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1859
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1860
1861
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natural selection increased in successive generations 1859 1860 1861 1866 1869
OMIT 1872

its 1859 1860 1861 1866 1869
were increased during successive generations, its 1872

1 blocks not present in 1859 1860; present in 1861 1866 1869 1872
The evidence that accidental mutilations can be inherited is at present very scanty; but the remarkable case observed by Brown-Séquard of epilepsy produced by injuring the spinal chord of guinea-pigs, being inherited, should make us cautious.

There is not sufficient evidence to induce us to believe that mutilations are ever inherited; and I should prefer explaining 1859
There is not sufficient evidence to induce me to believe that mutilations are ever inherited; and I should prefer explaining 1860
So that it will perhaps be safest to look at 1861 1866
Hence it will perhaps be safest to look at 1869 1872

by the long-continued 1859 1860
as due to the long-continued 1861 1866
as due to the 1869
not as cases of inherited mutilations, but as due to the 1872

disuse in their progenitors; 1859 1860 1861 1866
long-continued disuse; 1869 1872

the tarsi are almost always lost in 1859 1860 1861 1866
OMIT 1869 1872

they must be lost 1859 1860 1861 1866
are generally found with their tarsi lost, this must happen 1869 1872

cannot 1859 1860 1861 1866
the tarsi cannot be of much importance or 1869 1872

much 1859 1860 1869 1872
of much importance or be much 1861 1866

inhabiting 1859 1860 1861
(but more are now known) which inhabit 1866 1869
(but more are now known) inhabiting 1872

quadrupeds. We may
believe
imagine
that the
early
early
progenitor of the ostrich
genus had
had
habits like those of
the
a
bustard, and
that,
that
as natural selection increased in successive generations the size and weight of its
body
body,
its legs were used more, and its wings less, until they became incapable of flight.
Kirby has remarked (and I have observed the same fact) that the anterior tarsi, or feet, of many male dung-feeding beetles are
very
very
often broken off; he examined seventeen specimens in his own collection, and not one had even a relic left. In the Onites apelles the tarsi are so habitually lost, that the insect has been described as not having them. In some other genera they are present, but in a rudimentary condition. In the Ateuchus or sacred
bettle
beetle
of the Egyptians, they are totally deficient. There is not sufficient evidence to induce us to believe that mutilations are ever inherited; and I should prefer explaining the entire absence of the anterior tarsi in Ateuchus, and their rudimentary condition in some other genera, by the long-continued effects of disuse in their progenitors; for as the tarsi are almost always lost in many dung-feeding
beetles
beetles,
they must be lost early in
life;
life,
and
and
therefore cannot be much used by these insects.
In some cases we might easily put down to disuse modifications of structure which are wholly, or mainly, due to natural selection. Mr. Wollaston has discovered the remarkable fact that 200
beetles
beetles,
out of the 550 species inhabiting
Madeira
Madeira,
are so far deficient in wings that they cannot fly; and
that,
that
of the twenty-nine endemic genera, no less than twenty-three
genera
genera
have all their species in this condition! Several
facts,—
facts,
— namely,
namely,
that beetles in many parts of the world are
very
very
frequently blown to sea and perish; that the beetles in Madeira, as observed by Mr. Wollaston, lie much
con-
con-