin, C. R. 1875. The movements and habits of climbing plants. 2d ed. London: John Murray.
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Shoots of the Kidney-bean, cut off and placed in water, were similarly retarded, but in a less degree. I have repeatedly observed that carrying a plant from the greenhouse to my room, or from one part to another of the greenhouse, always stopped the movement for a time; hence I conclude that plants in a state of nature and growing in exposed situations, would not make their revolutions during very stormy weather. A decrease in temperature always caused a considerable retardation in the rate of
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F836
Book:
Darwin, C. R. 1875. The movements and habits of climbing plants. 2d ed. London: John Murray.
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of 3 hrs. 26 m. The shoots, however, sometimes stood still. It is considered a greenhouse plant; but when kept there, the petioles took several days to clasp a stick: in the hothouse a stick was clasped in 7 hrs. In the greenhouse a petiole was not affected by a loop of string, suspended during several days and weighing 2½ grains (163 mg.); but in the hothouse one was made to curve by a loop weighing 1.64 gr. (106.27 mg.); and, on the removal of the string, it became straight again. Another
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amount of inflection is utterly insignificant, as we shall hereafter see, compared with that caused by very weak solutions of several salts of ammonia. Plants which have lived for some time in a rather high temperature are far more sensitive to the action of water than those grown out of doors, or recently brought into a warm greenhouse. Thus in the above seventeen cases, in which the immersed leaves had a considerable number of tentacles inflected, the plants had been kept during the winter
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amount of inflection is utterly insignificant, as we shall hereafter see, compared with that caused by very weak solutions of several salts of ammonia. Plants which have lived for some time in a rather high temperature are far more sensitive to the action of water than those grown out of doors, or recently brought into a warm greenhouse. Thus in the above seventeen cases, in which the immersed leaves had a considerable number of tentacles inflected, the plants had been kept during the winter
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.* It is also anemophilous, that is, or it is fertilised by means of the wind; and of such plants only the common beet had alone been tried. Some plants of maize were grown raised in the greenhouse, and were crossed with pollen taken from a distinct plant; and a single plant, growing quite separately in a different part of the house, was allowed to fertilise itself spontaneously. The seeds thus obtained were placed on damp sand, and as they germinated were planted in pairs of equal age on the
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CUL-DAR91.95r-98r
Draft:
[Undated]
Drafts of Cross and self fertilisation, sheets 191-194, published pp. 101-103
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cool greenhouse. Early in January their heights were measured to the tips of their leaves. The thirteen crossed plants averaged 13.16 inches in height, and the twelve (for one had died) self-fertilised plants averaged 13.7 inches; or as 100 to 104. So that the taller self-fertilised plants exceeded by a little the crossed plants. Early in the spring the plants were gradually hardened off and turned out of their pots into the open ground without being disturbed. By the end of August the greater
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The bladders are filled with water. They generally, but by no means always, contain bubbles of air. According to the quantity of the contained water and air, they vary much in thickness, but are always somewhat compressed. At an early stage of growth, the flat or ventral surface faces the axis or stem; but the footstalks must have some power of movement; for in plants kept in my greenhouse the ventral surface was generally turned either straight or obliquely downwards. The Rev. H.M. Wilkinson
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The bladders are filled with water. They generally, but by no means always, contain bubbles of air. According to the quantity of the contained water and air, they vary much in thickness, but are always somewhat compressed. At an early stage of growth, the flat or ventral surface faces the axis or stem; but the footstalks must have some power of movement; for in plants kept in my greenhouse the ventral surface was generally turned either straight or obliquely downwards. The Rev. H.M. Wilkinson
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in my greenhouse, that the odour is attractive. In this latter case the leaves may be compared with a baited trap; in the former case with a trap laid in a run frequented by game, but without any bait. That the glands possess the power of absorption, is shown by their almost instantaneously becoming dark-coloured when given a minute quantity of carbonate of ammonia; the change of colour being chiefly or exclusively due to the rapid aggregation of their contents. When certain other fluids are
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, extremely tough cartilage were cut from the end of a slightly roasted leg-bone of a sheep. These were placed on three leaves, borne by poor, small plants in my greenhouse during November; and it seemed in the highest degree improbable that so hard a substance would be digested under such unfavourable circumstances. Nevertheless, after 48 hrs., the cubes were largely dissolved and converted into minute spheres, surrounded by transparent, very acid fluid. Two of these spheres were completely
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weather. Those with the weakest solutions should be made on plants which have been kept for a considerable time in a warm greenhouse, or cool hothouse; but this is by no means necessary for trials with solutions of moderate strength. I beg the reader to observe that the sensitiveness or irritability of the tentacles was ascertained by three different methods-indirectly by drops placed on the disc, directly by drops applied to the glands of the outer tentacles, and by the immersion of whole leaves; and
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in my greenhouse, that the odour is attractive. In this latter case the leaves may be compared with a baited trap; in the former case with a trap laid in a run frequented by game, but without any bait. That the glands possess the power of absorption, is shown by their almost instantaneously becoming dark-coloured when given a minute quantity of carbonate of ammonia; the change of colour being chiefly or exclusively due to the rapid aggregation of their contents. When certain other fluids are
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, extremely tough cartilage were cut from the end of a slightly roasted leg-bone of a sheep. These were placed on three leaves, borne by poor, small plants in my greenhouse during November; and it seemed in the highest degree improbable that so hard a substance would be digested under such unfavourable circumstances. Nevertheless, after 48 hrs., the cubes were largely dissolved and converted into minute spheres, surrounded by transparent, very acid fluid. Two of these spheres were completely
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weather. Those with the weakest solutions should be made on plants which have been kept for a considerable time in a warm greenhouse, or cool hothouse; but this is by no means necessary for trials with solutions of moderate strength. I beg the reader to observe that the sensitiveness or irritability of the tentacles was ascertained by three different methods-indirectly by drops placed on the disc, directly by drops applied to the glands of the outer tentacles, and by the immersion of whole leaves; and
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F2111
Book contribution:
Darwin, C. R. 1896. [Recollections of Darwin and correspondence with Romanes, 1875-1881]. In E. D. Romanes ed., The life and letters of George John Romanes. 6th impression. London: Longmans, 1908.
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so young a plant by putting it in a greenhouse. Perhaps, therefore, you might pot it, as soon as it arrives, and keep it till I go up. If you do not care to take charge of it altogether, I can then get a home for it somewhere in the South. It will not require a deep pot, for I see that I have cut through the end of one of the roots. It would be as well, before potting, to cut off the end of the other root also, so that the one half may not grow longer than the other, and thus perhaps assert an
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F836
Book:
Darwin, C. R. 1875. The movements and habits of climbing plants. 2d ed. London: John Murray.
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only one-eighth of a grain (8.1 mg.) sometimes acted and sometimes did not act. The sensitiveness extends from the blade of the leaf to the stem. I may here state that I ascertained in all cases the weights of the string and thread used by carefully weighing 50 inches in a chemical balance, and then cutting off measured lengths. The main petiole carries three leaflets; but their short, sub-petioles are not sensitive. A young, inclined shoot (the plant being in the greenhouse) made a large circle
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not reopen until nearly two days had passed, and was not even then in the least sensitive. After an additional day it recovered its powers, and closed on being touched and subsequently reopened. Another leaf behaved in nearly the same manner after a shorter exposure to this vapour. On the Manner in which Insects are caught. We will now consider the action of the leaves when insects happen to touch one of the sensitive filaments. This often occurred in my greenhouse, but I do not know whether
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not reopen until nearly two days had passed, and was not even then in the least sensitive. After an additional day it recovered its powers, and closed on being touched and subsequently reopened. Another leaf behaved in nearly the same manner after a shorter exposure to this vapour. On the Manner in which Insects are caught. We will now consider the action of the leaves when insects happen to touch one of the sensitive filaments. This often occurred in my greenhouse, but I do not know whether
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eight pairs of fine leaves, and the weather was favourable; the temperature of the room where the leaves were immersed varying from 75° to 81° (23°.8 to 27°.2 Cent.) In another trial with four pairs (included in the above twenty pairs), the temperature in my room was rather low, about 60° (15°.5 Cent.); but the plants had been kept for several days in a very warm greenhouse and thus rendered extremely sensitive. Special precautions were taken for this set of experiments; a chemist weighed for me a
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those placed in the solutions and in water, were taken from plants which had been kept in a very warm greenhouse during the winter. They were thus rendered extremely sensitive, as was shown by water exciting them much more than in the previous experiments. Before giving my observations, it may be well to remind the reader that, judging from thirty-one fine leaves, the average number of tentacles is 192, and that the outer or exterior ones, the movements of which are alone significant, are to the
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in amount; but one such mass retained exactly the same form as before after an interval of 5 hrs., so that it could hardly have consisted of living protoplasm. These glands seem to have very little or no power of absorption, certainly much less than those of the foregoing plants. Mirabilis longiflora. The stems and both surfaces of the leaves bear viscid hairs. young plants, from 12 to 18 inches in height in my greenhouse, caught so many minute Diptera, Coleoptera, and larvae, that they were
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eight pairs of fine leaves, and the weather was favourable; the temperature of the room where the leaves were immersed varying from 75° to 81° (23°.8 to 27°.2 Cent.) In another trial with four pairs (included in the above twenty pairs), the temperature in my room was rather low, about 60° (15°.5 Cent.); but the plants had been kept for several days in a very warm greenhouse and thus rendered extremely sensitive. Special precautions were taken for this set of experiments; a chemist weighed for me a
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those placed in the solutions and in water, were taken from plants which had been kept in a very warm greenhouse during the winter. They were thus rendered extremely sensitive, as was shown by water exciting them much more than in the previous experiments. Before giving my observations, it may be well to remind the reader that, judging from thirty-one fine leaves, the average number of tentacles is 192, and that the outer or exterior ones, the movements of which are alone significant, are to the
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in amount; but one such mass retained exactly the same form as before after an interval of 5 hrs., so that it could hardly have consisted of living protoplasm. These glands seem to have very little or no power of absorption, certainly much less than those of the foregoing plants. Mirabilis longiflora. The stems and both surfaces of the leaves bear viscid hairs. young plants, from 12 to 18 inches in height in my greenhouse, caught so many minute Diptera, Coleoptera, and larvae, that they were
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F880.1
Book:
Darwin, C. R. 1875. The variation of animals and plants under domestication. London: John Murray. 2d ed. vol. 1.
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the seedlings were not quite equal in merit to their parents. The red-fleshed orange, on the other hand, fails to reproduce itself. Gallesio also observed that the seeds of several other singular varieties all reproduced trees having a peculiar physiognomy, partly resembling their parent-forms. I can adduce another case: the myrtle leaved orange is ranked by all authors as a variety, but is very distinct in general aspect: in my father's greenhouse, during many years, it rarely yielded any fruit
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F880.1
Book:
Darwin, C. R. 1875. The variation of animals and plants under domestication. London: John Murray. 2d ed. vol. 1.
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common greenhouse; and Blanche Fleur seems as if made on purpose for growing in winter, like many bulbs, and to rest all summer. These odd constitutional peculiarities would enable a plant in a state of nature to become adapted to widely different circumstances and climates. Flowers possess little interest under our present point of view, because they have been almost exclusively attended to and selected for their beautiful colour, size, perfect outline, and manner of growth. In these particulars
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F880.2
Book:
Darwin, C. R. 1875. The variation of animals and plants under domestication. London: John Murray. 2d ed. vol. 2.
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, according to Godron,95 are sometimes barren and sometimes fertile; whilst those of Gloxinia are well known to yield plenty of seed. In our greenhouse Pelargoniums, the central flower of the truss is often peloric, and Mr. Masters informs me that he tried in vain during several years to get seed from these flowers. I likewise made many vain attempts, but sometimes succeeded in fertilising them with pollen from a normal ————————————————— 89 'Bastarderzeugung,' s. 356. 90 'Teoria della Riproduzione
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F880.2
Book:
Darwin, C. R. 1875. The variation of animals and plants under domestication. London: John Murray. 2d ed. vol. 2.
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this shows either pliability of organisation or some constitutional difference. The same individual cherry-tree, when forced, has been observed during successive years gradually to change its period of vegetation.75 Few pelargoniums can resist the heat of a stove, but Alba Multiflora will, as a most skilful gardener asserts, stand pine-apple top and bottom heat the whole winter; without looking any more drawn than if it had stood in a common greenhouse; and Blanche Fleur seems as if it had been
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F1249
Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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, left uncovered in greenhouse, produced capsules in number(about) as 100 to 100 LUPINUS LUTEUS crossed and self-fertilised plants of the 2nd generation, left uncovered in the greenhouse, produced seeds in number (judged from only a few pods) as 100 to 88 PHASEOLUS MULTIFLORUS crossed and self-fertilised plants, left uncovered in greenhouse, produced seeds in number(about) as 100 to 100 LATHYRUS ODORATUS crossed and self-fertilised plants of the 2nd generation, left uncovered in the greenhouse
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F1249
Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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the cases which follow that this result was accidental, owing to only a few plants having been measured, and to one of the self-fertilised plants having grown only to a height of 15 inches. The plants had been kept in the greenhouse, and from being drawn up to the light had to be tied to sticks in this and the following trials. They were measured to the summits of their flower-stems. The four crossed plants here average 29.68 inches, and the four self-fertilised 25.56 in height; or as 100 to
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F1249
Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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occasions with plants of the first generation of Nicotiana. The crossed plants always withstood the injurious effects of being suddenly removed into the open air after having been kept in the greenhouse better than did the self-fertilised. On several occasions they also resisted much better cold and intemperate weather. This was manifestly the case with some crossed and self-fertilised plants of Ipomoea, which were suddenly moved from the hothouse to the coldest part of a cool greenhouse. The
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F1249
Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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hemisphere, with a complete reversal of the seasons, were thus rendered slightly self-fertile, whereas they seem always to be completely self-sterile in their native home. Senecio cruentus (greenhouse varieties, commonly called Cinerarias, probably derived from several fruticose or herbaceous species much intercrossed*). Two purple-flowered varieties were placed under a net in the greenhouse, and four corymbs on each were repeatedly brushed with flowers from the other plant, so that their
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F1249
Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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flowers; as they crawled into them, they rubbed against the bristles which project from the anthers, and became dusted with pollen. Veronica agrestis (Scrophulariaceae). Covered-up plants produced an abundance of seeds. I do not know whether any insects visit the flowers; but I have observed Syrphidae repeatedly covered with pollen visiting the flowers of V. hederaefolia and chamoedrys. Mimulus luteus (Scrophulariaceae). Highly self-fertile. Calceolaria (greenhouse variety) (Scrophulariaceae
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F1249
Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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three long rows, however, were not of quite equal lengths, and the plants were much crowded, so that it would have been extremely difficult to have ascertained how many capsules were produced by them, even if I had been willing to undertake so laborious a task as to collect and count all the capsules. But this was feasible with the plants grown in pots in the greenhouse; and although these were much less fertile than those growing out of doors, their relative fertility appeared, after
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F1249
Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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maturity only a pair of plants, which were kept in a pot in the greenhouse. The crossed plant grew to a height of 33 inches, and the self-fertilised to that of 26 1/2 inches. The former produced, whilst still kept in the greenhouse, eight pods, containing on an average 2.77 seeds; and the latter only two pods, containing on an average 2.5 seeds. The average height of the two crossed plants of the two [page] 15
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F1249
Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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The self-fertilised peas were, however, quite as heavy as those from the crossed pods. From these two lots of seeds, the plants of the next generation were raised. Plants of the Second Generation. Many of the self-fertilised peas just referred to germinated on sand before any of the crossed ones, and were rejected. As soon as I got equal pairs, they were planted on the opposite sides of two large pots, which were kept in the greenhouse. The seedlings thus raised were the grandchildren of the
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F1249
Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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conditions in pots in the greenhouse. As long as they were cultivated in this manner, they grew well and were healthy and fertile. Their fertility even increased in the later generations, as if they were becoming habituated to illegitimate fertilisation. Plants of the first illegitimate generation when taken from the greenhouse and planted in moderately good soil out of doors grew well and were healthy; but when those of the two last illegitimate generations were thus treated they became exces- * See
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F1249
Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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; and to the summits of their male flowers, 53.96 and 43.45 inches; or as 100 to 80. Phalaris canariensis. Hildebrand has shown in the paper referred to under the last species, that this hermaphrodite grass is better adapted for cross-fertilisation than for self-fertilisation. Several plants were raised in the greenhouse close together, and their flowers were mutually intercrossed. Pollen from a single plant growing quite separately was collected and placed on the stigmas of the same plant. The
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F1249
Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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TABLE D.—continued. PETUNIA VIOLACEA left uncovered as above: offspring of plants self-fertilised for four generations and then crossed by a fresh stock, compared with plants of the 5th self- fertilised generation, produced seeds, as judged by the weight of an equal number of capsules as 100 to 46 CYCLAMEN PERSICUM crossed and self-fertilised plants, left uncovered in the greenhouse, produced capsules in number as 100 to 12 ANAGALLIS COLLINA crossed and self-fertilised plants, left uncovered
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F1249
Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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vigour to the crossed. My attention was now thoroughly aroused, for I could hardly doubt that the difference between the two beds was due to the one set being the offspring of crossed, and the other of self-fertilised flowers. Accordingly I selected almost by hazard two other plants, which happened to be in flower in the greenhouse, namely, [page] 1
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F1249
Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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. In the successive generations they were exposed to slightly different conditions as the seasons varied, and they were raised at different periods. But in other respects all were treated alike, being grown in pots in the same artificially prepared soil, being watered at the same time, and kept close together in the same greenhouse or hothouse. They were [page] 2
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F1249
Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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crossed and self-fertilised generations Small amount of pollen in the anthers of the self-fertilised plants of the later generations, and the sterility of their first produced flowers Uniform colour of the flowers produced by the self-fertilised plants The advantage from a cross between two distinct plants depends on their differing in constitution. A PLANT of Ipomoea purpurea, or as it is often called in England the convolvulus major, a native of South America, grew in my greenhouse. Ten flowers on
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F1249
Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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first in a cool greenhouse. Early in January their heights were measured to the tips of their leaves. The thirteen crossed plants averaged 13.16 inches in height, and the twelve (for one had died) self-fertilised plants averaged 13.7 inches, or as 100 to 104; so that the self-fertilised plants exceeded by a little the crossed plants. TABLE XXX. Weights of Plants after they had formed Heads. No. of Pot. Crossed Plants from Pollen of fresh Stock. Self-fertilised Plants of the Third Generation. I
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F1249
Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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inches in height; or as 100 to 84. When fully grown they were again measured, but from want of time only a single plant on each side was measured; so that I have thought it best to give the earlier measurements. At the later period the three tallest crossed plants still exceeded considerably in height the three tallest self-fertilised, but not in quite so great a degree as before. The pots were left uncovered in the greenhouse, but whether the flowers were intercrossed by bees or self-fertilised
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F1249
Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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inches. 467.5 368.0 The eight tallest crossed plants now averaged 58.43, and the eight tallest self-fertilised plants 46 inches in height, or as 100 to 79. There was also a great difference in the fertility of the two lots which were left uncovered in the greenhouse. On the 17th of September the capsules from all the plants were gathered, and the seeds counted. The crossed plants yielded 243, whilst the same number of self-fertilised plants yielded only 155 seeds, or as 100 to 64. Limnanthes
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F1249
Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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flowered before the crossed! The measurements are given in the following table: TABLE LII. Lupinus luteus. No. of Pot. Crossed Plants. Self-fertilised Plants. I. Inches. 33 2/8 30 4/8 30 Inches. 24 4/8 18 4/8 28 II. 29 4/8 30 26 25 III. 30 4/8 31 31 4/8 28 27 2/8 24 4/8 Total in inches. 246.25 201.75 The eight crossed plants here average 30.78, and the eight self-fertilised 25.21 inches in height; or as 100 to 82. These plants were left uncovered in the greenhouse to set their pods, but they
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F1249
Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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present case. The seeds were placed on sand, and after germinating were planted in pairs on the opposite sides of five pots, which were kept in the greenhouse. When the seedlings were from 2 to 3 inches in height, most of the crossed had a slight advantage over the self-fertilised. The plants were trained up sticks, and thus grew to a considerable height. In four out of the five pots a crossed plant flowered before any one of the self-fertilised. TABLE LXXII. Nemophila insignis; O means that the
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F1249
Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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5/72. The twelve crossed plants now averaged 33.28, and the ten self-fertilised 19.9 inches in height, or as 100 to 60; so that they differed somewhat less than before. The plants in Pots III. and V. were placed under a net in the greenhouse, two of the crossed plants in the latter pot being pulled up on account of the death of two of the self-fertilised; so that altogether six crossed and six self-fertilised plants were left to fertilise themselves spontaneously. The pots were rather small, and
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F1249
Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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flowers the stamens are considerably shorter than the pistil, in others equal to it in length. I suspected, therefore, but erroneously as it proved, that this plant was dimorphic, like Primula, Linum, etc., and in the year 1862 twelve plants, covered by a net in the greenhouse, were subjected to trial. The spontaneously self-fertilised flowers yielded 64 grains weight of seeds, but the product of fourteen artificially crossed flowers is here included, which falsely increases the weight of the
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F1249
Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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own pollen placed on the stigma never yielded nearly a full complement of seed; whilst those left uncovered produced fine capsules, showing that pollen from other plants must have been brought to them, probably by moths. Plants growing vigorously and flowering in pots in the greenhouse, never yielded a single capsule; and * 'Proceedings of the Academy of Natural Science of Philadelphia' August 2, 1870 page 90. † 'Annales des Sc. Nat.' 4th series Bot. tom. ix. cah. 5. [page] 18
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F1249
Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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large pot (III.), the seedlings being afterwards thinned, so that an equal number was left on each side; the three tallest on each side being measured. The pots were kept in the greenhouse, and the plants were trained up sticks. For some time the young crossed plants had no advantage in height over the self-fertilised; but their leaves were larger. When fully grown and in flower the plants were measured, as follows: TABLE LXXVI. Petunia violacea (First Generation). No. of Pot. Crossed Plants
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F1249
Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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there them re-potted in larger pots and in the following year to measure them carefully; but we shall see that this was partly frustrated by so few flower-stems being then produced. These plants were left uncovered in the greenhouse; and the twelve crossed plants produced forty capsules, whilst the twelve self-fertilised plants produced only five; or as 100 to 12. But this difference does not give a just idea of the relative fertility of the two lots. I counted the seeds in one of the finest
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F1249
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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also anemophilous, or is fertilised by the wind; and of such plants only the common beet had been tried. Some plants were raised in the greenhouse, and were crossed with pollen taken from a distinct plant; and a single plant, growing quite separately in a different part of the house, was allowed to fertilise itself spontaneously. The seeds thus obtained were placed on damp sand, and as they germinated in pairs of equal age were planted on the opposite sides of four very large pots
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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Total in inches. 428 00 392 63 The eleven crossed plants now averaged 38.9, and the eleven self-fertilised plants 35.69 inches in height; or as 100 to 92, which is the same ratio as before. Differently to what occurred with the maize, the crossed plants did not flower before the self-fertilised; and though both lots flowered very poorly from having been kept in pots in the greenhouse, yet the self-fertilised plants produced twenty-eight flower-heads, whilst the crossed produced only twenty! Two long
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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, Hildebrand observed || that H. grandiflorum was highly self-sterile, whilst H. procumbens was fairly self-fertile. ('Jahrb. für wiss. Botanik' B. 7 page 464.) Thunbergia alata kept by me in a warm greenhouse was self-sterile early in the season, but at a later period produced many spontaneously self-fertilised fruits. So it was with Papaver vagum: another species, P. alpinum, was found by Professor H. Hoffmann to be quite self-sterile excepting on one occasion; whilst P. somniferum has been with me
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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.) (Lobeliaceae). Five plants left unprotected in my greenhouse produced twenty-four fine capsules, containing altogether 12.2 grains weight of seed, and thirteen other very poor capsules, which were rejected. Five plants protected from insects, but otherwise exposed to the same conditions as the above plants, produced sixteen fine capsules, and twenty other very poor and rejected ones. The sixteen fine capsules contained seeds by weight in such proportion that twenty-four would have yielded 4.66
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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two other pots, IV. and V. These pots had been kept in the hothouse, but from want of room were, whilst the plants were young, suddenly moved during very cold weather into the coldest part of the greenhouse. They all suffered greatly, and never quite recovered. After a fortnight only two of the nine self-fertilised seedlings were alive, whilst seven of the crossed survived. The tallest of these latter plants when measured was 47 inches in height, whilst the tallest of the two surviving self
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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in a warm greenhouse. All three lots consequently suffered greatly, but the Chelsea-crossed plants much less than the other two lots. On the 3rd of October the Chelsea-crossed plants began to flower again, and continued to do so for some time; whilst not a single flower was produced by the plants of the other two lots, the stems of which were cut almost down to the ground and seemed half dead. Early in December there was a sharp frost, and the stems of Chelsea-crossed were now cut down; but on
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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bring the pollen-covered hairs, with which the lower surface is clothed, into contact with the stigma by either of which means self-fertilisation might be effected. Seeds from the above crossed and self-fertilised capsules, after germinating on bare sand, were planted in pairs on the opposite sides of five moderately-sized pots, which were kept in the greenhouse. The plants after a time appeared starved, and were therefore, without being disturbed, turned out of their pots, and planted in the
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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thus differs from Ipomoea, and almost certainly from Mimulus, as with these two species a cross between flowers on the same plant did no good. CALCEOLARIA. A bushy greenhouse variety, with yellow flowers blotched with purple. The flowers in this genus are constructed so as to favour or almost ensure cross-fertilisation;* and Mr. Anderson remarks† that extreme care is necessary to exclude insects in order to preserve any kind true. He adds the interesting statement, that when the corolla is cut
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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Early in the spring the plants were gradually hardened, and turned out of their pots into the open ground without being disturbed. By the end of August the greater number had formed fine heads, but several grew extremely crooked, from having been drawn up to the light whilst in the greenhouse. As it was scarcely possible to measure their heights, the finest plant on each side of each pot was cut down close to the ground and weighed. In Table XXX. we have the result. The six finest crossed
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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of an inch. We thus see that reversion to a more natural condition acted more powerfully in favouring the ultimate growth of these plants than did a cross; but it should be remembered that the cross was with a semi-sterile variety having a feeble constitution. Iberis umbellata. Var. Kermesiana. This variety produced plenty of spontaneously self-fertilised seed under a net. Other plants in pots in the greenhouse were left uncovered, and as I saw small flies visiting the flowers, it seemed
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Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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inches, and that of the seven self-fertilised plants 16.39, or as 100 to 86. But as the plants on the self-fertilised side grew very unequally, this ratio cannot be fully trusted, and is probably too high. In both pots a crossed plant flowered before any one of the self-fertilised. These plants were left uncovered in the greenhouse; but from being too much crowded they were not very productive. The seeds from all seven plants of both lots were counted; the crossed produced 206, and the self
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Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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plants of the first generation. The crossed plants in the present case, as in the last, were more fertile than the self-fertilised, both lots being left uncovered in the greenhouse. The thirty crossed plants produced 103 seed-bearing flowers-heads, as well as some heads which yielded no seeds; whereas the twenty-nine self-fertilised plants produced only 81 seed-bearing heads; therefore thirty such plants would have produced 83.7 heads. We thus get the ratio of 100 to 81, for the number of seed
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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yielded these capsules were grown in pots in the greenhouse, so that their absolute productiveness must not be compared with that of plants growing out of doors. The above three lots of seeds, namely, the self-fertilised, intercrossed, and English-crossed, were planted in an equal state of germination (having been as usual sown on bare sand) in nine large pots, each divided into three parts by superficial partitions. Many of the self-fertilised seeds germinated before those of the two crossed
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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pots, but only the two tallest plants on each side of each pot were measured to the tops of their stems. The pots were placed in the greenhouse, and the plants trained up sticks, so that they ascended to an unusual height. In three of the pots the crossed plants flowered first, but in the fourth at the same time with the self-fertilised. When the seedlings were between 6 and 7 inches in height, the crossed began to [page] 14
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Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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, and the pots, which were much too small, were kept in the greenhouse. The plants in consequence grew badly, and the self-fertilised suffered most in both pots. The two crossed plants when in flower during the following spring were 9 inches in height; one of the self-fertilised plants was 8, and the three others only 3 inches in height, being thus mere dwarfs. The two crossed plants produced thirteen pods, whilst the four self-fertilised plants produced only a single one. Some other self-fertilised
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Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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five crossed plants is 86 inches, and that of the five self-fertilised plants 82.35; or as 100 to 96. The pots were kept in the greenhouse, and there was little or no difference in the fertility of the two lots. Therefore as far as these few observations serve, the advantage gained by a cross is very small. * 'Kenntnis der Befruchtung' 1844 pages 573, 577. [page] 15
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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pollen from a distinct plant growing in the same row, and others were allowed to fertilise themselves under a net. The two lots of seeds thus obtained were sown on opposite sides of two large pots, but only four pairs came up at the same time. The pots were kept in the greenhouse. The seedlings of both lots when between 6 and 7 inches in height were equal. When nearly full-grown they were measured, as in the following table: TABLE LVII. Pisum sativum. No. of Pot. Crossed Plants. Self-fertilised
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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green as those of the crossed seedlings. The pots were kept in the greenhouse, and as the plants on the following spring (1868) looked unhealthy and had grown but little, they were plunged, still in their pots, into the open ground. The plants all suffered much from the sudden change, especially the self-fertilised, and two of the latter died. The remainder were measured, and I give the measurements in the following table, because I have not seen in any other species so great a difference between
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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flowers contained on an average 4.1 seeds, so that these were the most productive of all; and the seeds themselves looked finer even than those from the crossed perfect flowers. The seeds from the crossed perfect flowers and from the self-fertilised cleistogene flowers were allowed to germinate on sand; but unfortunately only two pairs germinated at the same time. These were planted on the opposite sides of the same pot, which was kept in the greenhouse. In the summer of the same year, when the
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Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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germinating on sand, were planted in pairs on the opposite sides of Pots II. and III., which were at first kept in the greenhouse and then turned out of doors. The plants were measured when in full flower. The following table, therefore, includes plants belonging to two generations. When the seedlings of the two lots were only 5 or 6 inches in height they were equal. In Pot III. one of the self-fertilised plants died before flowering, as has occurred in so many other cases. * The Compositae are
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Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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worth speaking of between the two lots in height; though other great advantages are derived, as we have seen, from cross-fertilisation. From being grown in pots and kept in the greenhouse, none of the plants produced any capsules. Lobelia ramosa.* Var. Snow-flake. The well-adapted means by which cross-fertilisation is ensured in this genus have been described by several authors.† The pistil as it slowly increases in length pushes the pollen out of the conjoined anthers, by the aid of a ring of
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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; and they flowered so much earlier in all the pots, that the height of the flower-stems could be fairly compared only in Pots I. and II. * Gärtner has shown that certain plants of Lobelia fulgens are quite sterile with pollen from the same plant, though this pollen is efficient on any other individual; but none of the plants on which I experimented, which were kept in the greenhouse, were in this peculiar condition. [page] 18
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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, whence my plants originally came, a fresh plant differing in no respect from mine except in the colour of the flowers, which was a fine purple. But this plant must have been exposed during at least four generations to very different conditions from those to which my plants had been subjected, as these had been grown in pots in the greenhouse. Eight flowers on the self-fertilised plants in Table LXXXI., of the last or fourth self-fertilised generation, were fertilised with pollen from this fresh
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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germination, were thickly sown in an eighth pot. The pots were kept in the greenhouse, and the plants trained up sticks. They were first measured to the tops of their stems when coming into flower; and the twenty-two Westerham-crossed plants then averaged 25.51 inches; the twenty-three intercrossed plants 30.38; and the twenty-three self-fertilised plants 23.40 inches in height. We thus get the following ratios: The Westerham-crossed plants in height to the self-fertilised as 100 to 91. The Westerham
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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green than those of the intercrossed and self-fertilised plants. Relative Fertility of the Three Lots of Plants. None of the plants in pots in the greenhouse ever produced a capsule; and this may be attributed in chief part to the exclusion of moths. Therefore the fertility of the three lots could be judged of only by that of the plants growing out of doors, which from being left uncovered were probably cross-fertilised. The plants in the three rows were exactly of the same age and had been
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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raised from the two plants, their heights have been added together in calculating the averages. I should state that by the accidental fall of a large bush in the greenhouse, several plants in both the series were much injured. These were at once measured together with their opponents and afterwards thrown away. The others were left to grow to their full height, and were measured when in flower. This accident accounts for the small height of some of the pairs; but as all the pairs, whether only partly
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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capsule for the crossed and self-fertilised flowers. The flowers hang downwards, and as the stigmas stand close beneath the anthers, it might have been expected that pollen would have fallen on them, and that they would have been spontaneously self-fertilised; but these covered-up plants did not produce a single capsule. On some other occasions uncovered plants in the same greenhouse produced plenty of capsules, and I suppose that the flowers had been visited by bees, which could hardly fail to
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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. These plants, which were grown in pots in the greenhouse, were evidently in a very sterile condition, and the seeds in both sets of capsules, especially in the self-fertilised, although numerous, were of so poor a quality that it was very difficult to determine which were good and which bad. But as far as I could judge, the crossed capsules contained on an average 6.3 good seeds, with a maximum in one of thirteen; whilst the self-fertilised contained 6.05 such seeds, with a maximum in one of
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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of the fourth illegitimate generation (i.e., great-great-grandchildren of plants which had been legitimately fertilised), growing vigorously in pots in the greenhouse, were legitimately fertilised with pollen from an almost wild short-styled cowslip, and these flowers yielded some fine capsules. Thirty other flowers on the same illegitimate plants were fertilised with their own pollen, and these yielded seventeen capsules, containing on an average thirty-two seeds. This is a high degree of
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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number of seedlings which thus perished, the surviving self-fertilised plants grew thinly in the row, and thus had an advantage over the crossed plants, which grew very thickly in the other row. The young plants in the two rows were protected by a little straw during the winter, and those in the two large pots were placed in the greenhouse. There was no difference between the two lots in the pots until the ensuing spring, when they had grown a little, and then some of the crossed plants were
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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height of all the plants of the ten generations 73 85 84 73 66 02 '' '' 77 Mimulus luteus three first generations, before the new and taller self-fertilised variety appeard 10 8 19 10 5 29 '' '' 65 Digitalis purpurea 16 51 33 8 35 87 '' '' 70 Calceolaria (common greenhouse variety) 1 19 50 1 15 00 '' '' 77 Linaria vulgaris 3 7 08 3 5 75 '' '' 81 Verbascum thapsus 6 65 34 6 56 50 '' '' 86 Vandellia nummularifolia crossed and self-fertilised plants, raised from perfect flowers 20 4 30 20 4 27
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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). Highly self-fertile. Leptosiphon androsaceus (Polemoniacae). Plants under a net produced a good many capsules. Primula mollis (Primulaceae). A non-dimorphic species, self-fertile: J. Scott, in 'Journal of the Linnean Society Botany' volume viii. 1864 page 120. Nolana prostrata (Nolanaceae). Plants covered up in the greenhouse, yielded seeds by weight compared with uncovered plants, the flowers of which were visited by many bees, in the ratio of 100 to 61. Ajuga reptans (Labiatae). Set a good
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Book:
Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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from plants which had been raised from seed procured from a garden at Chelsea. The Chelsea plants bore yellow flowers blotched with red, but differed in no other respect. They had been grown out of doors, whilst mine had been cultivated in pots in the greenhouse for the last eight generations, and in a different kind of soil. The seedlings raised from this cross with a wholly different stock may be called the Chelsea-crossed. The three lots of seeds thus obtained were allowed to germinate on
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Darwin, C. R. 1876. The effects of cross and self fertilisation in the vegetable kingdom. London: John Murray.
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third of their full size, and the crossed plants then averaged 3.87 inches, and the self-fertilised only 2.00 inches in height; or as 100 to 52. They were kept in the greenhouse, and did not grow vigorously. Whilst in flower they were again measured to the summits of their stems (see Table XLI.), with the following result: The average height of the fourteen crossed plants is here 5.58 inches, and that of the fourteen self-fertilised 2.37; or as 100 to 42. In four out of the five pots, a crossed
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F1277
Book:
Darwin, C. R. 1877. The different forms of flowers on plants of the same species. London: John Murray.
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tilised with their own pollen, produced 15 capsules, containing an average of 16 3 seeds. As already stated, the probable average with legitimate plants for a union of this nature is rather above 20 seeds. These plants were remarkably healthy and vigorous, as long as they were kept under highly favourable conditions in pots in the greenhouse; and such treatment greatly increases the fertility of the cowslip. When these same plants were planted during the next year (which, however, was an
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F1277
Book:
Darwin, C. R. 1877. The different forms of flowers on plants of the same species. London: John Murray.
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. These whilst still in the greenhouse produced smaller leaves and shorter flower-stalks than some legitimate plants with which they grew in competition; but it should be observed that the latter were the product of a cross with a fresh stock, a circumstance which by itself would have added much to their vigour.* When these illegitimate plants were transferred to fairly good soil out of doors, they became during the two following years much more dwarfed in stature and produced very few flower-stems
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CUL-DAR66.121-128
Note:
1877--1878
Leaves — Right-side cleaned of bloom [with tepid water]
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no injury, but on 27th whole space where sponge was quite brown much injured!!! I think this must have been effect of water, for bloom so apt to be removed, no flower-stem wd be perfect if its removal sufficed to cause injury (a) July 25 Œnantha fistulosa cleaned whole of 2 stalks leaves: bloom poor, — not worth trying — white wool — (Sept 11th bloom apparently removed.) Dec 13 no effect July 25 Aquilegia glandulosa — cleaned 2 leaflets, young plants, Greenhouse white wool— cannot be well
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CUL-DAR66.121-128
Note:
1877--1878
Leaves — Right-side cleaned of bloom [with tepid water]
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Aug 1. 12˚ 9˚ 30' Papaver somniferum (white flowered, not much bloom) sponged with tepid water right side of 4 leaves — white Thread. Sept 17th. The peduncle which were cleaned of bloom, when pods were cleaned, both on plants in greenhouse out-of doors look browner. Caladium (Mr Nash) Aug. 2d. 9˚ sponged with tepid water whole 1/2 leaf right side, between 2 of lat veins rubbed with pumice up to margin; propped up filled centre with water. Aug 8' 9' I see near place which was pumiced are
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Darwin, C. R. 1877. The different forms of flowers on plants of the same species. London: John Murray.
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cowslips of both forms, protected from insects in my greenhouse, did not set one pod, though artificially fertilised flowers in other pots produced an abundance. We thus see that the visits of insects are absolutely necessary for the fertilisation of Primula veris. If the corolla of the long-styled form had dropped off, instead of remaining attached in a withered state to the ovarium, the anthers attached to the lower part of the tube with some pollen still adhering to them would have been
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Darwin, C. R. 1877. The different forms of flowers on plants of the same species. London: John Murray.
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the spring of 1862 forty flowers were fertilised at the same time in both ways. The plants were accidentally exposed in the greenhouse to too hot a sun, and a large number of umbels perished. Some, however, remained in moderately good health, and on these there were twelve flowers which had been fertilised legitimately, and eleven which had been fertilised illegitimately. The twelve legitimate unions yielded seven fine capsules, containing on an average each 57 3 good seeds; whilst the eleven
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F1277
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Darwin, C. R. 1877. The different forms of flowers on plants of the same species. London: John Murray.
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, and the female organs of the long-styled form in a more or less complete state, they spontaneously produced a surprising number of capsules, which generally contained a large average of remarkably fine seeds. With ordinary cowslips, legitimately fertilised, I once obtained from plants cultivated in the greenhouse the high average, from seven capsules, of 58 7 seeds, with a maximum in one capsule of 87 seeds; but from plants grown out of doors I never obtained a higher average than 41 seeds. Now
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. The non-cleaned most decayed.) (33) Oxalis small-kind yellow-flowered, not coppery, greenhouse— 2 leaves. (go to sleep at night.) (June 20. 10. AM. Not cleaned. wetted. Cleaned slightly yellower) (June 21. 10 AM no difference ditto) (June 22. 2.30. cleaned decidedly most decayed) (
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CUL-DAR66.166v
Note:
1877.06.30
Greenhouse bushy Coronilla — Both surfaces of leaves silvery
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The Complete Work of Charles Darwin Online [166v] June 30th 77. Southampton — Greenhouse bushy. Coronilla — Both surfaces of leaves silvery. — It is rather of that young leaves very imperfectly silvery somewhat wetted by immersion.— It is only the young leaves which go to sleep the movement slight, the terminal leaflet turns a little up so as to be vertical or with lower surface obliquely towards the sky (as in clovers) with next 2 lateral leaflets turned a little towards each other, but they
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CUL-DAR66.113
Note:
1877.07.10--1877.07.26
Broccoli — Soaking / 9 am Frank placed 8 cleaned & 8 uncleaned leaves of
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The Complete Work of Charles Darwin Online [113] 1877 Broccoli Cabbages — Soaking July 10 9˚ A.m. Frank placed 8 cleaned 8 uncleaned leaves of Walcherer Broccoli placed in same quantity of water in Greenhouse — with cover to keep petioles (which were out of water) damp: The leaves kept immersed by lead weight.— (Also at 10˚. 45' 2 leaves 1 in each jar, with right side alone cleaned.) (July 12th 13th 14th 12˚- 15th do —16th do — 19th do — 20th do 8˚ 30' non-cleaned sides dry, no difference in
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F2093
Periodical contribution:
Timiriazev, Kliment. [1877]. A visit to Darwin at Down. From: Historical note K. A. Timiriazev: A visit to Darwin, with notes by Leon Bell. Archipelago 9 (2006): 44-58.
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insectivorous plants. In spite of the fact that it was a hot July day (albeit a cloudy one) and the greenhouse was within a short walking distance, his wife and son brought a short overcoat and the soft felt hat that are so famous now from his photographs. In front of the verandah there was a large English lawn trimmed like velvet on which you could nevertheless walk freely, sit or lie about. The flowerbeds were nothing special. The greenhouse in the opposite corner of the garden was small, the
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vertically. Sept 19' F. [Frank] measured angle of petiole at night day in greenhouse cd perceive no difference. Does not look like a Trifolium.— (over) Sept 29' observed at 8° 30' P.m. when leaves well asleep. Plant now grown 2-3 ft tall, terminal leaflet never shows any tendency to turn vertical during sleep on its own axis so as to become vertical, but often does not overlap neatly the 2 lateral leaflet which approach each other pretty closely. Movement closely like that of Trifolium. [105v
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CUL-DAR66.143
Note:
[1877].08.05--[1877].09.21
Papaver somniferus / 4h pm removed bloom from 5 young pods & peduncles in
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it very odd that the pods on plant s out of doors have their stigmas, both cleaned uncleaned equally dry. I rather suspect peduncles of those out of doors are rather browner. — Sept. 21. compared pods, not perfectly mature, in greenhouse out of doors, which had been cleaned with bloom — perhaps peduncles a shade browner. Put those on plant out of doors to dry in window — white wool cleaned pods. Poppy greenhous
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CUL-DAR66.143
Note:
[1877].08.05--[1877].09.21
Papaver somniferus / 4h pm removed bloom from 5 young pods & peduncles in
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The Complete Work of Charles Darwin Online [143] Papaver somniferum Aug. 5' 4˚ P.m. removed bloom from 5 young pods peduncles in greenhouse marked with white wool. Also 5 pods of nearly same size marked with black thread for comparing Some large some small — removed bloom with damp sponge. — Aug 7th 12˚ 35. I find all 3 pods with much bloom on them; cleaned them carefully again with brush under stigma. 12˚ 40' cleaned a 4th pod; marked with black white wool. Aug. 9th 10˚ 45. one of the pods
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The Complete Work of Charles Darwin Online [68] Down Aug 17th. 1877 Horace Dwarf Kidney Bean — Greenhouse Observed angles of both parts of petiole of two leaves at 12 midday and at 10. 15 P.M. of two leaves, one very young, the other fully developed. [Fig] [Table] I think that The movement of one part of the petiole relatively to the other is so small (2°1/2 nd 1 1/4°) that it may be accounted for by errors of observation, especially in the young leaf as the upper part of the petiole was very
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CUL-DAR209.14.151-154
Note:
1877.09.03--1877.09.13
Tropaeolum minus / Tropaeolum canariense
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If the horizon to the N.E late in evening had been brighter, I shd have said that leaves became vertical to face last trace of light remained fixed in this position till the light reappeared next morning. It is surprising what a little obscurity during the day prevents the leaves at night going to sleep, as shown by those on back of plants in greenhouse in the present experiment.— After the plants had gone to sleep to above extent they were placed in dark cupboards next morning (Sept 7' 9° A.m
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CUL-DAR209.14.151-154
Note:
1877.09.03--1877.09.13
Tropaeolum minus / Tropaeolum canariense
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leaves certainly less vertical than they were last night. Sept. 13' Put plants in greenhouse, went at night but cd see no signs of plain sleep. — (
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CUL-DAR209.14.151-154
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1877.09.03--1877.09.13
Tropaeolum minus / Tropaeolum canariense
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of plant, facing to light gone to sleep with petioles bent, but many to the back the depending leaves on both sides not gone to sleep. To On the left side, (of the leaves encircled with thread) only those on tall shoot gone to sleep. All this agrees with what I observed in greenhouse. viz that plants on inner or dark side did not go to sleep. no external difference perceptible in bending part of Petiole. (over) (a) (
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The Complete Work of Charles Darwin Online [71] Lupinus menziesii. —(California) Sept. 4 2° 30' P.M. morning [annotated diagram] (B) Pet Pet older at right angles to A The 2 leaves B at the same distance apart at night as at day [annotated diagram] asleep 2 opposite petioles A Pet Pet Black wool Stars sub-horizontal or generally slightly inclined to light 8° 15' P.m. The stars almost as horizontal as at during day. (Keep in greenhouse see if will then go to sleep) (of no use) Petioles rose a
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The Complete Work of Charles Darwin Online [171] Robinia 1877 Sept 8. 11° 30' Sponged 2 lateral 1 terminal on 2 nice leaves with water at 88° to 90°— marked red wool Bush in greenhouse put on drops of water — Also spheres on 4 other leaves not sponged. The sponged leaves on left side viewed from base. (Sept 20' freed leaves: Sept 21st no effect, or at most the faintest shade of yellow. Sept. 25' There can now be no doubt that all the leaves especially the sponged ones are yellowish more
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The Complete Work of Charles Darwin Online [79] Lupinus sub-carnosus Sept 11. shorter leaves form horizontal stars at night; taller ones with leaflets deflexed— put into greenhouse. First leaves have only 3 leaflets. L. Douglasi [Nachipalensis] (6) horizontal stars at night sent to greenhouse [80
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Lupinus subcarnosus Oct 5. 6th Kept as above in greenhouse. At night very little or no change in most of leaves; but the leaflets in some formed an obtuse a good cone of 103° [sketch] 103, so that each leaflet on the 2 leaves which were measured fell 38° 30' beneath horizon. — No change in the position of the petioles. [in margin:] Leaves fell at night L. polyphyllus, nanus, menziesii, speciosus, not in [illeg] albiform Hartwegii, all Kept in greenhouse. The leaves went very little or not at
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The Complete Work of Charles Darwin Online [49] Sept 14. 77 Nicotiana glauca Plant old terminal leaflets hardy g do not visibly go to sleep.— fixed long bristle to midrib traced course on horizontal planes of glass over it. — as shown from 7° 23' A.m. to 8° 30' P.m. — see diagram. F. also observed both leaves in water they certainly move like most sleeping plants — confirm Pfeffer on N. rustica. — The above leaf stood highly inclined with back to front of greenhouse upper surface to roof of do
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The Complete Work of Charles Darwin Online [81] Sleep. 1877 Lupinus arboreus Oct 5-6th. Kept in greenhouse— Four leaflets leaves observed previous afternoon next morning — The leaflets formed during day a horizontal star; but at night a cone of 108° 30' [sketch], a second leaf of 107°; so that the each leaflet fell about 36° beneath horizon. The petioles moved very little during sleep, so little in opposite directions, that safer to say in no marked manner [82
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1877 Lupinus arboreus Oct 11 moderately young leaf facing light, in nearly position which had stood in greenhouse considerably inclined — Bristle fixed along middle of one of longest leaflets, which extended transversely vertical glass 13 2/8 of inch to pointed card behind leaf. card from end of bristle 1 7/8 of inch. From 8° 40' to 10° 14' went down; then rose z:zagged greatly to 2. 49'; then rose to 3: 57. Then began to fall, which was incipient sleep movement; downward movement rapid, but
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The Complete Work of Charles Darwin Online (1 Lupinus speciosus Oct 11th 1877. Moderately old leaf quite free petiole spink from stock without any stem Leaf facing obliquely to light as in greenhouse.— Bristle fixed to one of longer leaflets — Glass to tip of bristle 11 3/8 inch; glass to paper mark behind 13 3/8. Traced on vertical glass. At first bristle went down below glass then rose above glass very high, so that I altered adjustment. First dot at 10°. 16' — see diagram (1) between this
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The Complete Work of Charles Darwin Online [115] Melilotus officinalis 1877 Oct. 24 Old Plant put facing study window in same position as during weeks in greenhouse, with most of leaves hanging down vertically facing light.— Traced on vertical glass.— Right-Hand Diagram B stem tied showing nutation movement of whole leaf, but there was little movement as terminal leaflet depended vertically, but there was some. This diagram magnified 7.5 times. Left-Hand. Diagram A This more interesting, as it
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The Complete Work of Charles Darwin Online [75] Oct. 29' 1877 Oxalis acetosella observed at night (Plant had been Kept in pot in greenhouse) Secured petioles of 2 nice youngis[h] leaves, close under leaflets to little sticks; with long bristles affixed longitudinally to central leaflets marked a mark behind, traced on vertical glass.— Diagram A shows that lea this leaflet after going to sleep moved a little laterally from side to side between 7° 10° 40' P.m, going thrice in one direction twice
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The Complete Work of Charles Darwin Online [20] Oct 30 (77) Seedling Onions (Used) Long glass-filament fixed to stem to diverticulum on distal portion which rises doubled back [sketch] Kept in dark traced on Horizontal glass; but the tracing failed for both points swept from wes N.E to S W so much that twice I had to move pot, then by 1° moved off glass, was so next morning— I can only conjecture that had in greenhouse bent to light was recovering all day its upright position, through negative
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The Complete Work of Charles Darwin Online [31v] self-fertilised capsules were however the heavier in the proportion ratio of 100 to 87. Three or four pairs of these two lots of Seeds in an equal state of germination were planted on the opposite sides of four pots: but only the two tallest plants on each side of each pot were measured to the tops of their stems. The pots were placed in the greenhouse, and the plants trained up sticks, so that they ascended to an unusual height. In thre
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The Complete Work of Charles Darwin Online [1] Nov. 25 to 27. 1877 A very pleasant visit at Down ─ C.D in particular force his own health: the return of the Litchfields, she looking very well he promising for total recovery:1 and W. Ds marriage all made the house very cheerful.2 Francis Galton there interesting. C.D was out hour by hour from 7.30 till dark walking slowly to his greenhouse alone ─ with straw hat and short cloak ─ eyes on ground ─ taking measurements of movements of plants which
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The Complete Work of Charles Darwin Online (1 Nov. 3d 77 Trifolium resupinatum (Used) Nutation of stem alone. Selected young shoot fixed filament of glass to one side, (Kept in almost darkness,) traced on [sketch] vertical glass during 7° from 9° 26 to 4° 30' — Tracing Tracing magnified but not very greatly — During this time went down thrice up twice or thrice or horizontally transversely — no doubt about nutation (Nov. 5th The plant was put yesterday into greenhouse, today I fixed glass
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hardly be normally an upward or closing movement during the morning, as they close during night; there certainly must be a rising movement in the Evening. Therefore I believe that the complete darkness altogether put out all the normal movement. It is especially noteworthy that after being kept dark all day, they the Cots did not go to sleep at night at 10° 30' P.m; being kept dark during whole night were not asleep next morning at 7° a.m.) (Nov. 7th After being all day in greenhouse but day very
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CUL-DAR209.4.199-200
Note:
1877.11.20--1877.11.26
Lathyrus nissolia [with diagram attached]
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same relative position in Ho warm greenhouse)— Next day Nov. 22d (B in diagram,) it moved from light from 7° 30', to 8° 25' then to N.W. then to N.E until 11° 34'; then from light till about 1° then to S.E. made one irregular circle.— I certainly think the leaves revolve, but much influenced by light. Plant A of diagram tip of nearly straight grass-leaf observed— made an open ellipse during same period, from N W. to S.E. moving a little to light in morning from it in afternoon opposed to course
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Nov. 29th The cots were widely open today in greenhouse, the short brown paper tube having been removed, ∴ I think the small angle which the Cots formed together yesterday was due to the direction in which ؟؟ light fell on them from above: to night 10° P.m, only 1 of the 4 white threads almost closed, the 3 others being much more closed than during the day.— Of the 12 with black threads none closed, though the cots much raised of most of seedlings— Nov. 30' 10° P.m. Cots. seem hardy at all
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unpainted side alone is seen.— It is the same appearance as from Heliotropism, with this remarkable distinction that there is no angular bend at about .1 or .2 above ground. Jan 17' 8° a.m.— somewhat more bowed — 1 Killed 1 I think accidentally injured.─ Pot put into greenhouse to see if will recover.—) Jan. 25' are growing well not 1 Killed— some days ago the blackened side became convex, on examining one under microscope, the paint, though still soft has contracted whole sheath has expanded
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F1782
Periodical contribution:
Darwin, C. R. 1877. Growth under difficulties. Gardeners' Chronicle 8 (29 December): 805.
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Darwin, C. R. 1877. Growth under difficulties. Gardeners' Chronicle and Agricultural Gazette 8 (29 December): 805. [page] 805 GROWTH UNDER DIFFICULTIES. [...] The following communication, with which we have been favoured by Mr. Darwin, affords us another illustration of similar character:— The enclosed branch of Cotyledon (Echeveria stolonifera was cut from a plant growing in my greenhouse, and was suspended on August 10 in my study, which is a dry room, and in which a fire burns most of the
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[calculations not transcribed] Used (while the seedling was kept in the greenhouse on Feb 9th, the Cots, in the middle of the day → (Feb 9th) open formed together an angle estimated at 30°; at 6° P.m. they were nearly closed next morning, at 8° a.m (Feb 10th) they were completely shut or asleep. The plants were then placed in a box with a glass cover, so as to be feebly illuminated from above whilst the circumnutation of the hypochil was observed; the cotyledons did not open in the least all
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CUL-DAR209.4.397-398
Draft:
[1877?].12.17--[1877?].12.18
Tropaeolum / Draft of Cross and self fertilisation.
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sometimes produced Corydalis C. halleri is only slightly self-sterile, and C. intermedia not at all so. *(*Botany Zeitung June 27 1873). In another Fumariaceous genus Hypercoum, Hildebrand observed *(*Jahrb. für Wiss. Botanik Bd VII p 464) that H grandiflorum was highly self-sterile, whilst H procumbens was fairly self-sterile. Thunbergia alata kept by me in a warm greenhouse was self-fertile sterile early in the season, but at a later period produced spontaneously many self-fertilised fruit. So it
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F1251
Book:
Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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greenhouse, produced capsules in number(about) as 100 to 100 LUPINUS LUTEUS crossed and self-fertilised plants of the 2nd generation, left uncovered in the greenhouse, produced seeds in number (judged from only a few pods) as 100 to 88 PHASEOLUS MULTIFLORUS crossed and self-fertilised plants, left uncovered in greenhouse, produced seeds in number(about) as 100 to 100 LATHYRUS ODORATUS crossed and self-fertilised plants of the 2nd generation, left uncovered in the greenhouse, but certainly self
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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the cases which follow that this result was accidental, owing to only a few plants having been measured, and to one of the self-fertilised plants having grown only to a height of 15 inches. The plants had been kept in the greenhouse, and from being drawn up to the light had to be tied to sticks in this and the following trials. They were measured to the summits of their flower-stems. The four crossed plants here average 29 68 inches, and the four self-fertilised 25 56 in height; or as 100 to
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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occasions with plants of the first generation of Nicotiana. The crossed plants always withstood the injurious effects of being suddenly removed into the open air after having been kept in the greenhouse better than did the self-fertilised. On several occasions they also resisted much better cold and intemperate weather. This was manifestly the case with some crossed and self-fertilised plants of Ipom a, which were suddenly moved from the hothouse to the coldest part of a cool greenhouse. The
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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hemisphere, with a complete reversal of the seasons, were thus rendered slightly self-fertile, whereas they seem always to be completely self-sterile in their native home. Senecio cruentus (greenhouse varieties, commonly called Cinerarias, probably derived from several fruticose or herbaceous species much intercrossed*). Two purple-flowered varieties were placed under a net in the greenhouse, and four corymbs on each were repeatedly brushed with flowers from the other plant, so that their
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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flowers; as they crawled into them, they rubbed against the bristles which project from the anthers, and became dusted with pollen. Veronica agrestis (Scrophulariace ). Covered-up plants produced an abundance of seeds. I do not know wheter any insects visit the flower; but I have observed Syrphid repeatedly covered with pollen visiting the flowers of V. hederaefolia and cham drys. Mimulus luteus (Scrophulariace ). Highly self-fertile. Caleeolaria (greenhouse variety) (Scrophulariace ). Highly
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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three long rows, however, were not of quite equal lengths, and the plants were much crowded, so that it would have been extremely difficult to have ascertained how many capsules were produced by them, even if I had been willing to undertake so laborious a task as to collect and count all the capsules. But this was feasible with the plants grown in pots in the greenhouse; and although these were much less fertile than those growing out of doors, their relative fertility appeared, after
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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a pair of plants, which were kept in a pot in the greenhouse. The crossed plant grew to a height of 33 inches, and the self-fertilised to that of 26 inches. The former produced, whilst still kept in the greenhouse, eight pods, containing on an average 2 77 seeds; and the latter only two pods, containing on an average 2 5 seeds. The average height of the two crossed plants of the two [page] 15
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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The self-fertilised peas were, however, quite as heavy as those from the crossed pods. From these two lots of seeds, the plants of the next generation were raised. Plants of the Second Generation. Many of the self-fertilised peas just referred to germinated on sand before any of the crossed ones, and were rejected. As soon as I got equal pairs, they were planted on the opposite sides of two large pots, which were kept in the greenhouse The seedlings thus raised were the grandachildren of the
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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conditions in pots in the greenhouse. As long as they were cultivated in this manner, they grew well and were healthy and fertile. Their fertility even increased in the later generations, as if they were becoming habituated to illegitimate fertilisation. Plants of the first illegitimate generation when taken from the greenhouse and planted in moderately good soil out of doors grew well and were healthy; but when those of the two last illegitimate generations were thus treated they became excessively
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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; and to the summits of their male flowers, 53 96 and 43 45 inches; or as 100 to 80. PHALARIS CANARIENSIS. Hildebrand has shown in the paper referred to under the last species, that this hermaphrodite grass is better adapted for cross-fertilisation than for self-fertilisation. Several plants were raised in the greenhouse close together, and their flowers were mutually intercrossed. Pollen from a single plant growing quite separately was collected and placed on the stigmas of the same plant. The
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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PETUNIA VIOLACEA left uncovered as above: offspring of plants self-fertilised for four generations and then crossed by a fresh stock, compared with plants of the 5th self- fertilised generation, produced seeds, as judged by the weight of an equal number of capsules as 100 to 46 CYCLAMEN PERSICUM crossed and self-fertilised plants, left uncovered in the greenhouse, produced capsules in number as 100 to 12 ANAGALLIS COLLINA crossed and self-fertilised plants, left uncovered in the greenhouse
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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vigour to the crossed. My attention was now thoroughly aroused, for I could hardly doubt that the difference between the two beds was due to the one set being the offspring of crossed, and the other of self-fertilised flowers. Accordingly I selected almost by hazard two other plants, which happened to be in flower in the greenhouse, namely, [page] 1
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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. In the successive generations they were exposed to slightly different conditions as the seasons varied, and they were raised at different periods. But in other respects all were treated alike, being grown in pots in the same artificially prepared soil, being watered at the same time, and kept close together in the same greenhouse or hothouse. They were [page] 2
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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and self-fertilised generations Small amount of pollen in the anthers of the self-fertilised plants of the later generations, and the sterility of their first-produced flowers Uniform colour of the flowers produced by the self-fertilised plants The advantage from a cross between two distinct plants depends on their differing in constitution. A PLANT of Ipom a purpurea, or as it is often called in England the convolvulus major, a native of South America, grew in my greenhouse. Ten flowers on
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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first in a cool greenhouse. Early in January their heights were measured to the tips of their leaves. The thirteen crossed plants averaged 13 16 inches in height, and the twelve (for on had died) self-fertilised plants averaged 13 7 inches, or as 100 to 104; so that the self-fertilised plants exceeded by a little the crossed plants. TABLE XXX. Weight of Plants after they had fromed Heads. No. of Pot. Crossed Plants fromPollen of fresh Stock. Self-fertilised Plantsof the Third Generation. Ounces
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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inches in height; or as 100 to 84. When fully grown they were again measured, but from want of time only a single plant on each side was measured; so that I have thought it best to give the earlier measurements. At the later period the three tallest crossed plants still exceeded considerably in height the three tallest self-fertilised, but not in quite so great a degree as before. The pots were left uncovered in the greenhouse, but whether the flowers were intercrossed by bees or self-fertilised
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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inches. 467 5 368 0 The eight tallest crossed plants now averaged 58 43, and the eight tallest self-fertilised plants 46 inches in height, or as 100 to79. There was also a great difference in the fertility of the two lots which were left uncovered in the greenhouse. On the 17th of September the capsules from all the plants were gathered, and the seeds counted. The crossed plants yielded 243, whilst the same number of self-fertilised plants yielded only 155 seeds, or as 100 to 64. LIMNANTHES DOUGLASH
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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flowered before the crossed! The measurements are given in the following table: TABLE LII. Lupinus luteus. No. of Pot. Crossed Plants. Self-fertilised Plants. Inches. Inches. I. 33 2/3 24 4/8 30 4/8 18 4/8 30 28 II. 29 4/8 26 30 25 III. 30 4/8 28 31 27 2/8 31 4/8 24 4/8 Total in inches. 246 25 201 75 The eight crossed plants here average 30 78, and the eight self-fertilised 25 21 inches in height; or as 100 to 82. These plants were left uncovered in the greenhouse to set their pods, but they
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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present case The seeds were placed on sand, and after germinating were planted in pairs on the opposite sides of five pots, which were kept in the greenhouse. When the seedlings were from 2 to 3 inches in height, most of the crossed had a slight advantage over the self-fertilised. The plants were trained up sticks, and thus grew to a considerable height. In four out of the five pots a crossed plant flowered before any one of the self-fertilised. TABLE LXXII. Nemophila insignis; O means that the
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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preceding table. The twelve crossed plants now averaged 33 28, and the ten self-fertilised 19 9 inches in height, or as 100 to 60; so that they differed somewhat less than before. The plants in Pots III. and V. were placed under a net in the greenhouse, two of the crossed plants in the latter pot being pulled up on account of the death of two of the self-fertilised; so that altogether six crossed and six self-fertilised plants were left to fertilise themselves spontaneously. The pots were rather
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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flowers the stamens are considerably shorter than the pistil, in others equal to it in length. I suspected, therefore, but erroneously as it proved, that this plant was dimorphic, like Primula, Linum, c., and in the year 1862 twelve plants, covered by a net in the greenhouse, were subjected to trial. The spontaneously self-fertilised flowers yielded 64 grains weight of seeds, but the product of fourteen artificially crossed flowers is here included, which falsely increases the weight of the self
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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sides of a large pot (III.), the seedlings being afterwards thinned, so that an equal number was left on each side; the three tallest on each side being measured. The pots were kept in the greenhouse, and the plants were trained up sticks. For some time the young crossed plants had no advantage in height over the self-fertilised; but their leaves were larger. When fully grown and in flower the plants were measured, as follows: TABLE LXXVI. Petunia violacea (First Generation). No. of Pot. Crossed
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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have them re-potted in larger pots and in the following year to measure them crefully; but we shall see that this was partly frustrated by so few flower-stems being then produced. These plants were left uncovered in the greenhouse; and the twelve crossed plants produced forty capsules, whilst the twelve self-fertilised plants produced only five; or as 100 to 12. But this difference does not give a just idea of the relative fertility of the two lots. I counted the seeds in one of the finest capsules
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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also anemophilous, or is fertilised by the wind; and of such plants only the common beet had been tried. Some plants were raised in the greenhouse, and were crossed with pollen taken from a distinct plant; and a single plant, growing quite separately in a different part of the house, was allowed to fertilise itself spontaneously. The seeds thus obtained were placed on damp sand, and as they germinated in pairs of equal age were planted on the opposite sides of four very large pots
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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Total in inches. 428 00 392 63 The eleven crossed plants now averaged 38 9, and the eleven self-fertilised plants 35 69 inches in height; or as 100 to 92, which is the same ratio as before. Differently to what occurred with the maize, the crossed plants did not flower before the self-fertilised; and though both lots flowered very poorly from having been kept in pots in the greenhouse, yet the self-fertilised plants produced twenty-eight flower-heads, whilst the crossed produced only twenty! Two long
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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observed || that H. grandiflorum was highly self-sterile, whilst H. procumbens was fairly self-fertile. Thunbergia alata kept by me in a warm greenhouse was self-sterile early in the season, but at a later period produced many spontaneously self-fertilised fruits. So it was with Papaver vagum: another species, P. alpinum, was found by Professor H. Hoffmann to be quite selfsterile excepting on one occasion; whilst P. somniferum has been with me always completely self-fertile. Eschscholtzia
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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greenhouse produced twenty-four fine capsules, containing altogether 12 2 grains weight of seed, and thirteen other very poor capsules, which were rejected. Five plants protected from insects, but otherwise exposed to the same conditions as the above plants, produced sixteen fine capsules, and twenty other very poor and rejected ones. The sixteen fine capsules contained seeds by weight in such proportion that twenty-four would have yielded 4 66 grains. So that the unprotected plants produced
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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two other pots, IV. and V. These pots had been kept in the hot-house, but from want of room were, whilst the plants were young, suddenly moved during very cold weather into the coldest part of the greenhouse. They all suffered greatly, and never quite recovered. After a fortnight only two of the nine self-fertilised seedlings were alive, whilst seven of the crossed survived. The tallest of these latter plants when measured was 47 inches in height, whilst the tallest of the two surviving self
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warm greenhouse. All three lots consequently suffered greatly, but the Chelsea-crossed plants much less than the other two lots. On the 3rd of October the Chelsea-crossed plants began to flower again, and continued to do so for some time; whilst not a single flower was produced by the plants of the other two lots, the stems of which were cut almost down to the ground and seemed half dead. Early in December there was a sharp frost, and the stems of Chelsea-crossed were now cut down; but on the
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bring the pollen-covered hairs, with which the lower surface is clothed, into contact with the stigma by either of which means self-fertilisation might be effected. Seeds from the above crossed and self-fertilised capsules, after germinating on bare sand, were planted in pairs on the opposite sides of five moderately-sized pots, which were kept in the greenhouse. The plants after a time appeared starved, and were therefore, without being disturbed, turned out of their pots, and planted in the
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thus differs from Ipom a, and almost certainly from Mimulus, as with these two species a cross between flowers on the same plant did no good. CALCEOLARIA. A bushy greenhouse variety, with yellow flowers blotched with purple. The flowers in this genus are constructed so as to favour or almost ensure cross-fertilisation;* and Mr. Anderson remarks that extreme care is necessary to exclude insects in order to preserve any kind true. He adds the interesting statement, that when the corolla is cut
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Early in the spring the plants were gradually hardened, and turned out of their pots into the open ground without being disturbed. By the end of August the greater number had formed fine heads, but several grew extremely crooked, from having been drawn up to the light whilst in the greenhouse. As it was scarcely possible to measure their heights, the finest plant on each side of each pot was cut down close to the ground and weighed. In the preceding table we have the result. The six finest
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of an inch. We thus see that reversion to a more natural condition acted more powerfully in favouring the ultimate growth of these plants than did a cross; but it should be remembered that the cross was with a semi-sterile variety having a feeble constitution. IBERIS UMBELLATA. Var. kermesiana. This variety produced plenty of spontaneously self-fertilised seed under a net. Other plants in pots in the greenhouse were left uncovered; and as I saw small flies visiting the flowers, it seemed
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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The average height of the seven crossed plants is here 19 12 inches, and that of the seven self-fertilised plants 16 39, or as 100 to 86. But as the plants on the self-fertilised side grew very unequally, this ratio cannot be fully trusted, and is probably too high. In both pots a crossed plant flowered before any one of the self-fertilised. These plants were left uncovered in the greenhouse; but from being too much crowded they were not very productive. The seeds from all seven plants of both
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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plants of the first generation. The crossed plants in the present case, as in the last, were more fertile than the self-fertilised, both lots being left unconvered in the greenhouse. The thirty crossed plants produced 103 seed-bearing flower-heads, as well as some heads which yielded no seeds; whereas the twenty-nine self-fertilised plants produced only 81 seed-bearing heads; therefore thirty such plants would have produced 83 7 heads. We thus get the ratio of 100 to 81, for the number of seed
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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yielded these capsules were grown in pots in the greenhouse, so that their absolute productiveness must not be compared with that of plants growing out of doors. The above three lots of seeds, viz., the self-fertilised, intercrossed,and English-crossed, were planted in an equal state of germination (having been as usual sown on bare sand) in nine large pots, each divided into three parts by superficial partitions. Many of the self-fertilised seeds germinated before those of the two crossed lots
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of their full size, and the crossed plants then averaged 3 87 inches, and the self-fertilised only 2 00 inches in height; or as 100 to 52. They were kept in the greenhouse, and did not grow vigorously. Whilst in flower they were again measured to the summits of their stems (see Table XLI.), with the following result: The average height of the fourteen crossed plants is here 5 58 inches, and that of the fourteen self-fertilised 2 37; or as 100 to 42. In four out of the five pots, a crossed plant
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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pot were measured to the tops of their stems. The pots were placed in the greenhouse, and the plants trained up sticks, so that they ascended to an unusual height. In three of the pots the crossed plants flowered first, but in the fourth at the same time with the self-fertilised. When the seedlings were between 6 and 7 inches in height, the crossed began to [page] 14
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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the pots, which were much too small, were kept in the greenhouse. The plants in consequence grew badly, and the self-fertilised suffered most in both pots. The two crossed plants when in flower during the following spring were 9 inches in height; one of the self-fertilised plants was 8, and the three others only 3 inches in height, being thus mere dwarfs. The two crossed plants produced thirteen pods, whilst the four self-fertilised plants produced only a single one. Some other self-fertilised
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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five crossed plants in 86 inches, and that of the five self-fertilised plants 82 35; or as 100 to 96. The pots were kept in the greenhouse, and there was little or no difference in the fertility of the two lots. Therefore as far as these few observations serve, the advantage gained by a cross is very small. * 'Kenntniss der Befruchtung,' 1844, pp. 573, 577. [page] 15
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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pollen from a distinct plant growing in the same row, and others were allowed to fertilise themselves under a net. The two lots of seeds thus obtained were sown on opposite sides of two large pots, but only four pairs came up at the same time. The pots were kept in the greenhouse. The seedlings of both lots when between 6 and 7 inches in height were equal. When nearly full-grown they were measured, as in the following table: Table LVII. Pisum sativum. No. of Pot. Crossed Plants. Self-fertilised
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green as those of the crossed seedlings. The pots were kept in the greenhouse, and as the plants in the following spring (1868) looked unhealthy and had grown but little, they were plunged, still in their pots, into the open ground. The plants all suffered much from the sudden change, especially the self-fertilised, and two of the latter died. The reminder were measured, and I give the measurements in the following table, because I have not seen in any other species so great a difference between
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flowers contained on an average 4 1 seeds, so that these were the most productive of all; and the seeds themselves looked finer even than those from the crossed perfect flowers. The seeds from the crossed perfect flowers and from the self-fertilised cleistogamic flowers were allowed to germinate on sand; but unfortunately only two pairs germinated at the same time. These were planted on the opposite sides of the same pot, which was kept in the greenhouse. In the summer of the same year, when the
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germinating on sand, were planted in pairs on the opposite sides of Pots II. and III., which were at first kept in the greenhouse and then turned out of doors. The plants were measured when in full flower. The following table, therefore, includes plants belonging to two generations. When the seedlings of the two lots were only 5 or 6 inches in height they were equal. In Pot III. one of the self-fertilised plants died before flowering, as has occurred in so many other cases. * The Composit are
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difference worth speaking of between the two lots in height; though other great advantages are derived, as we have seen, from cross-fertilisation. From being grown in pots and kept in the greenhouse, none of the plants produced any capsules. LOBELIA RAMOSA.* Var. Snow-flake. The well-adapted means by which cross-fertilisation is ensured in this genus have been described by several authors. The pistil as it slowly increases in length pushes the pollen out of the conjoined anthers, by the aid of a
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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; and they flowered so much earlier in all the pots, that the height of the flower-stems could be fairly compared only in Pots I. and II. * G rtner has shown that certain plants of Lobelia fulgens are quite sterile with pollen from the same plant, though this pollen is efficient on any other individual; but none of the plants on which I experimented, which were kept in the greenhouse, were in this peculiar condition. N 2 [page] 18
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, whence my plants originally came, a fresh plant differing in no respect from mine except in the colour of the flowers, which was a fine purple. But this plant must have been exposed during at least four generations to very different conditions from those to which my plants had been subjected, as these had been grown in pots in the greenhouse. Eight flowers on the self-fertilised plants in Table LXXXI., of the last or fourth self-fertilised generation, were fertilised with pollen from this fresh
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pot. The pots were kept in the greenhouse, and the plants trained up sticks. They were first measured to the tops of their stems when coming into flower; and the twenty-two Westerham-crossed plants then averaged 25 51 inches; the twenty-three intercrossed plants 30 38; and the twenty-three self-fertilised plants 23 40 inches in height. We thus get the following ratios: The Westerham-crossed plants in height to the self-fertilised as 100 to 91 The Westerham-crossed plants in height to the
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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of the intercrossed and self-fertilised plants. Relative Fertility of the Three Lots of Plants. None of the plants in pots in the greenhouse ever produced a capsule; and this may be attributed in chief part to the exclusion of moths. Therefore the fertility of the three lots could be judged of only by that of the plants growing out of doors, which from being left uncovered were probably cross-fertilised. The plants in the three rows were exactly of the same age and had been subjected to closely
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capsule for the crossed and self-fertilised flowers. The flowers hang downwards, and as the stigmas stand close beneath the anthers, it might have been expected that pollen would, have fallen on them, and that they would have been spontaneously self-fertilised; but these covered-up plants did not produce a single capsule. On some other occasions uncovered plants in the same greenhouse produced plenty of capsules, and I suppose that the flowers had been visited by bees, which could hardly fail to
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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. These plants, which were grown in pots in the greenhouse, were evidently in a very sterile condition, and the seeds in both sets of capsules, especially in the self-fertilised, although numerons, were of so poor a quality that it was very difficult to determine which were good and which bad. But as far as I could judge, the crossed capsules contained on an average 6 3 good seeds, with a maximum in one of thirteen; whilst the self-fertilised contained 6 05 such seeds, with a maximum in one of
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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-grandchildren of plants which had been legitimately fertilised), growing vigorously in pots in the greenhouse, were legitimately fertilised with pollen from an almost wild short-styled cowslip, and these flowers yielded some fine capsules. Thirty other flowers on the same illegitimate plants were fertilised with their own pollen, and these yielded seventeen capsules, containing on an average thirty-two seeds. This is a high degree of fertility; higher, I believe, than that which generally obtains with
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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numer of seedlings which thus perished, the surviving self-fertilised plants grew thinly in the row, and thus had an advantage over the crossed plants, which grew very thickly in the other row. The young plants in the two rows were protected by a little straw during the winter, and those in the two large pots were placed in the greenhouse. There was no difference between the two lots in the pots until the ensuing spring, when they had grown a little, and then some of the crossed plants were
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Darwin, C. R. 1878. The effects of cross and self fertilisation in the vegetable kingdom. 2d ed. London: John Murray.
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flowers, in number as 100 to 57 IBERIS UMBELLATA plants left uncovered in greenhouse; intercrossed plants of the 3rd generation, compared with self-fertilised plants of the 3rd generation, yielded seeds, in number as 100 to 75 IBERIS UMBELLATA plants from a cross between two varieties, compared with self-fertilised plants of the 3rd generation, yielded seeds, by weight as 100 to 75 PAPAVER VAGUM crossed and self-fertilised plants, left uncovered, produced capsules, in number as 100 to 99 ESCHSCHOLTZIA
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, Leptosiphon androsaceus (Polemoniace ). Plants under a net produced a good many capsules. Primula mollis (Primulace ). A homomorphic species, self-fertile: J. Scott, in 'Journal Linn. Soc. Bot.' vol. 1864, p. 120. Nolana prostrata. (Nolanace ). Plants covered up in the greenhouse, yielded seeds by weight compared with uncovered plants, the flowers of which were visited by many bees. in the ration of 100 ot 61. Ajuga reptans (Labiat ). Set a good many seeds; but none of the stems under a net
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from plants which had been raised from seed procured from a garden at chelsea. The Chelsea plants bore yellow flowers blotched with red, but differed in no other respect. They had been grown out of doors, whilst mine had been cultivated in pots in the greenhouse for the last eight generations, and in a different kind of soil. The seedlings raised from this cross with a wholly different stock may be called the Chelsea-crossed. The three lots of seeds thus obtained were allowed to germinate on
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-fertilised seedlings raised from the two plants, their heights have been added together in calculating the averages. I should state that by the accidental fall of a large bush in the greenhouse, several plants in both the series were much injured. These were at once measured together with their opponents and afterwards thrown away. The others were left to grow to their full height, and were measured when in flower. This accident accounts for the small height of some of the pairs; but as all the pairs
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40 as 100 to 54 Number and average height of all the plants of the ten generations 73 85 84 73 66 02 as 100 to 77 Mimulus luteus three first generations, before the new and taller self-fertilised variety appeard 10 8 19 10 5 29 as 100 to 65 Digitalis purpurea 16 51 33 8 35 87 as 100 to 70 Calceolaria (common greenhouse variety) 1 19 50 1 15 00 as 100 to 77 Linaria vulgaris 3 7 08 3 5 75 as 100 to 81 Verbascum thapsus 6 65 34 6 56 50 as 100 to 86 Vandellia nummularifolia crossed and self
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The Complete Work of Charles Darwin Online 4 Jan 29th /78 Phalaris canariensis Nice young seedlings raised in Greenhouse, exposed to light at 8° 30' a.m, removed at.— Day bright frosty. 5 painted with O. oil Lamp-black .25 down from tip all 5 perfectly vertical Oil 4 painted do .25 high from ground Lower part of all 4 quite vertical; upper unpainted part of 2 very slightly curved to light, but then with free seedlings curvature chiefly in lower part 2 (with narrow quills internally fastened
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CUL-DAR209.9.72-92
Note:
1878.02.00--1878.03.00
Radiation Cassia (various) / Cytisus / Oxalis (various) / Lotus
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(6 Radiation March 16 Cassia: (common-greenhouse Var.) A second Plant Exposed to temp of -3° -4 Cent for 30.' on early night of March 16. The 4 pinned out leaves with every leaflet quite dead both upper lower surfaces completely blackened. On whole large bush only 7 other large compound leaves blackened, only one A of the 7 with both surfaces blackened this was a younger leaf than any pinned down: The following case good [sketch] A B C D Branch z on branch (z) the uppermost young leaf A free
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CUL-DAR209.9.72-92
Note:
1878.02.00--1878.03.00
Radiation Cassia (various) / Cytisus / Oxalis (various) / Lotus
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11 Radiation 1878 Feb. 6 Melilotus italica 3 leaves pinned with upper surface (White W) 3 leaves with lower surface upwards (black wool.) — sky brilliantly clear at 5° 30' grass 30°; at 9° 30' temp 28° exposed from 5°. 30' to 10° 30' P.m. All six leaves much more injured, as seen next day in greenhouse, those that other upper leaves which went to sleep were nearly vertical, though some of the rather older leaves were a little spotted. Feb. 8th. The several leaves which were pinned open seem
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CUL-DAR68.58
Note:
1878.02.19--1878.03.05
Sea-kale / no bloom on cotyledons which keep closed for some days after
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sleep very late.— Put salt water on both Cots. of both seedlings. The Cots with salt opened in greenhouse in course of day. The above seedling did not go sleep. on night of 20', was observed at 10° 30' P.m at 5.15. a.m! Feb 22d 4° P.m put salt water on both Cots of silveryplant in the other pot. — Feb. 25' noon — The sea-kale plant look miserable March 5th Two quite dead, the third looks sickly Cease Mar 1
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from apogeotropism; but for For the first 11 hours it moved to a great distance greatly in a nearly straight line to one side from some another cause ; probably owing probably to its having been deflected by the light whilst standing in Greenhouse, had been drawn to one opposite side by light. At 7° 20' P.m on March 7' a fresh tracing was began continued during (see fig) for about 2 hours it followed nearly same course, then then bent to oth changed its course it a little; during the night it
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The Complete Work of Charles Darwin Online [47] Cyclamen Persica March 16' Pot with several pods bowed down toward ground laid horizontally in greenhouse the pods bowed soon bowed down at right angles to former course (is there not a note about this describing it more carefully???) so I assumed was geotropism— On 24th Pot still horizontal was turned upside down, so pods all pointed vertically up kept in dark cupboard in cool hot-house: by the Ap 1 ' 28th ie after one week hardly 4 days no or
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light greenhouse— on 29th the stem with pod curved down somewhat 2 stems with pods cut off considerably, but then doubt about their vigour. 31' rather more bowed down [48
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Lotus Jacobæus did they on following night of the 5th.— As temperature on the 4th. was generally 17 1/2 C, rather warmer than on previous days I doubt whether temperature was the cause: one more probably it was the want of sufficient illumination during several during the 3 previous days, as the Cots. zig-zagged all day growth had not ceased. During the 6th the pot was placed in greenhouse exposed to the sun at night the Cots slept pretty well, i.e. stood up at a high angle, but not I think so
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fertilisation in this genus, as has the Revd G. Henslow in the Journal of Linn. Socty, Bot. Vol IX, 1866, p. 327 and 355. Nicotiana tabacum (Solanaceæ) fully self-fertile. Ipomoea purpurea (Convolvulaceæ) highly self-fertile. Leptosiphon androsaceus (Polemoniaceæ). plants under a net produced a good many capsules. Nolana prostrata (Nolanaceæ), covered up plants in the greenhouse yielded seeds by weight compared with uncovered plants, the flowers of which were visited by many bees
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CUL-DAR67.53
Note:
1878.07.00--1878.12.00
Acacia retinoides 4 Phyllodia sponged both sides with tepid water both
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The Complete Work of Charles Darwin Online [53] Acacia retinoides 4 Phyllodia sponged both surface with tepid water both surfaces marked with black wool — on 3 main branches. Feb. 15. 1878 8˚ a.m. Recleaned both surfaces May 20th no effect. — Phyllodia are silvery both surface in water before they were sponged — Kept in greenhouse— growing vigorously July 11th 8˚ a.m. Green wool Branch — leaf with black wool, whole of lower surface up to apex thickly dotted with black smut — 2 or 3 dots on
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The Complete Work of Charles Darwin Online [81] Aug 1st 1878. Cassia f common Greenhouse var. floribunda? C. neglecta Pinched rubbed pulvinus blades of several leaves, but only one became slightly oblique this was a terminal leaflet of C. floribunda— So separate leaflets hardly sensitive— when leaf was shaken for 30 — The leaflets became somewhat depressed.— (C. neglecta did the same with leaflets shook whole seedlings, but no effect— some of leaflets in latter case slightly approached each
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previous generation. Nevertheless the self-fertilised seedlings were plainly inferior in height and vigour to the crossed. My attention was now thoroughly aroused, for I could hardly doubt that the difference between the two beds was due to the one set being the offspring of crossed, and the other of self-fertilised flowers. Accordingly I selected almost by hazard two other plants, which happened to be in flowers in the greenhouse, namely Mimulu
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The Complete Work of Charles Darwin Online (1 1878. Sept 26 Phalaris — Action of Light. after exposure to Darkness Light One pot of young seedlings all raised in darkness kept perfectly dark— another pot exposed yesterday in greenhouse, dark cloudy day, to light from above this morning bright to same light. — At 10. 20', both pots placed 38 inches before study N.E. window with linen muslin window blinds down.— a towel (bright-blue sky—) over case. — Sides blackened paper— roof back protected
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CUL-DAR209.7.133
Draft:
[1878].11.02--[1878].11.05
Tropaeolum majus / Draft of Cross and self fertilisation
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The Complete Work of Charles Darwin Online [133] Tropæolum majus— plants bow will lean to Light in greenhouse slew a little from them — reversed Pot placed before N.E. window — black on all other sides top. — filament with triangle fixed to penultimate internode— traced on Horizontal glass — Height of Plants? —Length of internodes? Distance from glass Left Hand [data not transcribed] Apheliotropism Right Hand [data not transcribed] (Used) [133v
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The Complete Work of Charles Darwin Online [116] Pea (Used) Top of petiole tied Dec. 29th 8° 20' [data not transcribed] 30th 6° 50 Kept in Study [sketch] Light in Hall Nice young trifid tendril — tip of longest branch in line with dot on top of stick. Horizontal glass— Kept dark above— N.E window with muslin blind— open to room behind — Clear morning — Plant placed in reversed position compared to position in greenhouse with respect to light [116v
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The Complete Work of Charles Darwin Online (1 In the greenhouse the cotyledons of numerous quite young seedlings were open in the middle of the day (Feb. 9th) formed together an angle, estimated at 30°—. At 6° P.m, when it was almost dark, they were nearly closed Perhaps introduce under Heliotrop— say here movement cd not be observed (2
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greenhouse, in 2 1/2 hours, though the day was very dull misty, the cotyledons of some few of the plants began to open, after an additional 1 1/2 hours they formed together were open to the extent of between 20° 30°. By this time the hypocotyls The stems of all the plants had become greatly bowed to the chief source of light: At 7° 30' P.m all the cotyledons were completely closed. On the following morning (Feb 12th) at 7° 30' a.m they were still closed, but at 9° partially open, by which time the sun
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CUL-DAR209.4.237-238
Draft:
[1878?].02.12--[1878?].02.13
Lychnis githago [with diagram] / Draft of Cross and self fertilisation.
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Lychnis githago (Used) Feb. 13' Cots open at 7° a.m. put in greenhouse till 1°, then in dark cupboard, not shut at all at 4° P.m. —5° perhaps a trace closed 6° not nearly shut 7. 20 very little shut 9° rather more shut than before, yet well open 10 40' P.m do. Feb 14' 6°. 50' Am well awake 10. 40. P.m. very much shut, but not quite closed Feb 15 7° a.m wide awake, but Cots never open horizontall
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CUL-DAR209.3.150-151
Draft:
[1878.06.?]13--[1878.06.?]16
Dianthus caryophyllus Carnation / Draft of Cross and self fertilisation, folio 641
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bristles which project from the anthers, and became dusted with pollen. Veronica agrestis (do) covered-up plants produced an abundance of seeds. I do not know whether any insects visit the flowers; but I have observed Syrphidæ repeatedly covered with pollen visiting the flowers of V. hederæfolia and chamoe ædrys. Mimulus luteus (do) highly self-fertile. Calceolaria (greenhouse var.) (do) do Verbascum thapsus (do) d
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CUL-DAR209.3.257-258
Draft:
[1878?].06.14--[1878?].06.16
Pelargonium zonale / Draft of Cross and self fertilisation, folio 642
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), covered plants produced some seeds, but the summer was wet and unfavourable. Galium aparine (Rubiaceæ), covered plants produced quite as many seeds as the uncovered. Apium petroselinum (Umbelliferæ), cov covered plants apparently were as productive as the uncovered. Zea mays (Gramineæ), a single plant in the greenhouse produced a good many grains
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The Complete Work of Charles Darwin Online (1 Vicia faba (Circumnutation of Leaf) Plant 18 inches high in pot, drawn up by having gone having grown in greenhouse.— Young leaf, measured to end of terminal leaflet 3.1 inch – with filament affixed to one midrib of one of 2 terminal leaflets: stem [sketch] secured to stick at base of leaf— observed under skylight. Temp. between 17° 18° C. See tracing — Apex of leaflet 7 1/4 from vertical glass.— Fell in morning — then rose greatly falling down to
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CUL-DAR67.57-58
Note:
1879
Trifolium resupinatum / 10h 30 a.m a little Plant with 5 leaves 3 long &
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Dec Jan. 7th 1879. Trifolium resupinatum. 10˚ 35' a.m. Plants growing — rather young in greenhouse, brought into study. 4 leaves white thread, bloom unbloomed have painted of opposite lateral leaflets painted with olive oil Lamp-black. — 10˚ P.m— All the painted ones went to sleep Jan 9th 8˚ 30' — There is no difference in the degree to which the bloomed unbloomd halves have absorbed the oil 10th 9˚ 15' a m Certainly no difference in degree to which grease has been absorbed by the two halve
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CUL-DAR157.40
Draft:
[1879--1880]
Draft of Movement in plants, Erythrina sleep, folio 506.
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the great nocturnal fall commenced. On the third day, the movement was equally great in amplitude, but was remarkably simple, for the leaflet rose in an almost perfectly straight line from 6 50 am to 3 P.m, then sank down, in a straight line until vertically dependent asleep. Apios tuberosa Phaseolus vulgaris: the leaflets of both these plants sink vertically down at night. The petiole of the a young leaf of the latter plant, cultivated in a pot in the greenhouse rose 16°, of an older leaf 10° at
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each leaflet could would necessary be 120° or a 1/3 of 360°; but as the vertical edges depend petiole this is not case as leaflets. No, no, this make angle bigger— There must be [sketch] space on summit, they must fold backwards. [sketch] In Ox. fragrans, the lower surfaces do not come into contact, but are separated by rather wide space, so that in greenhouse the summit of petioles look like bi 6 leaflets radiating from summit, Here the leaflets so much folded that in 2 of best [85v
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The Complete Work of Charles Darwin Online [page] [10] April. 1879 6 or 7' Radicles cot. open. 8' 10' P.m. not asleep. Red Cabbage 8th open this morning, well asleep at 9° 45' P.m. 9th 10° P.m transplanted into [pan] very slightly asleep. 10th somewhat raised, but doubtfully asleep. 15th Hardly a sign of sleep has been very cold weather 18th no sign of sleep (I may say that from a few observations we doubt whether sleep as long out of doors as in greenHouse
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The Complete Work of Charles Darwin Online [65] N.B. Lathyrus aphaca — I can detect no apheliotropism in tendrils, yet they circumnutate very little — top of stem is highly heliotropic — Nor can I in Greenhouse see any signs in L. maritimus.— Sept 2d. 1879
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The Complete Work of Charles Darwin Online [39] Lupinus luteus Sept. 2d— 1879 Lots of Plants Kept in greenhouse— brought into Study do not sleep well— yet enough movement to show that it is base of leaflets which move.— The exterior longer leaflets now have more much more power of movement than the shorter ones, which retained at night their diurnal position. From such complex twisting movements , the base of leaflets, as might have been expected are provided with a pulvinus Sept. 11th one pot
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The Complete Work of Charles Darwin Online [56] 1879 Hedera Helix Glass-filament with triangle fixed perpendicularly Branches of some young plants during middle of summer were found to be strongly apheliotropic sent branches directly away from light. On Sept. 2d these same plants had 2 trailing branches tied up perpendicularly, exposed to N.E. window were both found for first 4 days to be decidedly heliotropic. (so were several other plants in greenhouse; some of latter afterwards became
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The Complete Work of Charles Darwin Online [21] [in margin:] Sleep Coronilla rosea— pot in greenhouse. Sept 14. 1879— Leaves with many opposite leaflets— which are extended horizontally during day with their midribs at rt. angles to petiole. at night rise up so as to come nearly with younger leaves in close contact, at the same time they bend back toward base of petiole, so that these midribs now form angles of between 40° 50° with main petiole in a vertical plane, instead of angle of about 90
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CUL-DAR209.3.47-52
Note:
1879.11.02
(Miscellaneous): Carnation-tree / Petunia violacea / cabbage / Vicia faba
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The Complete Work of Charles Darwin Online [47] Shoots tied Horizontally in greenhouse (Cold water Feb 5th 12° 45 Red cords Black dots 5 1/2 c.t from tip, to show whether grow Tips red — ⨀ — apart — ⨀ (2 innermost leaves now in contact touched with red.) Feb 19th. 12.15 tips apart ⨀ —— ⨀ increase Feb 5' 12° 45' (String) Tips red ⨀ —— ⨀ apart (2 innermost leaves now in contact touched with red) 9th. 12° 15 tips apart ⨀ —— ⨀ increase Feb. 9th no apogeotropism Carnation-Tree. [47v
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(5) 9th noon — Purple thread open less than rectangle (6) 9th noon Black wool. open less than rectangle. Jan 9th 10° P.m all 6 seedlings with Cots. shut nicely closely; but then only risen less less than 45° well-watered. Jan 10' Placed in greenhouse; half dry in sun, half behind pot, but Cots. did not open more very few as much as rectangle. 9° P.m perpendicular, but tips of purple thread, black wool one of thick cotton thread. reflexed Jan 11th. 10° 30' P.m none of the 6 hardly at all closed
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F1280
Book:
Darwin, C. R. 1880. The different forms of flowers on plants of the same species. London: John Murray. 2s edition.
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tilised with their own pollen, produced 15 capsules, containing an average of 16 3 seeds. As already stated, the probable average with legitimate plants for a union of this nature is rather above 20 seeds. These plants were remarkably healthy and vigorous, as long as they were kept under highly favourable conditions in pots in the greenhouse; and such treatment greatly increases the fertility of the cowslip. When these same plants were planted during the next year (which, however, was an
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F1325
Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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germinating at the proper season, would not be subjected to darkness at this hour in the morning. The above amount of movement of the cotyledons is only temporary, lasting with plants kept in a warm greenhouse from four to six days; how long it would last with seedlings growing out of doors we do not know. Raphanus sativus. In the middle of the day the blades of the cotyledons of 10 seedlings stood at right angles to their hypocotyls, with their petioles a little divergent; at night the blades
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Book:
Darwin, C. R. 1880. The different forms of flowers on plants of the same species. London: John Murray. 2s edition.
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. These whilst still in the greenhouse produced smaller leaves and shorter flower-stalks than some legitimate plants with which they grew in competition; but it should be observed that the latter were the product of a cross with a fresh stock, a circumstance which by itself would have added much to their vigour.* When these illegitimate plants were transferred to fairly good soil out of doors, they became during the two following years much more dwarfed in stature and produced very few flower-stems
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Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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leaves were in a flaccid condition. There were about 150 free leaves on the plant, and none of these were injured, except 2 or 3 very young ones. But after two days, the plants having been brought back into the greenhouse, the 6 pinned-out leaves all recovered. Melilotus Taurica. Several plants were exposed for 5 h. during two nights to a clear sky and slight frost, accompanied by some wind; and 5 leaves which had been pinned out suffered more than those both above and below on the same branches
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surrounding legitimate plants, in the [words excised] It is [words excised] certain [rest excised] [Forms of flowers, p. 234: These whilst still in the greenhouse produced smaller leaves and shorter flower-stalks than some legitimate plants with which they grew in competition; but it should be observed that the latter were the product of a cross with a fresh stock,—a circumstance which by itself would have added much to their vigour.* When these illegitimate plants were transferred to fairly
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F1325
Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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Cycas pectinata (Cycadeæ). The large seeds of this plant in germinating first protrude a single leaf, which breaks through the ground with the petiole bowed into an arch and with the leaflets involuted. A leaf in this condition, which at the close of our observations was 2 1/2 inches in height, had its movements traced in a warm greenhouse by means of a glass filament bearing paper triangles attached across its tip. The tracing (Fig. 45) shows how large, complex, and rapid were the
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Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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the greenhouse became bent in the middle of the summer at right angles from the light. On Sept. 2nd some of these stems were tied up so as to stand vertically, and were placed before a north-east window; but to our surprise they were now decidedly heliotropic, for during 4 days they curved themselves towards the light, and their course being traced on a horizontal glass, was strongly zigzag. During the 6 succeeding days they circumnutated over the same small space at a slow rate, but there could
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Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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could be plainly seen when a straight narrow black line had been painted along it during the day. The two terminal leaflets in the daytime include rather less than a right angle; but their divergence increases greatly whilst they * I am informed by Mr. Dyer that Mr. Bentham believes that C. floribunda (a common greenhouse bush) is a hybrid raised in France, and that it comes very near to C. laevigata. It is no doubt the same as the form described by Lindley ('Bot. Reg.,' Tab. 1422) as C
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Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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(?) (cultivated var.) (Tropaeoleæ). Several plants in pots stood in the greenhouse, and the blades of the leaves which faced the front-lights were during the day highly inclined and at night vertical; whilst the leaves on the back of the pots, though of course illuminated through the roof, did not become vertical at night. We thought, at first, that this difference in their positions was in some manner due to heliotropism, for the leaves are highly heliotropic. The true explanation, however, is
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F1280
Book:
Darwin, C. R. 1880. The different forms of flowers on plants of the same species. London: John Murray. 2s edition.
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cowslips of both forms, protected from insects in my greenhouse, did not set one pod, though artificially fertilised flowers in other pots produced an abundance. We thus see that the visits of insects are absolutely necessary for the fertilisation of Primula veris. If the corolla of the long-styled form had dropped off, instead of remaining attached in a withered state to the ovarium, the anthers attached to the lower part of the tube with some pollen still adhering to them would have been
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Book:
Darwin, C. R. 1880. The different forms of flowers on plants of the same species. London: John Murray. 2s edition.
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in the spring of 1862 forty flowers were fertilised at the same time in both ways. The plants were accidentally exposed in the greenhouse to too hot a sun, and a large number of umbels perished. Some, however, remained in moderately good health, and on these there were twelve flowers which had been fertilised legitimately, and eleven which had been fertilised illegitimately. The twelve legitimate unions yielded seven fine capsules, containing on an average each 57 3 good seeds; whilst the eleven
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Book:
Darwin, C. R. 1880. The different forms of flowers on plants of the same species. London: John Murray. 2s edition.
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very rich soil, some in stiff poor clay, some in old peat, and some in pots in the greenhouse; so that these plants, 765 in number, as well as their parents, were subjected to diversified and un * I have given instances in my work 'On the Variation of Animals and Plants under Domestication,' ch. xv. 2nd edit. vol. ii. p. 69. [page] 6
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Book:
Darwin, C. R. 1880. The different forms of flowers on plants of the same species. London: John Murray. 2s edition.
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form of a cap. These cleistogamic flowers produced an abundance of seed. Later in the season perfect flowers appeared. With plants in a state of nature the flowers open only in the early morning, as I have been informed by Mr. Wallis, who particularly attended to the time of their flowering. In the case of D. Anglica, the still folded petals on some plants in my greenhouse opened just sufficiently to leave a minute aperture; the anthers dehisced properly, but the pollen-grains adhered in a mass
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Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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described were directed to nearly the same point; so that more light seemed to have come through the glass roof of the greenhouse on one side than on any other. All five tracings resembled each other to a certain extent, and it will suffice to give two of them. In A (Fig. 48) the seedling was only .45 of an Fig. 48. Asparagus officinalis: circumnutation of plumules with tips whitened and marks placed beneath, traced on a horizontal glass. A, young plumule; movement traced from 8.30 A.M. Nov. 30th
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Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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during cold weather in a sitting-room, and others in a greenhouse, but the temperature was not recorded. These six striking cases almost convinced us that the apex was sensitive, but of course we determined to make many more trials. As we had noticed that the radicles grew much more quickly when subjected to considerable heat, and as we imagined that heat would increase their sensitiveness, vessels with germinating beans suspended in damp air were placed on a chimney-piece, where they were subjected
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Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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18 inches in height was observed. The bead changed its course greatly eleven times in the course of 10 h. 30 m. (Fig. 75), and there could be no doubt about the circumnutation of the stem. Fig. 75. Deutzia gracilis: circumnutation of stem, kept in darkness, traced on horizontal glass, from 8.30 A.M. to 7 P.M. March 20th. Movement of bead originally magnified about 20 times, here reduced to half scale. (9.) Fuchsia (greenhouse var., with large flowers, probably a hybrid) (Onagrarieae, Fam. 100
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Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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having been removed a few days before from the greenhouse to the hot-house, was observed with especial interest, as it seemed so little probable that the stem would circumnutate. The branches are flat, or flabelliform; but some of them are triangular in section, with the three sides hollowed out. A branch of this latter shape, 9 inches in length and 1 1/2 in diameter, was chosen for observation, as less likely to circumnutate than a flabelliform branch. The movement of the bead at the end of the
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Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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growing bush, at an angle of 35 above the horizon, was selected for observation. For the first 11 h. it moved to a considerable distance in a nearly straight line to one side, owing probably to its having been previously deflected by the light whilst standing in the greenhouse. At 7.20 P.M. on March 7th a fresh tracing was begun and continued for the next 43 h. 40 m. (see Fig. 80). During the first 2 h. it followed nearly the same direction as before, and then changed it a little; during the night
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Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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the frond was placed near to the vertical glass, and would probably have been greater and more rapid had the day been warmer. For the plant was brought out of a warm greenhouse and observed under a skylight, where the temperature was between 15 and 16 C. We have seen in Chap. I. that a frond of this Fern, as yet only slightly lobed and with a rachis only .23 inch in height, plainly circumnutated.* Fig. 119. Nephrodium molle: circumnutation of rachis, traced from 9.15 A.M. May 28th to 9 A.M. 29th
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Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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: Erythrina crista-galli, out of doors and nailed against a wall, seemed in fairly good health, but the leaflets did not sleep, whilst those on another plant kept in a warm greenhouse were all vertically dependent at night. In a kitchen-garden the leaflets of Phaseolus vulgaris did not sleep during the early part of the summer. Ch. Royer says,‡ referring I suppose to the native plants in France, that they do not sleep when the temperature is below 5 C. or 41 F. In the case of several sleeping
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Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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and albifrons, though observed out of doors and in the greenhouse, did not change the position of their leaves sufficiently at night to be said to sleep. From observations made on two sleeping species, it appears that, as with Tropæolum majus, the leaves must be well illuminated during the day in order to sleep at night. For several plants, kept all day in a sitting-room with north-east windows, did not sleep at night; but when the pots were placed on the following day out of doors, and were
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Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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sommet, les inférieures vers la base du petiole commun; but the leaflets on a young plant observed by us in the greenhouse merely sank vertically downwards at night. The leaflets are raised in Sphaerophysa salsola, Colutea arborea, and Astragalus uliginosus, but are depressed, according to Linnaeus, in Glycyrrhiza. The leaflets of Robinia pseudo-acacia likewise sink vertically down at night, but the petioles rise a little, viz., in one case 3 , and in another 4 . The circumnutating movements
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Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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of the secondary trifoliate leaves sink vertically at night. On young plants of Sida rhombaefolia, only a few inches in height, the leaves did not sleep, though on rather older plants they rose up vertically at night. On the other hand, the leaves on very young plants of Cytisus fragrans slept in a conspicuous manner, whilst on old and vigorous bushes kept in the greenhouse, the leaves did not exhibit any plain nyctitropic movement. In the genus Lotus the basal stipule-like leaflets rise up
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Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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that which they have previously received from above and on all sides. Analogous facts have been given with respect to the nyctitropic and periodic movements of plants. Of two pots containing seedlings of Phalaris which had germinated in darkness, one was still kept in the dark, and the other was exposed (Sept. 26th) to the light in a greenhouse during a cloudy day and on the following bright morning. On this morning (27th), at 10.30 A.M., both pots were placed in a box, blackened within and open
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Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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rise did not coincide with that of the great diurnal fall, so that narrow ellipses were described, as is the usual rule with circumnutating organs. The cotyledons are provided with a pulvinus, and its development will hereafter be described. Mimosa pudica (Leguminosæ). The cotyledons rise up vertically at night, so as to close together. Two seedlings were observed in the greenhouse (temp. 16 to 17 C. or 63 to 65 F.). Their hypocotyls were secured to sticks, and glass filaments bearing little
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Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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3 to 6 P.M. descended; whereas cotyledons which have been exposed all the day to the light rise in the evening so as to stand vertically at night; but this statement applies only to young seedlings. For instance, six seedlings in the greenhouse had their cotyledons partially open for the first time on the morning of November 15th, and at 8.45 P.M. all were completely closed, so that they might properly be said to be asleep. Again, on the morning of November 27th, the cotyledons of four other
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Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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different directions, there could be no doubt that the arched hypocotyl circumnutated. At the close of the 34 h. the upper part began to rise and straighten itself, dragging the cotyledons out of the ground, so that the movements of the bead could no longer be traced on the glass. Quercus (American sp.) (Cupuliferæ). Acorns of an American oak which had germinated at Kew were planted in a pot in the greenhouse. This transplantation checked their growth; but after a time one grew to a height of five
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Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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A filament was next fixed to a cotyledon only 1/4 of an inch in height, which was illuminated exclusively from above, and as it was kept in a warm greenhouse, it grew rapidly; and now there could be no doubt about its circumnutation, for it described a figure of 8 as well as two small ellipses in 5 1/2 hours. Nephrodium molle (Filices). A seedling fern of this species came up by chance in a flowerpot near its parent. The frond, as yet only slightly lobed, was only .16 of an inch in length and
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Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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covered with between 300 and 400 leaves, had been kept all winter in the greenhouse. Seven leaves were pinned horizontally open, and were exposed on March 16th for 2 h. to a clear sky, the temperature on the surrounding grass being -4 C. (24 to 25 F.). Next morning all seven leaves were found quite killed, so were many of the free ones which had previously gone to sleep, and about 100 of them, either dead or browned and injured were picked off. Some leaves showed that they had been slightly injured
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Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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whilst some, but not many, of those which had formerly stood facing the back of the room and which had never before been well illuminated or gone to sleep, now assumed a vertical position at night. On the next day (12th) the plant was turned round into its original position, so that the same leaves faced the light as formerly, and these now went to sleep in the usual manner. We will only add that with some young seedlings kept in the greenhouse, the blades of the first pair of true leaves (the
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Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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Apios tuberosa (Tribe 8). The leaflets sink vertically down at night. Phaseolus vulgaris (Tribe 8). The leaflets likewise sink vertically down at night. In the greenhouse the petiole of a young leaf rose 16 , and that of an older leaf 10 at night. With plants growing out of doors the leaflets apparently do not sleep until somewhat late in the season, for on the nights of July 11th and 12th none of them were asleep; whereas on the night of August 15th the same plants had most of their leaflets
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Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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movement of this kind in one member of the genus. Again, with Chenopodium album (Chenopodieæ), the upper young leaves of some seedlings, about 4 inches in height, were horizontal or sub-horizontal during the day, and at 10 P.M. on March 7th were quite, or almost quite, vertical. Other seedlings raised in the greenhouse during the winter (Jan. 28th) were observed day and night, and no difference could be perceived in the position of their leaves. According to Bouché ('Bot. Zeitung,' 1874, p. 359) the
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Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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upright flower-heads, arising from branches in all sorts of positions, on plants growing in a warm greenhouse, were marked with thread, and after 24 h. six of them were vertically dependent; these therefore had travelled through 180 in this time. Ten were extended sub-horizontally, and these had moved through about 90 . Three very young peduncles had as yet moved only a little downwards, but after an additional 24 h. were greatly inclined. At the time when the flower-heads reach the ground
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Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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occasionally observed and was less strongly marked. It should be stated that these peduncles were observed under a double skylight in the house, and that they generally moved downwards very much more slowly than those on plants growing out of doors or in the greenhouse. Fig. 191. Trifolium subterraneum: circumnutating movement of peduncle, whilst the flower-head was burying itself in sand, with the reflexed tips of the calyx still visible; traced from 8 A.M. July 26th to 9 A.M. on 27th. Glass
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Book:
Darwin, C. R. 1880. The different forms of flowers on plants of the same species. London: John Murray. 2s edition.
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, and the female organs of the long-styled form in a more or less complete state, they spontaneously produced a surprising number of capsules, which generally contained a large average of remarkably fine seeds. With ordinary cowslips, legitimately fertilised, I once obtained from plants cultivated in the greenhouse the high average, from seven capsules, of 58 7 seeds, with a maximum in one capsule of 87 seeds; but from plants grown out of doors I never obtained a higher average than 41 seeds. Now
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Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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during the winter in the greenhouse, was exposed during 5 h. at night to a slight frost and clear sky. Four leaves had been pinned out, and these died after a few days; but so did many of the free leaves. Therefore nothing certain could be inferred from this trial, though it indicated that the horizontally extended leaves suffered most. Another large pot with many plants was next exposed for 1 h., the temperature on the surrounding grass being lower, viz., 3 to -4 C. Ten leaves had been pinned out
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Book:
Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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the early spring from shoots on some cut-down plants in the greenhouse, slept in a totally different manner from the normal one; for the three leaflets, instead of twisting on their own axes so as to present their lateral edges to the zenith, turned upwards and stood vertically with their apices pointing to the zenith. They thus assumed nearly the same position as in the allied genus Trifolium; and on the same principle that embryological characters reveal the lines of descent in the animal
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Darwin, C. R. 1880. The power of movement in plants. London: John Murray.
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traced from 6.40 A.M. June 8th, to 8 A.M. on the 10th. In order to observe the nyctitropic movements of this plant, it is necessary that it should have grown in a warm greenhouse, for out of doors in our climate it does not sleep. We see in the tracing (Fig. 152) that the leaflet oscillated twice up and down between early morning and noon; it then fell greatly, afterwards rising till 3 P.M. At this latter hour the great nocturnal fall commenced. On the second day (of which the tracing is not
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F2095
Book contribution:
Candolle, Alphonse de. 1887. [Recollection of Darwin]. In Bettany, G. T. 1887. Life of Charles Darwin. London: Walter Scott, pp. 148-150.
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remain of the former labours of the owner. Darwin used simple means. He was not one who would have demanded to have palaces built in order to accommodate laboratories. I looked for the greenhouse in which such beautiful experiments on hybrid plants had been made. It contained only a vine. One thing struck me, although it is not rare in England, where animals are loved. A heifer and a colt were feeding close to us with the tranquillity which tells of good masters, and I heard the joyful barking of
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The Complete Work of Charles Darwin Online [251] Ap. 20th 80/ Trifolium subterraneum— Geotropism. Thread were tied on 19 flowers, all placed on upright trailing branches all directed vertically upwards in the complete darkness in greenhouse after 24 hr of these 6 had become about quite or almost vertically down had moved 180° — so were nearly horizontal so had had moved about 90°, 1 was declined about 45° from the vertical 2 alone were still vertical. After additional 24h the 45° was almost
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CUL-DAR209.5.155-156
Draft:
[1880].04.24
Radicles of Peas with squares of card gummed to tips / Draft of Forms of flowers.
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ratio only of 5 to 100! There can therefore be no doubt that illegitimate fertilisation continued during successive generation affecting the powers of growth fertility of t P. veris to an extraordinary degree; more especially when the plants are exposed to th ordinary conditions of life, instead of being protected in a greenhouse. (one line open
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CUL-DAR67.60-64
Note:
1880.12.24--1881.01.27
Commelina coelestis / When seeds germinate — Cot? projects first -
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It is curious that to the Cotyledon never become green the distal part f serves slowly solely to convey nutriments from the seed to the seedlings Jan 11' The Cots in Greenhouse have become pale green. — In seedling with 2 leaves seed-coats almost empty with knob within still white: I doubt whether knob can ever escape from the seed-coats:) no sign of tube in radicle — but base of Cot. looks a little thickened: see to this. — (
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F3396
Book contribution:
Darwin, C. R. 1884. [Letter extract from 1881 and recollection of Darwin's words]. In R. Meldola, The presidential address: Darwin and modern evolution. Transactions of the Essex Field Club 3: 64-93.
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continued his studies in this country, it was always living animals and plants that were appealed to when possible; his most important experiments and observations were made upon materials growing in his greenhouse or garden, or were collected from the surrounding country; and in his broader generalisations, which necessitated a wider survey of facts, he always made use of the observations of those to whom Nature had spoken face to face. Even when failing health compelled him to abandon the great
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A673
Book contribution:
Richter, Hans. 1882. [English translation of recollection of Darwin in 1881]. In Otto Zacharias, Charles R. Darwin und die culturhistorische Bedeutung seiner Theorie vom Ursprung der Arten. Berlin: E. Staude, pp. 5-6. Translated for Darwin Online by Anders Hansson.
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of various sorts and a large greenhouse. The garden turns into a deer park, an actual forest with lush meadows that stretch along a hillside in refreshing verdure. Once we had returned, we met Darwin in the dining room and on his request I played the piano again. He complained that he had been forced to suspend his work, the doctors gave him no peace, he must go to the sea, and he was completely inconsolable that he was not allowed to work there. On his farewell he promised to come to London
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CUL-DAR67.76-77
Note:
1881.07.25--1882.01.07
Acacia lophantha / 12h 15 p.m The 6 pairs of terminal leaflets on the 2
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The Complete Work of Charles Darwin Online [76] 1881. Acacia lophantha July 25' 12˚ 15' P.m. The 6 pairs of terminal leaflets on the 2 terminal pinnæ 6 pairs on the next pinna on moderately young leaf in cool greenhouse, was sponged with water at about 96° F, so that drops wd adhere; but the surface was still a little greasy, so that water aggregated readily, but yet drops stuck well on in manner in which it wd not do on unwashed leaves. These 18 leaflets had drops on, occasionally wet
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CUL-DAR67.86-87
Note:
1881.07.25--1881.10.11
Robinia pseudo-acacia / 12h 15 pm Plant in pot in cool greenhouse growing
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The Complete Work of Charles Darwin Online [86] 1881 Robinia pseudo-acacia.— July 25' 12˚ 15 P.m.— Plant in pot in cool greenhouse, growing well — 3 leaflets on very young, but nearly full size, on uppermost leaf, marked with white thread. These 3 opposite ones — sponged with water about 96°, till bloom removed, but they yet remained very greasy. — Also 4 leaflets, on Right side (viewed from apex) 3d leaf from apex, similarly treated, as well as the opposite ones, making 7 with drops 7 without
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CUL-DAR67.82-83
Note:
1881.07.26--1881.08.14
Clarkia elegans / Dark red anthers with red filaments exterior face
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In the genus Enchiridium, which is close to Clarkia there are 4 stamens, so 4 quite aborted — In several other allied genera there are 8 stamens. Clarkia plants in pots in Greenhous
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CUL-DAR67.78-79
Note:
1881.07.26--1881.10.28
Cassia floribunda / Drops put on at 11h am / 2 pairs of young tender
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The Complete Work of Charles Darwin Online [78] 1881. Cassia floribunda July 26 Drops put on at 11˚. a.m. 2 pair of young tender leaflets fully expanded on uppermost leaf not fully expanded — 2 pairs on 2d leaf almost fully expanded 5 pair of leaflets on 3d fully expanded leaf all on right side, viewed from apex — Sponged well with water at 96°. Middle portion of leaf near midrib extremely greasy — Plant in cool greenhouse Leaves quite free not pinned or covered Drops allowed often to dry
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CUL-DAR52.F26-F44
Note:
1881.09.04--1881.10.02
Aggregation / Geranium [application of carbonate of ammonia, alcohol,
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aggregation, much pulpy green matter. After alcohol, a good many, but not full complement of Chloro-grains visible; these generally heaped together, I think sometimes confluent; often indistinct. — Sept. 16th a leaf with horse-shoe mark from greenhouse was put into sol (4 to 1000) at 12˚ 45 15th, examined at 11˚ on 16th — no true aggregation, but in very many pallisade cells, bodies of the most diversified shapes, many [sketch] many like this [sketches], apparently denser fluid, with the Chlo
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12% |
CUL-DAR52.F26-F44
Note:
1881.09.04--1881.10.02
Aggregation / Geranium [application of carbonate of ammonia, alcohol,
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epidermis! (Sept 17th The latter slice was left in the iodide all night, there are yet some spheres in the palissade cells in the epidermis.) Sept 17. The 2 foregoing leaves which had been immersed in 4 to 1000 on the 15' at 12˚ 45, reexamined this morning— The Greenhouse leaf p. L now with abundant spheres in epidermis cells — swollen chloro. grains perh a few spheres— left in alcohol till colourless the spheres in epidermis certainly not acted on some spheres in parenchyma, which do not look like
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The Complete Work of Charles Darwin Online [76] Roots Dec. 29th Solanum (Empress) greater hardy sp which produces large berry woody greenhouse young plant — 4 roots were immersed for 20 1/2° 21° in sol of 4 to 1000— in one of these [illeg] In cells lying immediately beneath epidermis for some distance some way up root [cut visible] minute aggregation mass only of some granular sometimes hyaline brown matter. — Moreover In many of isolated elongated cells near vascular bundle isolated dark
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F839
Book:
Darwin, C. R. 1882. The movements and habits of climbing plants. London: John Murray.
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shoots, cut off a plant, and placed in water made circles in 3 hrs. 10 m., 5 hrs. 20 m., and 7 hrs. 15 m. Wistaria Chinensis (Leguminos ), in greenhouse, moves against the sun. H. M. May 13, 1st circle was made in 3 5 May 13, 2nd circle was made in 3 20 May 16, 3rd circle was made in 2 5 May 24, 4th circle was made in 3 21 May 25, 5th circle was made in 2 37 May 25, 6th circle was made in 2 35 Phaseolus vulgaris (Leguminos ), in greenhouse, moves against the sun. H. M. May, 1st circle was made in 2
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F839
Book:
Darwin, C. R. 1882. The movements and habits of climbing plants. London: John Murray.
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weighing only 82 of a grain (53 14 mg.) We have seen that the petioles of some other leaf-climbing plants are affected by one-thirteenth of this latter weight. In this species, and in no other leaf-climber seen by me, a full-grown leaf is capable of clasping a stick; but in the greenhouse the movement was so extraordinarily slow that the act required several weeks; on each succeeding week it was clear that the petiole had become more and more curved, until at last it firmly clasped the stick
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F839
Book:
Darwin, C. R. 1882. The movements and habits of climbing plants. London: John Murray.
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Asparagus (unnamed species from Kew) (Liliace ) moves against the sun, placed in hothouse. H. M. Dec. 26, 1st circle was made in 5 0 Dec. 27, 2nd circle was made in 5 40 Tamus communis (Dioscoreace ). A young shoot from a tuber in a pot placed in the greenhouse: follows the sun. H. M. July, 7, 1st circle was made in 3 10 July 7, 2nd circle was made in 2 38 July 8, 3rd circle was made in 3 5 July 8, 4th circle was made in 2 56 July 8, 5th circle was made in 2 30 July 8, 6th circle was made in 2
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F839
Book:
Darwin, C. R. 1882. The movements and habits of climbing plants. London: John Murray.
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Shoots of the Kidney-bean, cut off and placed in water, were similarly retarded, but in a less degree. I have repeatedly observed that carrying a plant from the greenhouse to my room, or from one part to another of the greenhouse, always stopped the movement for a time; hence I conclude that plants in a state of nature and growing in exposed situations, would not make their revolutions during very stormy weather. A decrease in temperature always caused a considerable retardation in the rate of
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F839
Book:
Darwin, C. R. 1882. The movements and habits of climbing plants. London: John Murray.
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hrs. 26 m. The shoots, however, sometimes stood still. It is considered a greenhouse plant; but when kept there, the petioles took several days to clasp a stick: in the hothouse a stick was clasped in 7 hrs. In the greenhouse a petiole was not affected by a loop of string, suspended during several days and weighing 2 grains (163 mg.); but in the hothouse one was made to curve by a loop weighing 1 64 gr. (106 27 mg.); and, on the removal of the string, it became straight again. Another petiole
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F839
Book:
Darwin, C. R. 1882. The movements and habits of climbing plants. London: John Murray.
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only one-eighth of a grain (8 1 mg.) sometimes acted and sometimes did not act. The sensitiveness extends from the blade of the leaf to the stem. I may here state that I ascertained in all cases the weights of the string and thread used by carefully weighing 50 inches in a chemical balance, and then cutting off measured lengths. The main petiole carries three leaflets; but their short, sub-petioles are not sensitive. A young, inclined shoot (the plant being in the greenhouse) made a large circle
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F1800
Periodical contribution:
Darwin, C. R. 1882. The action of carbonate of ammonia on the roots of certain plants. [Read 16 March] Journal of the Linnean Society of London (Botany) 19: 239-261, 2 text figures.
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roots of a fern, Nephrodium molle, were immersed for 20 hours in a solution of 4 to 1000; and this caused the deposition of some brown granular matter in the cells near their tips; and more or less confluent globules could be seen in the underlying parenchyma-cells. So it was with an unnamed greenhouse species of fern; and in this case the almost loose cells of the root-cap contained brown granules. The roots of a Ranunculus (R. acris ?) similarly treated exhibited near their tips brown granular
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F1800
Periodical contribution:
Darwin, C. R. 1882. The action of carbonate of ammonia on the roots of certain plants. [Read 16 March] Journal of the Linnean Society of London (Botany) 19: 239-261, 2 text figures.
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were numerous; but not one was seen which arose from a cell containing granules. Roots of Oxalis sepium, comiculata, and of a greenhouse species with small yellow flowers were immersed in a solution of 7 to 1000, and granular matter was deposited in the layer of cells underlying the exterior layer. This occurred in the case of O. sepium in 20 minutes. With O. corniculata the cells with granules were isolated—that is, did not form rows; and the granules were either brown or of a bluish-green
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A314
Pamphlet:
Miall, L. C. 1883. The life and work of Charles Darwin: a lecture delivered to the Leeds Philosophical and Literary Society, on February 6th, 1883. Leeds: Richard Jackson.
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researches. Darwin used simple means. I looked for the greenhouse, in which those beautiful researches on vegetable hybrids had been made; it contained nothing but a vine. One thing struck me, although it was nothing uncommon in England, where animals are petted. A heifer and a colt were feeding close to us, with a familiarity which told of kind masters, and I heard the joyful barking of dogs. 'Here,' said I, 'the history of the variations of animals has been written; and, no doubt, the observations are
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A106
Periodical contribution:
Wallace, A. R. 1883. The Debt of Science to Darwin. Century Magazine 25, 3 January: 420-432.
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series between distinct great geological formations were held to show that the older forms of life had been destroyed, and were replaced by a new creation of a more advanced organization suited to the altered conditions of the world. THE GREENHOUSE IN WHICH MR. DARWIN'S EXPERIMENTS AND OBSERVATIONS WERE MADE. And thus, perhaps, we might have gone on to this day, ever accumulating fresh masses of fact, while each set of workers became ever more and more occupied in their own departments of study, and
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A317
Book:
Woodall, Edward. 1884. Charles Darwin. A paper contributed to the Transactions of the Shropshire Archæological Society. London: Trübner.
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ILLUSTRATIONS. PORTRAIT, from a photograph by Captain Darwin ... Facing Title. MR. DARWIN'S BIRTH PLACE ... ... ... ... ... 12 SHREWSBURY SCHOOL (where Mr. Darwin was educated) ... 16 LAGOON-ISLAND ... ... ... ... ... ... ... 32 DOWN HOUSE ... ... ... ... ... ... ... 37 THE GREENHOUSE AT DOWN ... ... ... ... ... 40 FACSIMILE LETTER
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18% |
F1281
Book:
Darwin, C. R. 1884. The different forms of flowers on plants of the same species. 3d thousand. Preface by Francis Darwin. London: John Murray.
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tilised with their own pollen, produced 15 capsules, containing an average of 16 3 seeds. As already stated, the probable average with legitimate plants for a union of this nature is rather above 20 seeds. These plants were remarkably healthy and vigorous, as long as they were kept under highly favourable conditions in pots in the greenhouse; and such treatment greatly increases the fertility of the cowslip. When these same plants were planted during the next year (which, however, was an
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17% |
A317
Book:
Woodall, Edward. 1884. Charles Darwin. A paper contributed to the Transactions of the Shropshire Archæological Society. London: Trübner.
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researches. Darwin used simple means. I looked for the greenhouse, in which those beautiful researches on vegetable hybrids had been made; it contained nothing but a vine. One thing struck me, although it was nothing uncommon in England, where animals are petted. A heifer and a colt were feeding close to us, with a familiarity which told of kind masters, and I heard the joyful barking of dogs. Here, said I, the history of the variations of animals has been written; and, no doubt, the
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15% |
F1281
Book:
Darwin, C. R. 1884. The different forms of flowers on plants of the same species. 3d thousand. Preface by Francis Darwin. London: John Murray.
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. These whilst still in the greenhouse produced smaller leaves and shorter flower-stalks than some legitimate plants with which they grew in competition; but it should be observed that the latter were the product of a cross with a fresh stock, a circumstance which by itself would have added much to their vigour.* When these illegitimate plants were transferred to fairly good soil out of doors, they became during the two following years much more dwarfed in stature and produced very few flower-stems
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CUL-DAR140.3.1--159
Draft:
[1884]
'Reminiscences of My Father's Everyday Life' (partial fair copy)
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maintained it was all affectation saying marking by 15 30 40 instead of 1,2,3. He often knocked up a stray ball for us with the curved handle of his stick. He took no share in the management of the garden which was my mother's business; he sometimes regretted that he did not — like my his own father look after things especially as regards the greenhouse. Nor did he see after horses or cows or any outdoor things — and thus things went on very slackly, so that a miller cheated us for years in
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F1281
Book:
Darwin, C. R. 1884. The different forms of flowers on plants of the same species. 3d thousand. Preface by Francis Darwin. London: John Murray.
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cowslips of both forms, protected from insects in my greenhouse, did not set one pod, though artificially fertilised flowers in other pots produced an abundance. We thus see that the visits of insects are absolutely necessary for the fertilisation of Primula veris. If the corolla of the long-styled form had dropped off, instead of remaining attached in a withered state to the ovarium, the anthers attached to the lower part of the tube with some pollen still adhering to them would have been
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F1281
Book:
Darwin, C. R. 1884. The different forms of flowers on plants of the same species. 3d thousand. Preface by Francis Darwin. London: John Murray.
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in the spring of 1862 forty flowers were fertilised at the same time in both ways. The plants were accidentally exposed in the greenhouse to too hot a sun, and a large number of umbels perished. Some, however, remained in moderately good health, and on these there were twelve flowers which had been fertilised legitimately, and eleven which had been fertilised illegitimately. The twelve legitimate unions yielded seven fine capsules, containing on an average each 57 3 good seeds; whilst the eleven
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F1281
Book:
Darwin, C. R. 1884. The different forms of flowers on plants of the same species. 3d thousand. Preface by Francis Darwin. London: John Murray.
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very rich soil, some in stiff poor clay, some in old peat, and some in pots in the greenhouse; so that these plants, 765 in number, as well as their parents, were subjected to diversified and un * I have given instances in my work 'On the Variation of Animals and Plants under Domestication,' ch. xv. 2nd edit. vol. ii. p. 69. [page] 6
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F1281
Book:
Darwin, C. R. 1884. The different forms of flowers on plants of the same species. 3d thousand. Preface by Francis Darwin. London: John Murray.
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form of a cap. These cleistogamic flowers produced an abundance of seed. Later in the season perfect flowers appeared. With plants in a state of nature the flowers open only in the early morning, as I have been informed by Mr. Wallis, who particularly attended to the time of their flowering. In the case of D. Anglica, the still folded petals on some plants in my greenhouse opened just sufficiently to leave a minute aperture; the anthers dehisced properly, but the pollen-grains adhered in a mass
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9% |
A317
Book:
Woodall, Edward. 1884. Charles Darwin. A paper contributed to the Transactions of the Shropshire Archæological Society. London: Trübner.
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work, but near the metropolis, he spent his fruitful days; and, except the record of his published works, there is little to tell of the forty years passed in this quiet retreat. Happy in his home, with children growing up about him, with 1 For the engraving of Down House, and the Greenhouse (given farther on), we are indebted to the publishers of The Century, in which they appeared (Vol. XXV., No 3), to illustrate an interesting paper by Mr. Wallace. [page] 3
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7% |
A1025
Periodical contribution:
Jordan, David Starr. 1884. [Recollection of Darwin]. Some gossip about Darwin. American Naturalist 18 (January): 108-10.
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first; had his wig and gown and all, but had to give it up on account of bad health. He would have made a hornament to the profession. Francis Darwin is a doctor, and used to work with his father in the greenhouse. He is soon to marry a lady who lectures on Botany in Oxford. [Cambridge] For the first twenty years after Mr. Darwin's return from South America, his health was very bad --much more than later. He had a stomach disease which resulted from sea-sickness while on the voyage around the
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7% |
A77
Periodical contribution:
Hague, J. D. 1884. A reminiscence of Mr. Darwin. Harper's new Monthly Magazine. 69, Issue 413 October: 759-763.
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invitation of Francis Darwin to go with him for a walk about the grounds, in the course of which we followed some of his father's favorite rambles along shaded paths in a neighboring field, coming back finally to the greenhouse, where some interesting experiments on the revolving movement of plants were at that time in progress. The work of the forenoon was the careful observation of a number of tender shoots that were growing in pots, each under a separate bell glass, and all ranged on a table exposed
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7% |
F1281
Book:
Darwin, C. R. 1884. The different forms of flowers on plants of the same species. 3d thousand. Preface by Francis Darwin. London: John Murray.
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, and the female organs of the long-styled form in a more or less complete state, they spontaneously produced a surprising number of capsules, which generally contained a large average of remarkably fine seeds. With ordinary cowslips, legitimately fertilised, I once obtained from plants cultivated in the greenhouse the high average, from seven capsules, of 58 7 seeds, with a maximum in one capsule of 87 seeds; but from plants grown out of doors I never obtained a higher average than 41 seeds. Now
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6% |
A77
Periodical contribution:
Hague, J. D. 1884. A reminiscence of Mr. Darwin. Harper's new Monthly Magazine. 69, Issue 413 October: 759-763.
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little further on were welcomed immediately by Mr. and Mrs. Darwin to a spacious and cheerful parlor or family room, whose broad windows and outer door opened upon a wide and partly sheltered piazza at the rear of the house, evidently a favorite sitting-place, judging from the comfortable look of easy-chairs assembled there, beyond which was a pleasing vista of fresh green lawn, bright flower beds, and blossoming shrubbery, gravel-paths, and a glass greenhouse, or perhaps botanical laboratory
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25% |
Sunday, 23 January 1887 Monday, 24 January 1887 Alice Hutcheson came Tuesday, 25 January 1887 went to Greenhouse !! 48º Wednesday, 26 January 1887 !! Thursday, 27 January 1887 Friday, 28 January 1887 Saturday, 29 January 1887 Alice H. went January – February 188
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the owner. Darwin used simple means. He was not one who would have demanded to have palaces built in order to accommodate laboratories. I looked for the greenhouse in which such beautiful experiments on hybrid plants had been made. It contained only a vine. One thing struck me, although it is not rare in England, where animals are loved. A heifer and a colt were feeding close to us with the tranquillity which tells of good masters, and I heard the joyful barking of dogs. 'Truly,' I said to
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F1452.3
Book:
Darwin, Francis ed. 1887. The life and letters of Charles Darwin, including an autobiographical chapter. vol. 3. London: John Murray.
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you give me any light? Are such plants commoner in warm than in colder climates? I ask because I often walk out in heavy rain, and the leaves of very few wild dicotyledons can be here seen with drops of water rolling off them like quicksilver. Whereas in my flower garden, greenhouse, and hothouses there are several. Again, are bloom-protected plants common on your dry western plains? Hooker thinks that they are common at the Cape of Good Hope. It is a puzzle to me if they are common under very
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9% |
F1452.1
Book:
Darwin, Francis ed. 1887. The life and letters of Charles Darwin, including an autobiographical chapter. vol. 1. London: John Murray.
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gemmules. He was delightfully tender to Polly, and never showed any impatience at the attentions she required, such as to be let in at the door, or out at the verandah window, to bark at naughty people, a self-imposed duty she much enjoyed. She died, or rather had to be killed, a few days after his death.* My father's midday walk generally began by a call at the greenhouse, where he looked at any germinating seeds or experimental plants which required a casual examination, but he hardly ever
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F1452.3
Book:
Darwin, Francis ed. 1887. The life and letters of Charles Darwin, including an autobiographical chapter. vol. 3. London: John Murray.
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of the 'Fertilisation of Orchids,' that he built himself a greenhouse. He wrote to Sir J. D. Hooker (Dec. 24th, 1862): And now I am going to tell you a most important piece of news!! I have almost resolved to build a small hot-house; my neighbour's really first-rate gardener has suggested it, and offered to make me plans, and see that it is well done, and he is really a clever fellow, who wins lots of prizes, and is very observant. He believes that we should succeed with a little patience; it
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13% |
F1225
Book:
Darwin, C. R. 1888. Insectivorous plants. 2d ed. Revised by Francis Darwin. London: John Murray.
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insignificant, as we shall hereafter see, compared with that caused by very weak solutions of several salts of ammonia. FIG. 9. (Drosera rotundifolia.) Leaf (enlarged) with ail the tentacles closely inflected, from Immersion in a solution of phosphate of ammonia (one part to 87,500 of water). Plants which have lived for some time in a rather high temperature are far more sensitive to the action of water than those grown out of doors, or recently brought into a warm greenhouse. Thus in the above
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F1225
Book:
Darwin, C. R. 1888. Insectivorous plants. 2d ed. Revised by Francis Darwin. London: John Murray.
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in thickness, but are always somewhat compressed. At an early stage of growth, the flat or ventral surface faces the axis or stem; but the footstalks must have some power of movement; for in plants kept in my greenhouse the ventral surface was generally turned either straight or obliquely downwards. The Rev. H. M. Wilkinson examined plants for me in a state of nature, and found this commonly to be the case, but the younger bladders often had their valves turned upwards. The general appearance
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F1225
Book:
Darwin, C. R. 1888. Insectivorous plants. 2d ed. Revised by Francis Darwin. London: John Murray.
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. Whether insects alight on the leaves by mere chance, as a resting-place, or are attracted by the odour of the secretion, I know not. I suspect, from the number of insects caught by the English species of Drosera, and from what I have observed with some exotic species kept in my greenhouse, that the odour is attractive. In this latter case the leaves may be compared with a baited trap; in the former case with a trap laid in a run frequented by game, but without any bait. That the glands possess the
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F1225
Book:
Darwin, C. R. 1888. Insectivorous plants. 2d ed. Revised by Francis Darwin. London: John Murray.
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opaque. From these facts it is clear that areolar tissue is easily and quickly digested by the secretion; but that it does not greatly excite the leaves. Cartilage. Three cubes (1/20 of an inch or 1 27 mm.) of white, translucent, extremely tough cartilage were cut from the end of a slightly roasted leg-bone of a sheep. These wero placed on three leaves, borne by poor, small plants in my greenhouse during November; and it seemed in the highest degree improbable that so hard a substance would be
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F1225
Book:
Darwin, C. R. 1888. Insectivorous plants. 2d ed. Revised by Francis Darwin. London: John Murray.
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On the Manner in which Insects are caught. We will now consider the action of the leaves when insects happen to touch one of the sensitive filaments. This often occurred in my greenhouse, but I do not know whether insects are attracted in any special way by the leaves. They are caught in large numbers by the plant in its native country. As soon as a filament is touched, both close with astonishing quickness; and as they stand at less than a right angle to each other, they have a good chance of
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F1225
Book:
Darwin, C. R. 1888. Insectivorous plants. 2d ed. Revised by Francis Darwin. London: John Murray.
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affected; nevertheless, it would be unfair to exclude them. I then waited until I had got eight pairs of fine leaves, and the weather was favourable, the temperature of the room where the leaves were immersed varying from 75 to 81 (23 8 to 27 2 Cent.). In another trial with four pairs (included in the above twenty pairs), the temperature in my room was rather low, about 60 (15 3 Cent.); but the plants had been kept for several days in a very warm greenhouse and thus rendered extremely sensitive
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7% |
F1225
Book:
Darwin, C. R. 1888. Insectivorous plants. 2d ed. Revised by Francis Darwin. London: John Murray.
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or 0000387 mg. Lastly, leaf No. 11, which had after 24 hrs. all its tentacles, except one, closely inflected, as well as the blade, bore the unusually large number of 252 tentacles; and, on the same principle as before, each gland could have absorbed only the 1/200800 of a grain or 0000322 mg. With respect to the following experiments, I must premise that the leaves, both those placed in the solutions and in water, were taken from plants which had been kept in a very warm greenhouse during the
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7% |
F1225
Book:
Darwin, C. R. 1888. Insectivorous plants. 2d ed. Revised by Francis Darwin. London: John Murray.
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be made on plants which have been kept for a considerable time in a warm greenhouse, or cool hothouse; but this is by no means necessary for trials with solutions of moderate strength. I beg the reader to observe that the sensitiveness or irritability of the tentacles was ascertained by three different methods indirectly by drops placed on the disc, directly by * When my first observations were made on the nitrate of ammonia, fourteen years ago, the powers of the spectroscope had not been
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7% |
F1225
Book:
Darwin, C. R. 1888. Insectivorous plants. 2d ed. Revised by Francis Darwin. London: John Murray.
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protoplasm. These glands seem to have very little or no power of absorption, certainly much less than those of the foregoing plants. Mirabilis longiflora. The stems and both surfaces of the leaves bare viscid hairs. Young plants, from 12 to 18 inches in height in my greenhouse, caught so many minute Diptera, Coleoptera, and larv , that they were quite dusted with them. The hairs are short, of unequal lengths, formed of a single row of cells, surmounted by an enlarged cell which secretes viscid
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A873
Book contribution:
Geikie, A. 1888. The life and letters of Charles Darwin. Littell's Living Age 176, 2271 (7 January): 3-10.
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during a succession of years a series of volumes, in which some of his observations and conclusions were worked out in fuller detail. His books on the fertilization of orchids, on the movements and habits of climbing plants, on the variation of animals ana plants under domestication, on the effects of cross and self fertilization in the vegetable kingdom, on the different forms of flowers on plants of the same species, were mainly based on his own quiet work in the greenhouse and garden at Down
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A223
Periodical contribution:
Nash, L. A. 1890. Some memories of Charles Darwin. Overland Monthly (October): 404-408.
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an old pony, oftener on foot, looking for the English likenesses or parallels of the plants he was then studying. In his greenhouse were still the orchids made famous by his researches into the modes of their fertilization. Room was needed for another set of plants, and he gave his orchids to us. How he would gaze on a plant from under his heavy eyebrows! Those eyebrows used to trouble his wife when his photograph was taken: she used to say the photographers gave him no eyes at all. Speaking of
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A268
Book:
Holder, Charles Frederick. 1892. Charles Darwin: his life and work. New York: G. P. Putnam's sons.
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tongue, suggests the general nature of Darwin's botanical studies. His researches were conducted in his laboratory, in pots of plants at his window, in his aquarium, in his greenhouse, in his garden. He worked with instruments of precision, recorded his observations with exactness, and employed every mechanical device for making his results reveal important truths of which the genius of man would seem to be capable. Darwin looked upon plants as living things. He did not study their forms so
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A268
Book:
Holder, Charles Frederick. 1892. Charles Darwin: his life and work. New York: G. P. Putnam's sons.
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ness of his character were apparent, while the delight he manifested in stating facts of interest was excelled only by the eagerness with which he sought them from others, whether while strolling through the greenhouse or sitting round the generously spread table. Going to him as a young entomologist with no claim on his favour, he seemed to take delight in manifesting appreciation. I had occasion, in my Third Report on the Insects of Missouri, published in the spring of that year, to discuss
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F1461
Book:
Darwin, Francis ed. 1892. Charles Darwin: his life told in an autobiographical chapter, and in a selected series of his published letters [abridged edition]. London: John Murray.
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-day walk generally began by a call at the greenhouse, where he looked at any germinating seeds or experimental plants which required a casual examination, but he hardly ever did any serious observing at this time. Then he went on for his constitutional either round the Sand-walk or outside his own grounds in the immediate neighbourhood of the house. The Sand-walk was a narrow strip of land 1 acre in extent, with a gravel-walk round it. On one * The basket in which she usually lay curled up near
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F1461
Book:
Darwin, Francis ed. 1892. Charles Darwin: his life told in an autobiographical chapter, and in a selected series of his published letters [abridged edition]. London: John Murray.
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instantaneously. . . . A cursed insect or something let my last flower off last night. Professor de Candolle has remarked* of my father, Ce n'est pas lui qui aurait demand de construire des palais pour y loger des laboratories. This was singularly true of his orchid work, or rather it would be nearer the truth to say that he had no laboratory, for it was only after the publication of the Fertilisation of Orchids, that he built himself a greenhouse. He wrote to Sir J. D. Hooker (December 24th, 1862): And now
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F2113
Book contribution:
Darwin, C. R. 1896. [Recollections of Darwin]. In E. R. Lankester. 'Charles Robert Darwin'. In C. D. Warner ed. Library of the world's best literature ancient and modern. New York: R. S. Peale & J. A. Hill, vol. 2, pp. 4385-4393.
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. Here most of his children were born, five sons and three daughters. One little girl died in childhood; the rest grew up around him and remained throughout his life in the closest terms of intimacy and affection with him and their mother. Here he carried on his experiments in greenhouse, garden, and paddock; here he collected his library and wrote his great books. He became a man of well-considered habits and method, carefully arranging his day's occupation so as to give so many hours to noting
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F1548.2
Book:
Darwin, Francis & Seward, A. C. eds. 1903. More letters of Charles Darwin. A record of his work in a series of hitherto unpublished letters. London: John Murray. Volume 2
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. H. Crüger, of Trinidad] to ask him to observe what the insects did in the flowers of Melastomaceae: he says not proper season yet, but that on one species a small bee seemed busy about the horn-like appendages to the anthers. It will be too good luck if my study of the flowers in the greenhouse has led me to right interpretation of these appendages. Letter 628. TO J.D. HOOKER. Down, November 28th [1871]. If you had come here on Sunday I should have asked you whether you could give me seed or
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F1548.1
Book:
Darwin, Francis & Seward, A. C. eds. 1903. More letters of Charles Darwin. A record of his work in a series of hitherto unpublished letters. London: John Murray. Volume 1
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could inform me on the points required; but no one other person has so often responded to my miscellaneous queries. I believe I have now in my greenhouse L. trigynum,3 which came up from seed purchased as L. flavum, from which it is wholly different in foliage. I have just sent in a paper on Dimorphism of Linum to the Linnean Society,4 and so I do not doubt your memory is right about L. trigynum: the functional difference in the two forms of Linum is really wonderful. I assure you I quite long to
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F1548.1
Book:
Darwin, Francis & Seward, A. C. eds. 1903. More letters of Charles Darwin. A record of his work in a series of hitherto unpublished letters. London: John Murray. Volume 1
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open, immersed in it, in the cellar for uniform temperature. I must plant out of doors, as I have no greenhouse. I told you I had inserted notice in the Gardeners' Chronicle, and to-day I have heard from Berkeley that he has already sent an assortment of seeds to Margate for some friend to put in salt water; so I suppose he thinks the experiment worth trying, as he has thus so very promptly taken it into his own hands.1 Reading this over, it sounds as if I were offended!!! which I need not say is
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F1548.1
Book:
Darwin, Francis & Seward, A. C. eds. 1903. More letters of Charles Darwin. A record of his work in a series of hitherto unpublished letters. London: John Murray. Volume 1
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startling one, when carried to the length of supporting plants of cold temperate regions; and I must confess that, much as I should like it, I can hardly stomach keeping the tropical genera alive in so very cool a greenhouse [pencil note by C.D., Not so very cool, but northern ones could range 1. Salter, Linn. Soc. Journal, I., 1857, page 140, On the Vitality of Seeds after prolonged Immersion in the Sea. It appears that in 1843 the mud was scraped from the bottom of the channels in Poole
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F1548.1
Book:
Darwin, Francis & Seward, A. C. eds. 1903. More letters of Charles Darwin. A record of his work in a series of hitherto unpublished letters. London: John Murray. Volume 1
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the Cameroons, and as Seemann describes, in low mountains of Panama. It is, as you say, absurd to suppose that such a genus as Dipterocarpus1 could have been developed since the Glacial era; but do you feel so sure, as to oppose2 a large body of considerations on the other side, that this genus could not have been slowly accustomed to a cooler climate? I see Lindley says it has not been brought to England, and so could not have been tried in the greenhouse. Have you materials to show to what
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F1548.2
Book:
Darwin, Francis & Seward, A. C. eds. 1903. More letters of Charles Darwin. A record of his work in a series of hitherto unpublished letters. London: John Murray. Volume 2
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1 1/2 hour, I found an indusium with hairs on the outer edge perfectly clogged with pollen, and horns protruded, which before the 1 1/2 hour had not one grain of pollen outside the indusium, and no trace of protruding horns. So you will see how I wish to know whether the horns are the true stigmatic surfaces. I would try the case experimentally by putting pollen on the horns, but my greenhouse is so cold, and my plant so small, and in such a little pot, that I suppose it would not seed... The
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F1548.2
Book:
Darwin, Francis & Seward, A. C. eds. 1903. More letters of Charles Darwin. A record of his work in a series of hitherto unpublished letters. London: John Murray. Volume 2
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complex and still more strange; as the age and position of the pistils comes into play, in relation to the two kinds of pollen. These facts seem to me so curious that I do not scruple to ask you to see whether you can lend me any Melastomad just before flowering, with a not very small flower, and which will endure for a short time a greenhouse or sitting-room; when fertilised and watered I could send it to Mr. Turnbull's to a cool stove to mature seed. I fully believe the case is worth investigation
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F1548.2
Book:
Darwin, Francis & Seward, A. C. eds. 1903. More letters of Charles Darwin. A record of his work in a series of hitherto unpublished letters. London: John Murray. Volume 2
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I will send up early to-morrow two plants [of Dionoea] with five goodish leaves, which you will know by their being tied to sticks. Please remember that the slightest touch, even by a hair, of the three filaments on each lobe makes the leaf close, and it will not open for twenty-four hours. You had better put 1/4 in. of water into the saucers of the pots. The plants have been kept too cool in order to retard them. You had better keep them rather warm (i.e. temperature of warm greenhouse) for a
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F1548.2
Book:
Darwin, Francis & Seward, A. C. eds. 1903. More letters of Charles Darwin. A record of his work in a series of hitherto unpublished letters. London: John Murray. Volume 2
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protect the leaflets from rain. Accordingly I syringed the plant for two minutes, and it was really beautiful to see how each leaflet on the younger leaves twisted its short sub-petiole, so that the blade was immediately directed at an angle between 45 and 90 deg to the horizon. I could not resist the pleasure of just telling you why I want to know the name of the Cassia. I should add that it is a greenhouse plant. I suppose that there will not be any better flowers till next summer or autumn
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A237.2
Book:
Wallace, A. R. 1905. My life: A record of events and opinions. London: Chapman and Hall. vol. 2.
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been able to satisfy my craving for enjoying new forms of plant-life every year, partly by raising numbers of seeds of hardy and greenhouse plants, always trying some of the latter in sheltered places out-of-doors, and partly by exchanges or by gifts from friends, so that every year I have the great pleasure of watching the opening of some of nature's gems which were altogether new to me, or of others which increase year by year in beauty. In one end of my greenhouse I have a large warmed tank
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A237.2
Book:
Wallace, A. R. 1905. My life: A record of events and opinions. London: Chapman and Hall. vol. 2.
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among residents who had come to the place for its general educational advantages or for the charm of its rural scenery. We had here about half an acre of ground with oak trees and hazel bushes (from which I named our place Nutwood Cottage ), and during the eight years we lived there I thoroughly enjoyed making a new garden, in which, and a small greenhouse, I cultivated at one time or another more than a thousand species of plants. The soil was a deep bed of the Lower Greensand formation, with a
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A237.2
Book:
Wallace, A. R. 1905. My life: A record of events and opinions. London: Chapman and Hall. vol. 2.
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that rarity in America, a greenhouse attached to a small country house. All down the valley in alluvial flats the Western plane tree (Platanus occidentalis) had a remarkable appearance, its upper half being pure white, exactly as if whitewashed. This is the colour of the young bark before it flakes off, as it does on the trunk and larger limbs. The peculiar appearance is not noticed by Loudon, so perhaps it is not produced in our less sunny climate. I reached Coalburg at 3 p.m., where Mr
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A237.2
Book:
Wallace, A. R. 1905. My life: A record of events and opinions. London: Chapman and Hall. vol. 2.
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America. I had spent the morning in the fine Golden Gate Park, where I saw some eucalyptus trees over sixty feet high, with numerous acacias and other greenhouse plants growing out-of-doors. I also had a fine view of the extensive sandhills, covered with huge clumps of blue and yellow tree-lupines, which produced a splendid effect. The interesting s ances I had here will be described later on. Returning to Stockton, I went with my brother and his daughter for a few days in the Yosemite Valley. The
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A237.2
Book:
Wallace, A. R. 1905. My life: A record of events and opinions. London: Chapman and Hall. vol. 2.
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catalogues for rarities and curiosities, and I find that I attempted the cultivation in this garden, or in a very small greenhouse and verandah, of about fifteen hundred species of plants, some of which, of course, never reached flowering size, others survived only a few years; but the delight of watching the growth of these, to me, new forms of vegetable life, and seeing them flower even once or twice, was so great that no trouble was spared to obtain it. My gardening has always been to me pure
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A237.2
Book:
Wallace, A. R. 1905. My life: A record of events and opinions. London: Chapman and Hall. vol. 2.
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of 400 or 500 a year, I think it probable that I should not have written another book, but should have gone to live further in the country, enjoyed my garden and greenhouse (as I always have done), and limited my work to a few lectures and review articles, but to a much less extent than I actually have done. It was the necessity of earning money, owing to my diminishing income, that caused me to accept invitations to lecture, which I always disliked; and the same reason caused me to seek out
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A297
Book:
Darwin, Francis & E. Hamilton Acton. 1909. Practical physiology of plants. Cambridge: University Press.
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30 ASSIMILATION. [CH. II smeared with wax-mixture, and the junction with the glass plate is made secure by a little embankment of Fig. 8. Exp. 35. wax-mixture melted into the angle with a hot wire. The aperture of the bell is closed by a rubber cork pierced for the tube T, which contains soda-lime. The apparatus should be placed out of doors or in a brightly lighted greenhouse. A control-plant must be fitted up in a similar way except that G may be dispensed with and that T must be filled with
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A297
Book:
Darwin, Francis & E. Hamilton Acton. 1909. Practical physiology of plants. Cambridge: University Press.
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24 ASSIMILATION. [CH. II water-plants (e.g. Elodea, Potamogeton) which have been placed in the dark room for about four days. The control-plants must be grown either out of doors or in a greenhouse. (28) Effect of dull light. Sachs' method may be used to demonstrate a fact, the knowledge of which is of practical value to the physiologist1, namely, that plants in a laboratory suffer from want of light far more than would be readily supposed—and that accordingly experimental plants cannot be too
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A297
Book:
Darwin, Francis & E. Hamilton Acton. 1909. Practical physiology of plants. Cambridge: University Press.
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CH. Il] PHOTOGRAPHIC METHOD. 25 (30) Gardiner's experiment1. A plant growing in a flower-pot (for convenience of moving) is placed in the dark for 24 hours, or until the leaves are found to be free from starch. One of the leaves is now covered with a photographic negative and left exposed to bright light out of doors, or in a greenhouse, until the evening, when the leaf is tested for starch. Tt will be found that an accurate copy of the photograph has been printed in starch. (31) Effect of
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A297
Book:
Darwin, Francis & E. Hamilton Acton. 1909. Practical physiology of plants. Cambridge: University Press.
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cultures keep reasonably free from algre. Fig. 11 gives the result of an experiment carried on in a greenhouse in the winter. Three jars S, K, W, were prepared, in each of which six fronds were placed. S contained 0-25 °/o Sachs' mixture of salts; K contained 0'25 °/0 potassium nitrate, while W contained only distilled water, a drop of dialysed iron being added to each culture. The amount of increase is shown in the figure, the difference in root production as well as in the amount of frond is
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A297
Book:
Darwin, Francis & E. Hamilton Acton. 1909. Practical physiology of plants. Cambridge: University Press.
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tension1 which consists in turgescent tissues may be made in summer or spring on any rapidly growing juicy shoot, e.g. elder (Sambucus), or with certain leafstalks, e.g. that of the rhubarb (Rheum). In winter it is sometimes difficult to find suitable material: if a greenhouse is available, the leaf stalks of Richardia will answer well. It is best to get fairly long shoots, i.e. not less than 1 See Sachs' Text-book, Sect. 1-i, 15. The whole discussion should be studied. [page break
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A297
Book:
Darwin, Francis & E. Hamilton Acton. 1909. Practical physiology of plants. Cambridge: University Press.
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containing the plant might be transferred. (234) Mimosa : effect of darkness. If Mimosa is kept in the dark for several days it loses its sensitiveness. The plant should be kept in a damp atmosphere in a greenhouse at a temperature of at least 16°—17° 0. The best plan is to place the flower-pot in a tray of wet sawdust and to invert over it a tin cylinder, the rim of which should sink into the sawdust. We find that 4 or 5 days are needed to destroy sensitiveness. In [page break
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A297
Book:
Darwin, Francis & E. Hamilton Acton. 1909. Practical physiology of plants. Cambridge: University Press.
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well. The only difficulty is to provide a suitable atmosphere in which to hang up the sieve. If the air is too dry the roots wither before they have time to bend; if too damp the surface of the sieve does not supply a sufficiently strong contrast to the surrounding air. A greenhouse atmosphere answers fairly well, or, as Sachs recommends, a large cupboard or 1 See however Elfving's interesting paper in Nature, March 15, 1894, where references to his original paper and to Errera's work on the
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A297
Book:
Darwin, Francis & E. Hamilton Acton. 1909. Practical physiology of plants. Cambridge: University Press.
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the following morning. The tracing therefore (if the broken line be neglected) practically represents the circumnutation during a day of about 12 hours. (259) Circumnutation: twining plants*. The observations should be made either in a greenhouse or indoors, on Humulus lupulus (the hop) and Phaseolus multifloras. The basal part of the plant should be tied to a stick stuck in the soil of the pot, and 6 inches or a foot 1 The tracing was made by a slightly different method to that here described
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A162
Book:
Seward, A. C. ed. 1909. Darwin and modern science. Essays in commemoration of the centenary of the birth of Charles Darwin and of the fiftieth anniversary of the publication of The origin of species. Cambridge: Cambridge University Press.
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including several elementary species. The legitimately produced plants are thus true 1 Under very favourable conditions (in a greenhouse) the fertility of the plants of the fourth generation increases—a point, which in view of various theoretical questions, deserves further investigation. 2 Forms of Flowers, p. 242. [page] 41
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A279
Pamphlet:
Darwin, George and Francis Darwin eds. 1909. Darwin celebration, Cambridge, June, 1909. Speeches delivered at the banquet held on June 23rd. Cambridge: Cambridge Daily News.
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and the sand walk are practically unaltered, and the house is the same except in respect of furniture and fixtures: unfortunately the greenhouse in which he worked was so dilapidated that it had to be renewed. The railway station is within four miles of the house, and there is a good road to the village, and it is not the case, as has been stated by a German writer, that the house can only be reached by a mule track. If I might be allowed a very few minutes more there is one other subject I
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A546
Periodical contribution:
Anon. 1909. A visit to Darwin's village: reminiscences of some of his humble friends. Evening News (12 February): 4.
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.' 'I don't suppose you do,' I said to him. Quite candid on both sides. Darwin's House. I bade Mr. Ffinden farewell and drove off to Darwin's house. It is only a little way out of the village, and stands close to the road. Eighteen months ago it was taken by two ladies, who have set up a girls' school. One of the heads received me, and readily offered to take me over the grounds and through the house. First we went down to the little greenhouse and the laboratory and the little dark house he had
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F3385
Book contribution:
Darwin, C. R. 1914. [Letters to John Lubbock and Lubbock's recollections of Darwin]. In Horace Gordon Hutchinson, Life of Sir John Lubbock, Lord Avebury. 2 vols. London.
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[…]. On the Sunday afternoon Sir John Lubbock, our host, took us all up to the hill-top whence in his quiet Kentish village Darwin was shaking the world. The illustrious pair, born in the same year, had never met before. Mr. Gladstone, as soon as seated, took Darwin's interest in lessons of massacre for granted, and launched forth his thunderbolts with unexhausted zest. His great, wise, simple, and truth-loving listener, then, I think, busy on digestive powers of the drosera in his greenhouse
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F1553.2
Book:
Litchfield, H. E. ed. 1915. Emma Darwin, A century of family letters, 1702-1896. London: John Murray. Volume 2
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visited my father's old home, the Mount, and were accompanied by the owner as we were shown over the house. This was meant in all kindness, but I remember my father's deep disappointment as he said, If I could have been left alone in that greenhouse for five minutes, I know I should have been able to see my father in his wheel-chair as vividly as if he had been there before me. [page] 19
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F1592.2
Book:
Marchant, James ed. 1916. Alfred Russel Wallace letters and reminiscences. London: Cassell. Volume 2.
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, I have been thinking of a plan of making a little bed for them on the top of the new rockery where there is now nothing particular. Will you please plant them out carefully in the zinc tray of peat and sphagnum that stands outside near the little greenhouse door? Just lift up the sphagnum and see if the earth beneath is moist, if not give it a soaking. Then put them all in, the short-rooted ones in the sphagnum only, the others through into the peat. Then give them a good syringing and put the
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F1592.2
Book:
Marchant, James ed. 1916. Alfred Russel Wallace letters and reminiscences. London: Cassell. Volume 2.
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enthusiasm for orchids: TO MISS VIOLET WALLACE Parkstone, Dorset. November 25, 1894. My dear Violet, I have found a doctor at Poole (Mr. Turner) who has two nice orchid houses which he attends to entirely himself, and as I can thus get advice and sympathy from a fellow maniac (though he is a public vaccinator!) my love of orchids is again aroused to fever-heat, and I have made some alterations in the greenhouse which will better adapt it for orchid growing, and have bought a few handsome kinds
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F1592.2
Book:
Marchant, James ed. 1916. Alfred Russel Wallace letters and reminiscences. London: Cassell. Volume 2.
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The next letter refers to the discovery of a rare moth and some beetles in the root of an orchid. It was certainly a strange yet pleasant coincidence that these creatures should find themselves in Dr. Wallace's greenhouse, where alone they would be noticed and appreciated as something uncommon. TO MR. W. G. WALLACE Old Orchard, Broadstone, Wimborne. February 23, 1909. My dear Will, In my last letter I did not say anything about my morning at the Nat. Hist. Museum. What I enjoyed most was
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F1592.2
Book:
Marchant, James ed. 1916. Alfred Russel Wallace letters and reminiscences. London: Cassell. Volume 2.
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charmed a geologist. However, we have stopped that for the future by a drain under the doorstep. The new breakfast-room is being papered and will look tidy soon. A man has been to measure for the stairs. The front porch door is promised for to-morrow, and the stairs, I suppose, in another week. A lot of fresh pointing is to be done, and all the rain-water pipes and the rain-water cistern with its overflow pipes, and then the greenhouse, and then all the outside painting after which we shall rest for a
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F1592.2
Book:
Marchant, James ed. 1916. Alfred Russel Wallace letters and reminiscences. London: Cassell. Volume 2.
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next month he was very bright and cheerful, though, as some of his letters to his old friend Dr. Richard Norris and to Dr. Littledale show, he had been becoming increasingly weak. TO MISS NORRIS Old Orchard, Broadstone, Dorset. December 10, 1912. My dear Miss Norris, I am very sorry to hear that your father is so poorly. The weather is terribly gloomy, and I have not been outside my rooms and greenhouse for more than an hour a week perhaps, for the last two months, and [page] 13
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A488
Book:
Nash, Wallis. 1919. A lawyer's life on two continents. Boston: Richard G. Badger, the Gorham Press. [Darwin reminiscences only]
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will find the journal for 18— of such a German entomological society. About page 357, on the right of the book and midway down, I think you will find the description of the insect and its life history. And there or thereabouts, we did find it. In the garden behind the house was the greenhouse with a mixed collection of the plants familiar to the world. Their habits were studied, changed modes of growth in the face of natural and artificial obstacles —their insect friends and enemies—influences on
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A283
Pamphlet:
Darwin, Francis. 1920. The story of a childhood. Edinburgh: Privately printed.
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said home. No. 29. August 13, 1878, Leith Hill. Meggy is a nurse-like little girl, and took great care of him. He was enjoying swimming very much, i.e. slipping about on his smooth little tail on the polished oak floor. He is very fond of Jane, and said we should see Na at Down, when we got there. He calls all cats Na, because he thinks they are Jane's [Na's] cats. No. 30. August 29, 1878. I asked B. this morning whether he would like to come to the greenhouse with me, and all he would say was
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A179
Book:
Ward, Henshaw. 1927. Charles Darwin: The man and his warfare. London: John Murray.
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tuft of hair as being in accordance with his theory of pangenesis; her father had been a red bull-terrier, thus the red hair appearing after the burn showed the presence of latent red gemmules. He was delightfully tender to Polly, and never showed any impatience at the attentions she required. My father's midday walk generally began by a call at the greenhouse, where he looked at any germinating seeds or experimental plants which required a casual examination, but he hardly ever did any serious
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A1111
Periodical contribution:
Anon. 1927. [Recollection of Darwin by gardener Henry Wheeler]. Darwin at home: a crusty, snuff-taking recluse. Sunday Post (4 September): 3.
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, but Darwin's repugnance for it. When we could not get flies in the winter, I had to feed the inhuman thing on tiny bits of meat, an occupation that always made me shiver. Darwin kept most of his specimens in his study, where he spent most of his time, but if you wandered round the grounds you would sometimes come unexpectedly upon evidence of one of his strange experiments. For instance, his greenhouse was full of the most beautiful blossoms, tended by my father-in-law, but if you passed right
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CUL-DAR262.28.6
Note:
[1929]
Recollection of Darwin. Episode [concerns supplying chloride of tin]
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departed, passed into the drawing-room. Parslow then took me out to the low greenhouse (now demolished) where Mr Darwin was bending over some plants. After his usual cordial enquiry after my father's health, I told him what I had got but which he had evidently forgotten about. He however became quite excited and said I must try it at once He then pushed back some plants he had been examining, and took done two or three others– meanwhile sending Parslow for pipette etc. while I explained the point
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F2554
Periodical contribution:
W.A.H. 1930. Charles Darwin: Some reminiscences [with words attributed to Darwin]. Telegraph, (Brisbane, Queensland), (10 March): 20.
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taking leave of Mr. Darwin we were invited to see the greenhouses where his father carried on many of those patient, wonderful and far-reaching investigations into plant life, which have revolutionised the Science of Botany. DARWIN'S GREENHOUSE. The greenhouse was a small and unpretentious structures about 30 feet by 12 feet. Here we saw the Fly-trap (Dioea Muscipula), basins of water containing Upricularia, Sun-dews (Drosera Rotundefolia), all of which were just recovering from the series of
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A346
Periodical contribution:
Blackman, F. F. 1932. Obituary notice of Francis Darwin. (With portrait). 1848-1925. Proceedings of the Royal Society of London 110: i-xxi.
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brought from a greenhouse into a dry laboratory, and thereby exhibit a reduction of water-loss. This reduction we should attribute to biological regulation by the reaction of the guard-cells. As we have indicated higher up, the behaviour when the humidity at the sink is lowered partly depends on the productive power of the source. Aloi had pointed out, and Darwin confirmed, that if the soil of a pot plant is abnormally saturated with water then a higher rate of water-loss initiated by a drier sink may
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A346
Periodical contribution:
Blackman, F. F. 1932. Obituary notice of Francis Darwin. (With portrait). 1848-1925. Proceedings of the Royal Society of London 110: i-xxi.
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opening of stomata associated with it was not tested by the porometer, as the humidity of the air inside the porometer would require special controlling complications. This initial phase may last perhaps 20 minutes. After 30 minutes or so there sets in a marked reduction of water-loss when the potted plant is brought from the greenhouse to the drier laboratory. Numerous measurements with the horn-hygroscope prove the generality of this effect. The reduction comes on before the eye detects any
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A871
Periodical contribution:
Anon. 1932. Darwin at Down: the house and garden. The Times (19 April): 19.
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. Some of the landmarks are still to be seen: the bank against which Darwin liked to lie: the great limes overshadowing it; the beam of the children's swing between the yew trees. The azalea bed still flourishes; the long shady border, with its flowering shrubs, has been partly renovated, with generous help from Kew Gardens. The greenhouse and the little laboratory attached to it still stand. The flat round stone on the surface of the lawn, used in connexion with the study of the work of earth
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A540
Pamphlet:
Howarth, O. J. R. and E. K. Howarth. [1933]. A history of Darwin's parish: Downe, Kent. With a foreword by Sir Arthur Keith. Southampton: Russell & Co.
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ever it reveals any hurt, that is more than half hidden, often under some word of humour. He never left Downe for long: his days followed a regular order, and Mrs. Darwin's loving task was to help him in the fight against bodily weakness. The hours given daily to the actual writing of books were not very long; but there was work upon plants in the greenhouse or at the window-shelf in the ' old study', and other observations of nature; there was a very large correspondence. For recreation he rode in
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A540
Pamphlet:
Howarth, O. J. R. and E. K. Howarth. [1933]. A history of Darwin's parish: Downe, Kent. With a foreword by Sir Arthur Keith. Southampton: Russell & Co.
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it. Outside, the greenhouse and experimental laboratory survive, the orchard, the bank beneath the big lime trees [where Darwin used often to lie], the Sand-walk, and the view from it over the ' quiet little valley ' with its woods, which may man preserve ! For that view helped to draw Darwin and his wife to Down. We see still the famous ' worm-stone ' at the edge of the lawn, used in making the observations on which was based the work on vegetable mould and earthworms; still see, too, the hard
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F1571
Book:
Barlow, Nora ed. 1945. Charles Darwin and the voyage of the Beagle. London: Pilot Press.
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the site and had built the solid comfortable red brick residence soon after he had married Susannah Wedgwood in 1796. There Dr. Robert lived for the rest of his life, and there Charles was born in 1809 and grew to manhood, familiar with the lawns and big trees and the terrace overlooking the meadows below, the orchard and newly planned greenhouse, and the flower garden where his sisters worked. During the winter months, when Charles was enduring the summer in the Antipodes, he would imagine the
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A6590
Pamphlet:
Anon. [1946.] Historical and descriptive catalogue of the Darwin Memorial at Down House Downe, Kent. British Association for the Advancement of Science. [London: n.p.] 35pp.
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which used to mark their terms of office. Outside the house, much of the garden remains as Darwin knew it. Here are his greenhouse and experimental laboratory, and a curious area of concrete, an early example of a hard tennis court (1881). Above all, the shaw or little wood known as the Sandwalk, from the sandy loam with which its path was dressed, was a usual scene of Darwin's daily exercise; it was known also as his 'thinking-path,' for here as he walked he thought over his work. The
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CUL-DAR132.1
Printed:
[1946]
'Historical and descriptive catalogue of the Darwin Memorial at Down House, Downe, Kent' [London]: 35pp
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which used to mark their terms of office. Outside the house, much of the garden remains as Darwin knew it. Here are his greenhouse and experimental laboratory, and a curious area of concrete, an early example of a hard tennis court (1881). Above all, the shaw or little wood known as the Sandwalk, from the sandy loam with which its path was dressed, was a usual scene of Darwin's daily exercise; it was known also as his 'thinking-path,' for here as he walked he thought over his work. The
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F1598
Book:
Barlow, Nora ed. 1967. Darwin and Henslow. The growth of an idea. London: Bentham-Moxon Trust, John Murray.
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Alegra in this country: the temperature of the former, must be that of a warm greenhouse, even plants of this country would require some protection (the olive orange bear fruit here). Also a bag of the sweepings of a Granary; it will be a Botanical problem to find out to what country the weeds belong: It might be curious to observe whether Europaean weeds have undergone any change by their residence in this country.1 If they are like the men, I will answer for it they are not much improved. I
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A668
Book:
Atkins, Hedley. 1976. Down: the home of the Darwins; the story of a house and the people who lived there. London: Royal College of Surgeons [Phillimore].
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Bob, a black and white half-bred retriever. This was the dog that used to accompany him round the Sandwalk and who put on a miserable expression when Charles would delay things by first of all pottering about in the greenhouse. Bob's hot house face has been immortalised in Charles' book, The Expression of the Emotions. In March 1870 Bob was taken ill and Emma writes to Henrietta, Poor Bobby is better to-day and has eaten a little. He looked so human, lying under a coat with his head on a pillow
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10.30 12 or Returned to study, which period he considered 12.15 am the end of his working day. 12 noon Walk, starting with visit to greenhouse, then round the sandwalk, the number of times depending on his health, usually alone or with a dog. 12.45 pm Lunch with whole family, which was his main meal of the day. After lunch read The Times and answered his letters. 3 pm Rested in his bedroom on the sofa and smoked a cigarette, listened to a novel or other light literature read by ED. 4 pm Walked
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main entrance of E side. 1881 billiard room converted to new study. ALTERATIONS TO LAND: 1844 New garden wall built; 1845 Mound under yews on W side removed, mound added at E side as wind protection; 1846 Sandwalk wood planted on land rented from Sir John William Lubbock; 1862 Feb. New greenhouse completed, superintended by Horwood, Sir John W. Lubbock's gardener; 1874 Sandwalk wood exchanged for a piece of pasture with Sir John Lubbock; 1881 Bought strip of field beyond orchard from Sydney Sales
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F167c
Book:
Keynes, Richard Darwin ed. 1979. The Beagle record. Cambridge: Cambridge University Press.
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am very anxious to hear how the spirit withstands evaporation, an insect case, which would require airing, a small box of stones, which may be left in statu quo, a bundle of seeds, which I send as a most humble apology for my idleness in Botony. They were collected in Port Alegra in this country: the temperature of the former must be that of a warm greenhouse, even plants of this country would require some protection (the olive orange bear fruit here). Also a bag of the sweepings of a Granary
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A691
Pamphlet:
[Titheradge, Philip]. 1981. The Charles Darwin memorial at Down House, Downe, Kent. [St. Ives, Cornwall: B. Tempest & Co.]
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.' Of the lime trees few remain and the orchard beyond is now a private garden but the yew trees are still to be seen, as is the mulberry thought to have been planted in 1609 and mentioned by Gwen Raverat in 'Period Piece'. Although the sand has washed away from Charles's 'thinking path' you may still follow in his footsteps through the 'Sandwalk Wood'. On your way up the path to the top of the garden you will pass Darwin's greenhouse which retains the original Victorian ironwork. The work of
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A2955
Pamphlet:
Freeman, R. B. 1982. Darwin and Gower Street: an exhibition in the Flaxman Gallery of the Library, University College London, Monday 19 April 1982. London: UCL.
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described as one of the most often printed and least often read books in the world. The Voyage of the Beagle is the second, not quite so often printed, but much more often read. This case shows the extraordinary width of his work, a classical work on taxonomy, a still readable biography, and above all the experimental work on plants which he undertook once he was settled in at Down House, with his own garden and greenhouse facilities. [page] 2
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availability of a heated greenhouse which he had built at Down House in 1862. On the movements and habits of climbing plants had first appeared in the Journal of the Linnean Society, in 1865, although it was also available as a slender book at the same time. After more work, a greatly enlarged second edition appeared ten years later, and finally, in 1880, he extended the ideas to show that the same functional methods occurred in flowering plants in general; in this book, Power of movement in
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F3275
Book:
Gregorio, Mario A. -Di, ed. 1990. Charles Darwin's marginalia, vol. 1. Edited by Mario A. Di Gregorio, with the assistance of N.W. Gill. New York; London: Garland.
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-32m 324 S-20m 325 3-6m 329 13-15m, 16-20m, 21-25m 347 3-9m 350 3-20m 355 26-30m 364 25-30m 377 23-2 6m (Gaudin)/14u souvent apétale /w dimorphic 385 2-4m 400 3-8m/4u Austriaticum 401 21-25m\22m\m\ 24u c'estlies 403 16-18m\18u dix carpelles 404 19-20m 407 22-22m 408 18-23xv Fermond 22-27m 493 23-26m, 29-33m (Linnaeus) 518 37-39m 521 18-20m, 30-32m 523 37-41m, wb I have seen this differ in Greenhouse vars 525 16-21m/16-27w strange notion but shows what a relation there is in position of anther
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F3275
Book:
Gregorio, Mario A. -Di, ed. 1990. Charles Darwin's marginalia, vol. 1. Edited by Mario A. Di Gregorio, with the assistance of N.W. Gill. New York; London: Garland.
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flowers organs of fructification not very fixed, especially in Exotics 332 Urn, 13-20m/w From this it almost follows that artificial self-fructification was done in House 333 l-2m/wt Many plants more fertile in wild state than in Garden or greenhouse. 22-Ï2« Gràsern\u.s.w./13u*/ 14u*/15w Nothing 5-15m/w In some, rich food makes more seed, in others a withdrawal of food. In former, those with dark. 26« Henschel /w Has written on the above 335 ll-12w aid of insects overrated by some, underated by
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A27b
Book:
Freeman, R. B. 2007. Charles Darwin: A companion. 2d online edition, compiled by Sue Asscher.
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Darwin, Charles Robert, 1809-1882. HABITS, continued. 10.30-12 or 12.15pm Returned to study, which period he considered the end of his working day. 12 noon Walk, starting with visit to greenhouse, then round the sandwalk, the number of times depending on his health, usually alone or with a dog. 12.45pm Lunch with whole family, which was his main meal of the day. After lunch read The Times and answered his letters. 3pm Rested in his bedroom on the sofa and smoked a cigarette, listened to a
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A27b
Book:
Freeman, R. B. 2007. Charles Darwin: A companion. 2d online edition, compiled by Sue Asscher.
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to drawingroom. 1877 New billiard room added and new main entrance of east side. 1881 Billiard room converted to new study. ALTERATIONS TO LAND: 1844 New garden wall built. 1845 Mound under yews on west side removed, mound added at east side as wind protection. 1846 Sandwalk wood planted on land rented from Sir John William Lubbock. 1863 Feb. New greenhouse completed, superintended by John Horwood, Mr Turnbull's gardener at The Rookery. 1874 Sandwalk wood exchanged for a piece of pasture with
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F2043
Book:
Wyhe, John van ed. 2009. Charles Darwin's shorter publications 1829-1883. With a foreword by Janet Browne and Jim Secord. Cambridge: University Press.
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to his greenhouse to begin an experiment that might test a statement that had recently come to hand. Or he might turn to friends and relations for verification. Always, his mind was alert to the tiny fact, the unobserved point that might contribute to his larger insight into the living world. This trait was evident in Darwin's character from very early on, and still charms readers today. Just before the Beagle voyage took place, his uncle Josiah Wedgwood called him ‘a man of enlarged curiosity'
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A715
Periodical contribution:
Pasquarè, G., Chiesura, G., Battaglia, T.A., Guaraldi Vinassa de Regny, I. and Pezzotta, F. 2009. Charles Darwin geologist at Santiago (Cape Verde Islands): a field reappraisal. Acta Vulcanologica 20-21: 223-231.
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not a botanical garden, a greenhouse of tropical plants, but a barren volcanic island that immediately challenges Darwin's scarce preparation in geological science. It is to be his baptism in geology, and already after the first day his immersion in complex stratigraphic, morphologic and volcanologic problems will be as good as total, so much so, that in the comments in his Notebooks, Nora Barlow writes A brief line, Wild cats, Kingfisher , is the only zoological note in the Cape de Verde; his
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