Origin of Species Variorum

→ from 1859 1860 1861 1866 1872

in all its proportions from 1869

→ in all its proportions, 1859 1860 1861 1866

in 1872

OMIT 1869

→ as much 1859 1860 1861 1866 1869

the same proportions 1872

→ The 1859 1860 1861 1866 1869

These facts are explained by the above 1872

→ two principles above given seem to me to 1859 1860 1861

two principles above given, namely that variations do not generally supervene at a very early age, and that they are inherited at a corresponding age whatever that may have been, seem to me to 1866

above two principles 1869

two principles. 1872

OMIT 1872

several facts regarding the later developmental stages of our domestic varieties. 1866

facts. Fanciers select for breeding their dogs, horses, pigeons, &c., when nearly grown up: they are indifferent whether the desired qualities are acquired earlier or later in life, if the full-grown animal possesses them. 1869

→ And the cases just given, more especially that of pigeons, seem to show that the characteristic differences which give value to each breed, and which have been 1859 1860 1861

Fanciers select their horses, dogs, and pigeons, for breeding, when they are nearly grown up: they are indifferent whether the desired qualities and structures have been 1866

And the cases just given, more especially that of pigeons, show that the characteristic differences which give value to the breeds and which have been 1869

Fanciers select their dogs, horses, pigeons, &c., for breeding, when nearly grown up: they are indifferent whether the desired qualities are 1872

→ accumulated by man's selection, have not generally first appeared at an early period of life, and have been inherited by the offspring at a corresponding not early period. 1859 1861

accumulated by mans selection, have not generally first appeared at an early period of life, and have been inherited by the offspring at a corresponding not early period. 1860

acquired earlier or later in life, if the full-grown animal possesses them. 1866 1872

accumulated by mans selection, have not generally appeared at a very early period of life, and have been inherited at a corresponding not early period. 1869

→ of the short-faced tumbler, which when twelve hours old had acquired its proper proportions, proves that this is not the universal rule; for here 1859 1860 1861

just given, more especially that of pigeons, seem to show that 1866

of the short-faced tumbler, which when twelve hours old possessed its proper characters, proves that this is not the universal rule; for here 1869

just given, more especially that of the pigeons, show that 1872

→ must either have appeared at an earlier period than usual, or, if not so, the differences must 1859 1860 1861 1869

which give value to each breed, and which 1866

which 1872

→ inherited, not at the corresponding, but 1859 1860 1861

accumulated by man's selection, have not generally first appeared 1866

inherited, not at a corresponding, but 1869

accumulated by man's selection, and which give value to his breeds, do not generally appear 1872

→ earlier age. 1859 1860 1861 1869

early period of life, and have been inherited by the offspring at a corresponding not early period. 1866

early period of life, and are inherited at a corresponding not early period. 1872

↑ 1 blocks not present in 1859 1860 1861 1869; present in 1866 1872

But the case of the short-faced tumbler, which when twelve hours old had acquired its proper proportions, proves that this is not the universal rule; for here the characteristic differences must either have appeared at an earlier period than usual, or, if not so, the differences must have been inherited, not at the corresponding, but at an earlier age.

→ facts and the above two principles— which latter, though not proved true, can be shown to be in some degree probable— 1859 1860 1861

facts and the above two principles 1866

two principles 1869 1872

→ on my theory 1859 1860 1861 1866

OMIT 1869 1872

→ one parent-species, and of which the several new species have become 1859 1860 1861 1866

ancient form and 1869 1872

→ in accordance with their diverse 1859 1860 1861 1866

for different 1869 1872

→ at a rather late 1859 1860 1861 1866

in the several species at a not early 1869 1872

young. But there was one remarkable exception to this rule, for the young of the short-faced tumbler differed →from the young of the wild rock-pigeon and of the other breeds, →in all its proportions, almost exactly →as much as in the adult state.

→The →two principles above given seem to me to

~~explain~~

explain

~~these~~

these

→facts in regard to the later embryonic stages of our domestic varieties. Fanciers select their horses, dogs, and pigeons, for breeding, when they are nearly grown up: they are indifferent whether the desired qualities and structures have been acquired earlier or later in life, if the full-grown animal possesses them. →And the cases just given, more especially that of pigeons, seem to show that the characteristic differences which give value to each breed, and which have been →accumulated by man's selection, have not generally first appeared at an early period of life, and have been inherited by the offspring at a corresponding not early period.

And

But

the

cases

case

→of the short-faced tumbler, which when twelve hours old had acquired its proper proportions, proves that this is not the universal rule; for here the characteristic differences →must either have appeared at an earlier period than usual, or, if not so, the differences must have been →inherited, not at the corresponding, but at

a very

→earlier age. ↑

Now let us apply these →facts and the above two principles— which latter, though not proved true, can be shown to be in some degree probable— to species in a state of nature. Let us take a

group

genus

of birds, descended →on my theory from some →one parent-species, and of which the several new species have become modified through natural selection →in accordance with their diverse habits. Then, from the many slight successive

~~steps of~~

steps of

variations

variation

having supervened →at a rather late age, and having been inherited at a corresponding