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further, might get 1859 1860 1861 1866
by the wind, gain 1869 1872

those producing seed less 1859 1860 1861 1866
those less well 1869
others less well 1872

dispersal; and this process could not possibly go on in fruit which did not open. 1859 1860 1861 1866
wide dispersal. 1869 1872

Subtitle not present 1859 1860 1861 Compensation and Economy of Growth . 1866 1869 1872
seeds,
seeds
which were a little better
adapted
fitted
to be wafted further, might get an advantage over those producing seed less fitted for dispersal; and this process could not possibly go on in fruit which did not open.
The elder Geoffroy and Goethe propounded, at about the same
time
period,
their law of compensation or balancement of growth; or, as Goethe expressed it, "in order to spend on one side, nature is forced to economise on the other side." I think this holds true to a certain extent with our domestic productions: if nourishment flows to one part or organ in excess, it rarely flows, at least in excess, to another part; thus it is difficult to get a cow to give much milk and to fatten readily. The same varieties of the cabbage do not yield abundant and nutritious foliage and a copious supply of oil-bearing seeds. When the seeds in our fruits become atrophied, the fruit itself gains largely in size and quality. In our poultry, a large tuft of feathers on the head is generally accompanied by a diminished comb, and a large beard by diminished wattles. With species in a state of nature it can hardly be maintained that the law is of universal application; but many good observers, more especially botanists, believe in its truth. I will not, however, here give any instances, for I see hardly any way of distinguishing between the effects, on the one hand, of a part being largely developed through natural selection and another and adjoining part being reduced by this same process or by disuse, and, on the other hand, the actual withdrawal of nutriment from one part owing to the excess of growth in another and adjoining part.
I suspect, also, that some
of the
of the
cases of compensation which have been advanced, and likewise some other facts, may be merged under a more general principle, namely, that natural selection is continually trying to economise
in
in
every part of the organisation. If under