Origin of Species Variorum

→ in a state of nature— had been 1859 1860 1861

in a state of nature — had been 1866 1869

under nature— were 1872

→ much greater variability, as well as the greater frequency of monstrosities, 1859 1860 1861 1866

fact of variations and monstrosities occurring much more frequently 1869 1872

→ leads me to believe that deviations of structure are in some way due 1859 1860

leads to the belief that deviations of structure are in some way due 1861 1866

and the greater variability of species having wide ranges than of those having restricted ranges, lead 1869

and the greater variability of species having wide ranges than of those with restricted ranges, lead 1872

→ nature of 1859 1860 1861 1866

conclusion that variability is directly related to 1869

conclusion that variability is generally related to 1872

→ the parents and their more remote ancestors have 1859 1860 1861 1866

each species has 1869 1872

→ I have remarked in 1859 1860 1861 1866

In 1869 1872

→ chapter— but a long catalogue of facts which cannot be here given would be necessary 1859 1860 1861

chapter — but a long catalogue of facts which cannot be here given would be necessary 1866

chapter I attempted 1869 1872

→ the truth of the remark— that 1859 1860 1861

the truth of the remark — that 1866

that changed conditions act in two ways, directly on the whole organization or on certain parts alone, and indirectly through 1869

that changed conditions act in two ways, directly on the whole organisation or on certain parts alone, and indirectly through 1872

CHAPTER V.

LAWS OF VARIATION.

Effects of

changed

external

conditions — Use and disuse, combined with natural selection; organs of flight and of vision — Acclimatisation —

Correlated

Correlation

variation

of growth

— Compensation and economy of growth — False correlations — Multiple, rudimentary, and lowly organised structures variable — Parts developed in an unusual manner are highly variable: specific characters more variable than generic: secondary sexual characters variable — Species of the same genus vary in an analogous manner — Reversions to

long lost

long-lost

characters — Summary.

I HAVE hitherto sometimes spoken as if the

variations—

variations —

so common and multiform

with

organic beings under domestication, and in a lesser degree

with

those →in a state of nature— had been due to chance. This, of course, is a wholly incorrect expression, but it serves to acknowledge plainly our ignorance of the cause of each particular variation. Some authors believe it to be as much the function of the reproductive system to produce individual differences, or

~~very~~

very

slight deviations of structure, as to make the child like its parents. But the →much greater variability, as well as the greater frequency of monstrosities, under domestication

~~or cultivation,~~

or cultivation,

than under nature, →leads me to believe that deviations of structure are in some way due to the →nature of the conditions of

life

life,

to which →the parents and their more remote ancestors have been exposed during several

successive generations.

generations.

→I have remarked in the first →chapter— but a long catalogue of facts which cannot be here given would be necessary to show →the truth of the remark— that the reproductive

system.

~~eminently~~

~~susceptible~~

~~changes~~

~~the~~

~~conditions~~

~~life;~~

~~and~~