Origin of Species Variorum

→ be 1859 1860

have been 1861 1866 1869

have been very 1872

→ Its action depends on there being 1859 1860 1861 1866

The result depends on there being 1869

It can act only when there are 1872

→ polity of nature, 1859 1860 1861 1866 1869

natural polity of a district 1872

→ some of the inhabitants of the country undergoing 1859 1860 1861

some of the inhabitants of the country under- going 1866

some of the inhabitants of the country under-going 1869

the 1872

→ kind. 1859 1860 1861 1866 1869

of its existing inhabitants. 1872

↑ 1 blocks not present in 1859 1860 1861 1866 1872; present in 1869

But the effects of natural selection will probably still oftener depend on some few of the inhabitants becoming slowly modified; the mutual relations of the other inhabitants being thus disturbed.

→ On the other hand, 1859 1860 1861 1866

But 1869 1872

→ often only at 1859 1860 1861

generally at only 1866

in effecting changes, at 1869

only at 1872

prevented,

vented,

so that new places in the polity of each island will have

had to

be filled up by

the modification

modifications

of the old inhabitants; and time will

have been

allowed for the varieties in each to become well modified and perfected. When, by renewed elevation, the islands

were

shall be

reconverted

re-converted

into a continental area, there will again →be severe competition: the most favoured or improved varieties will

have been

enabled to spread: there will

have been

much extinction of the less improved forms, and the relative proportional numbers of the various inhabitants of the

reunited

renewed

continent will again

have been

changed; and again there will

have been

a fair field for natural selection to improve still further the inhabitants, and thus

to produce

produce

new species.

That natural selection

always

generally

~~will always~~

will always

acts

act

with extreme

slowness

slowness,

I fully admit. →Its action depends on there being places in the →polity of nature, which can be better

filled through

occupied by

→some of the inhabitants of the country undergoing

modifications

modification

of some →kind. The existence of such places will often depend on physical changes, which are generally very slow, and on the immigration of better adapted forms

being

having been

checked. ↑ But the action of natural selection will probably still oftener depend on some

few of

the inhabitants becoming slowly modified; the mutual relations of many of the other inhabitants being thus disturbed. Nothing can be effected, unless favourable variations occur, and variation itself is apparently always a

~~very~~

very

slow process. The process will often be greatly retarded by free intercrossing. Many will exclaim that these several causes are amply sufficient

~~wholly~~

wholly

neutralise

stop

the

power

action

of natural selection. I do not believe so. →On the other hand, I do believe that natural selection

always