Origin of Species Variorum

→ crossed and 1859 1860 1861 1866

first crossed, and the maximum produced 1869 1872

→ parent-species 1859 1860 1866 1869 1872

parent species in 1861

→ error seems to me 1859 1860 1861 1866 1869

serious error here intervene: a plant, 1872

→ here introduced: a plant to be 1859 1860 1861

here introduced: a plant, to be 1866 1869

OMIT 1872

→ during several years 1859 1860 1861

OMIT 1866 1869 1872

→ the primrose and cowslip, which we have such good reason to believe to be 1859 1860 1861

some forms, such as the common red and blue pimpernels (Anagallis arvensis and cœrulea), which the best botanists rank as 1866 1869 1872

→ only once or twice succeeded in getting fertile seed; as he 1859 1860 1861

OMIT 1866 1869 1872

→ common red and blue pimpernels (Anagallis arvensis and cœrulea), which the best botanists rank as varieties, 1859 1860 1861

OMIT 1866 1869 1872

→ together; and 1859 1860 1861

we may doubt whether many other species are really so sterile, when intercrossed, 1866

we may doubt whether many species are really so sterile, when intercrossed, 1869 1872

→ came to the same concluson in several other analogous cases; it seems to me that we may well be permitted to doubt whether many other species are really so sterile, when intercrossed, as Gärtner 1859

came to the same conclusion in several other analogous cases; it seems to me that we may well be permitted to doubt whether many other species are really so sterile, when intercrossed, as Gärtner 1860

came to the same conclusion in several other analogous cases, it seems to me that we may well be permitted to doubt whether many other species are really so sterile, when intercrossed, as Gärtner 1861

OMIT 1866 1869 1872

unhesitatingly ranks them as varieties. Gärtner, also, makes the rule equally universal; and he disputes the entire fertility of Kölreuter's ten cases. But in these and in many other cases, Gärtner is obliged carefully to count the seeds, in order to show that there is any degree of sterility. He always compares the maximum number of seeds produced by two species when →crossed and by their hybrid offspring, with the average number produced by both pure →parent-species

a state of nature. But

~~a serious~~

a serious

causes

cause

of →error seems to me to be →here introduced: a plant to be

hybridised,

hybridised

must be castrated, and, what is often more important, must be secluded in order to prevent pollen being brought to it by insects from other plants. Nearly all the plants

experimented

experimentised

on by Gärtner were potted, and

~~apparently~~

apparently

were kept in a chamber in his house. That these processes are often injurious to the fertility of a plant cannot be doubted; for Gärtner gives in his table about a score of cases of plants which he castrated, and artificially fertilised with their own pollen, and (excluding all cases such as the Leguminosæ, in which there is an acknowledged difficulty in the manipulation) half of these twenty plants had their fertility in some degree impaired. Moreover, as Gärtner →during several years repeatedly crossed →the primrose and cowslip, which we have such good reason to believe to be varieties, and →only once or twice succeeded in getting fertile seed; as he found

them

the

→common red and blue pimpernels (Anagallis arvensis and cœrulea), which the best botanists rank as varieties, absolutely

sterile,

sterile

→together; and as he →came to the same concluson in several other analogous cases; it seems to me that we may well be permitted to doubt whether many other species are really so sterile, when intercrossed, as Gärtner

believed.

believes.