F373    Book:     Darwin, C. R. 1859. On the origin of species by means of natural selection, or the preservation of favoured races in the struggle for life. London: John Murray. 1st ed., 1st issue.   Text   Image   PDF

CHAPTER VI. DIFFICULTIES ON THEORY. Difficulties on the theory of descent with modification — Transitions — Absence or rarity of transitional varieties — Transitions in habits of life — Diversified habits in the same species — Species with habits widely different from those of their allies — Organs of extreme perfection — Means of transition — Cases of difficulty — Natura non facit saltum — Organs of small importance — Organs not in all cases absolutely perfect — The law of Unity of Type and

PC-Virginia-Francis-F373    Printed:    1859   On the origin of species by means of natural selection, or the preservation of favoured races in the struggle for life. London: John Murray. [Francis Darwin's copy]  London   Text   Image   PDF

CHAPTER XIII. MUTUAL AFFINITIES OF ORGANIC BEINGS: MORPHOLOGY: EMBRYOLOGY: RUDIMENTARY ORGANS. CLASSIFICATION, groups subordinate to groups — Natural system — Rules and difficulties in classification, explained on the theory of descent with modification — Classification of varieties — Descent always used in classification — Analogical or adaptive characters — Affinities, general, complex and radiating — Extinction separates and defines groups — MORPHOLOGY, between members of the same class

F373    Book:     Darwin, C. R. 1859. On the origin of species by means of natural selection, or the preservation of favoured races in the struggle for life. London: John Murray. 1st ed., 1st issue.   Text   Image   PDF

as the presence of peculiar species of bats, and the absence of all other mammals, on oceanic islands, are utterly inexplicable on the theory of independent acts of creation. The existence of closely allied or representative species in any two areas, implies, on the theory of descent with modification, that the same parents formerly inhabited both areas; and we almost invariably find that wherever many closely allied species inhabit two areas, some identical species common to both still exist

F373    Book:     Darwin, C. R. 1859. On the origin of species by means of natural selection, or the preservation of favoured races in the struggle for life. London: John Murray. 1st ed., 1st issue.   Text   Image   PDF

suddenly appear in certain formations, has been urged by several palæontologists, for instance, by Agassiz, Pictet, and by none more forcibly than by Professor Sedgwick, as a fatal objection to the belief in the transmutation of species. If numerous species, belonging to the same genera or families, have really started into life all at once, the fact would be fatal to the theory of descent with slow modification through natural selection. For the development of a group of forms, all of which have

F373    Book:     Darwin, C. R. 1859. On the origin of species by means of natural selection, or the preservation of favoured races in the struggle for life. London: John Murray. 1st ed., 1st issue.   Text   Image   PDF

of growth, but in order to excrete horny matter, as that the rudimentary nails on the fin of the manatee were formed for this purpose. On my view of descent with modification, the origin of rudimentary organs is simple. We have plenty of cases of rudimentary organs in our domestic productions,—as the stump of a tail in tailless breeds,—the vestige of an ear in earless breeds,—the reappearance of minute dangling horns in hornless breeds of cattle, more especially, according to Youatt, in young

F373    Book:     Darwin, C. R. 1859. On the origin of species by means of natural selection, or the preservation of favoured races in the struggle for life. London: John Murray. 1st ed., 1st issue.   Text   Image   PDF

inferences briefly recapitulated. That many and grave objections may be advanced against the theory of descent with modification through natural selection, I do not deny. I have endeavoured to give to them their full force. Nothing at first can appear more difficult to believe than that the more complex organs and instincts should have been perfected, not by means superior to, though analogous with, human reason, but by the accumulation of innumerable slight variations, each good for the individual

PC-Virginia-Francis-F373    Printed:    1859   On the origin of species by means of natural selection, or the preservation of favoured races in the struggle for life. London: John Murray. [Francis Darwin's copy]  London   Text   Image   PDF

the recent period all existing islands have been nearly or quite joined to some continent. This view would remove many difficulties, but it would not, I think, explain all the facts in regard to insular productions. In the following remarks I shall not confine myself to the mere question of dispersal; but shall consider some other facts, which bear on the truth of the two theories of independent creation and of descent with modification. The species of all kinds which inhabit oceanic islands

F373    Book:     Darwin, C. R. 1859. On the origin of species by means of natural selection, or the preservation of favoured races in the struggle for life. London: John Murray. 1st ed., 1st issue.   Text   Image   PDF

life, and that the offspring of slightly modified forms or varieties acquire from being crossed increased vigour and fertility. So that, on the one hand, considerable changes in the conditions of life and crosses between greatly modified forms, lessen fertility; and on the other hand, lesser changes in the conditions of life and crosses between less modified forms, increase fertility. Turning to geographical distribution, the difficulties encountered on the theory of descent with modification are

PC-Virginia-Francis-F373    Printed:    1859   On the origin of species by means of natural selection, or the preservation of favoured races in the struggle for life. London: John Murray. [Francis Darwin's copy]  London   Text   Image   PDF

forms — On the succession of the same types within the same areas — Summary of preceding and present chapters. LET us now see whether the several facts and rules relating to the geological succession of organic beings, better accord with the common view of the immutability of species, or with that of their slow and gradual modification, through descent and natural selection. New species have appeared very slowly, one after another, both on the land and in the waters. Lyell has shown that it is

PC-Virginia-Francis-F373    Printed:    1859   On the origin of species by means of natural selection, or the preservation of favoured races in the struggle for life. London: John Murray. [Francis Darwin's copy]  London   Text   Image   PDF

than, parts of high physiological importance. Rudimentary organs may be compared with the letters in a word, still retained in the spelling, but become useless in the pronunciation, but which serve as a clue in seeking for its derivation. On the view of descent with modification, we may conclude that the existence of organs in a rudimentary, imperfect, and useless condition, or quite aborted, far [page] 456 SUMMARY. CHAP. XIII

PC-Virginia-Francis-F373    Printed:    1859   On the origin of species by means of natural selection, or the preservation of favoured races in the struggle for life. London: John Murray. [Francis Darwin's copy]  London   Text   Image   PDF

be given in relation to the distribution of marine animals. On the theory of descent with modification, the great law of the long enduring, but not immutable, succession of the same types within the same areas, is at once explained; for the inhabitants of each quarter of the world will obviously tend to leave in that quarter, during the next succeeding period of time, closely allied though in some degree modified descendants. If the inhabitants of one continent formerly differed greatly from

F373    Book:     Darwin, C. R. 1859. On the origin of species by means of natural selection, or the preservation of favoured races in the struggle for life. London: John Murray. 1st ed., 1st issue.   Text   Image   PDF

embryo becomes at any period of life active and has to provide for itself;—of the embryo apparently having sometimes a higher organisation than the mature animal, into which it is developed. I believe that all these facts can be explained, as follows, on the view of descent with modification. It is commonly assumed, perhaps from monstrosities often affecting the embryo at a very early period, that slight variations necessarily appear at an equally early period. But we have little evidence on

F2293    Book contribution:     Darwin, C. R. 1890. [Letters to and about Adam Sedgwick and the Origin of Species]. In J. W. Clark and T. M. Hughes eds., The life and letters of the Reverend Adam Sedgwick. 2 vols. Cambridge: University Press, vol. 1, pp. 380-1, vol. 2, pp. 356-9.   Text

its written records tell us of its history. Take the case of the bee cells. If your development produced the successive modification of the bee and its cells (which no mortal can prove) final cause would stand good as the directing cause under which the successive generations acted and gradually improved— Passages in your book, like that to which I have alluded (and there are others almost as bad) greatly shocked my moral taste. I think in speculating upon organic descent, you over state the

F376    Book:     Darwin, C. R. 1860. On the origin of species by means of natural selection, or the preservation of favoured races in the struggle for life. London: John Murray. 2d ed., second issue.   Text   Image   PDF

As descent has universally been used in classing together the individuals of the same species, though the males and females and larvæ are sometimes extremely different; and as it has been used in classing varieties which have undergone a certain, and sometimes a considerable amount of modification, may not this same element of descent have been unconsciously used in grouping species under genera, and genera under higher groups, though in these cases the modification has been greater in degree

F376    Book:     Darwin, C. R. 1860. On the origin of species by means of natural selection, or the preservation of favoured races in the struggle for life. London: John Murray. 2d ed., second issue.   Text   Image   PDF

, families, orders, and classes. On this same view of descent with modification, all the great facts in Morphology become intelligible,—whether we look to the same pattern displayed in the homologous organs, to whatever purpose applied, of the different species of a class; or to the homologous parts constructed on the same pattern in each individual animal and plant. On the principle of successive slight variations, not necessarily or generally supervening at a very early period of life, and being

F376    Book:     Darwin, C. R. 1860. On the origin of species by means of natural selection, or the preservation of favoured races in the struggle for life. London: John Murray. 2d ed., second issue.   Text   Image   PDF

those groups, which have within known geological periods undergone much modification, should in the older formations make some slight approach to each other; so that the older members should differ less from each other in some of their characters than do the existing members of the same groups; and this by the concurrent evidence of our best palæontologists seems frequently to be the case. Thus, on the theory of descent with modification, the main facts with respect to the mutual affinities of

F376    Book:     Darwin, C. R. 1860. On the origin of species by means of natural selection, or the preservation of favoured races in the struggle for life. London: John Murray. 2d ed., second issue.   Text   Image   PDF

All the foregoing rules and aids and difficulties in classification are explained, if I do not greatly deceive myself, on the view that the natural system is founded on descent with modification; that the characters which naturalists consider as showing true affinity between any two or more species, are those which have been inherited from a common parent, and, in so far, all true classification is genealogical; that community of descent is the hidden bond which naturalists have been

F380    Book:     Darwin, C. R. 1860. The origin of species by means of natural selection, or the preservation of favoured races in the struggle for life. New York: D. Appleton. New edition, revised and augmented.   Text   Image   PDF

time with a pre-existing closely allied species. And I now know from correspondence, that this coincidence he attributes to generation with modification. The previous remarks on single and multiple centres of creation do not directly bear on another allied question, namely whether all the individuals of the same species have descended from a single pair, or single hermaphrodite, or whether, as some authors suppose, from many individuals simultaneously created. With those organic beings which

F380    Book:     Darwin, C. R. 1860. The origin of species by means of natural selection, or the preservation of favoured races in the struggle for life. New York: D. Appleton. New edition, revised and augmented.   Text   Image   PDF

there has been during the long course of ages much migration from one part of the world to another, owing to former climatal and geographical changes, and to the many occasional and unknown means of dispersal, then we can understand, on the theory of descent with modification, most of the great leading facts in Distribution. We can see why there should be so striking a parallelism in the distribution of organic beings throughout space, and in their geological succession throughout time; for in

F376    Book:     Darwin, C. R. 1860. On the origin of species by means of natural selection, or the preservation of favoured races in the struggle for life. London: John Murray. 2d ed., second issue.   Text   Image   PDF

seem to me simply to follow on the theory of descent with modification through natural selection. We can thus understand how it is that new species come in slowly and successively; how species of different classes do not necessarily change together, or at the same rate, or in the same degree; yet in the long run that all undergo modification to some extent. The extinction of old forms is the almost inevitable consequence of the production of new forms. We can understand why when a species has

F380    Book:     Darwin, C. R. 1860. The origin of species by means of natural selection, or the preservation of favoured races in the struggle for life. New York: D. Appleton. New edition, revised and augmented.   Text   Image   PDF

; and this by the concurrent evidence of our best pal ontologists seems frequently to be the case. Thus, on the theory of descent with modification, the main facts with respect to the mutual affinities of the extinct forms of life to each other and to living forms, seem to me explained in a satisfactory manner. And they are wholly inexplicable on any other view. On this same theory, it is evident that the fauna of any great period in the earth's history will be intermediate in general character

F376    Book:     Darwin, C. R. 1860. On the origin of species by means of natural selection, or the preservation of favoured races in the struggle for life. London: John Murray. 2d ed., second issue.   Text   Image   PDF

of bones being the same in the hand of a man, wing of a bat, fin of the porpoise, and leg of the horse,—the same number of vertebræ forming the neck of the giraffe and of the elephant,—and innumerable other such facts, at once explain themselves on the theory of descent with slow and slight successive modifications. The similarity of pattern in the wing and leg of a bat, though used for such different purpose,—in the jaws and legs of a crab,—in the petals, stamens, and pistils of a flower, is

F376    Book:     Darwin, C. R. 1860. On the origin of species by means of natural selection, or the preservation of favoured races in the struggle for life. London: John Murray. 2d ed., second issue.   Text   Image   PDF

existence of groups subordinate to groups, is the same with varieties as with species, namely, closeness of descent with various degrees of modification. Nearly the same rules are followed in classifying varieties, as with species. Authors have insisted on the necessity of classing varieties on a natural instead of an artificial system; we are cautioned, for instance, not to class two varieties of the pine-apple together, merely because their fruit, though the most important part, happens to be nearly

F376    Book:     Darwin, C. R. 1860. On the origin of species by means of natural selection, or the preservation of favoured races in the struggle for life. London: John Murray. 2d ed., second issue.   Text   Image   PDF

CHAPTER VI. DIFFICULTIES ON THEORY. Difficulties on the theory of descent with modification — Transitions — Absence or rarity of transitional varieties — Transitions in habits of life — Diversified habits in the same species — Species with habits widely different from those of their allies — Organs of extreme perfection — Means of transition — Cases of difficulty — Natura non facit saltum — Organs of small importance — Organs not in all cases absolutely perfect — The law of Unity of Type and

F376    Book:     Darwin, C. R. 1860. On the origin of species by means of natural selection, or the preservation of favoured races in the struggle for life. London: John Murray. 2d ed., second issue.   Text   Image   PDF

complex laws in their degrees and kinds of resemblance to their parents,—in being absorbed into each other by successive crosses, and in other such points,—as do the crossed offspring of acknowledged varieties. On the other hand, these would be strange facts if species have been independently created, and varieties have been produced by secondary laws. If we admit that the geological record is imperfect in an extreme degree, then such facts as the record gives, support the theory of descent with